Equine anatomy encompasses the gross and

microscopic The microscopic scale () is the scale of objects and events smaller than those that can easily be seen by the naked eye, requiring a lens (optics), lens or microscope to see them clearly. In physics, the microscopic scale is sometimes regarded a ...

anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...

of horses, ponies and other

equids Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', ...

, including

donkey The domestic donkey is a hoofed mammal in the family Equidae, the same family as the horse. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a ...

mule The mule is a domestic equine hybrid between a donkey and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two pos ...

s and

zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zeb ...

s. While all anatomical features of equids are described in the same terms as for other animals by the International Committee on Veterinary Gross Anatomical Nomenclature in the book ''

Nomina Anatomica Veterinaria The ''Nomina Anatomica Veterinaria'' (often abbreviated as ''NAV'') is an standardized nomenclature. It is made by World Association of Veterinary Anatomists (WAVA). It is used as the standard reference for anatomical ( zootomical) terminology in ...

'', there are many

horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...

-specific

colloquial Colloquialism (), also called colloquial language, everyday language or general parlance, is the linguistic style used for casual (informal) communication. It is the most common functional style of speech, the idiom normally employed in conver ...

terms used by

equestrians Equestrianism (from Latin , , , 'horseman', 'horse'), commonly known as horse riding (Commonwealth English) or horseback riding (American English), includes the disciplines of riding, driving, and vaulting. This broad description includes the ...

External anatomy

Back The human back, also called the dorsum, is the large posterior area of the human body, rising from the top of the buttocks to the back of the neck. It is the surface of the body opposite from the chest and the abdomen. The vertebral column run ...

: the area where the saddle sits, beginning at the end of the

withers The withers is the ridge between the shoulder blades of an animal, typically a quadruped. In many species, it is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, cattle ar ...

, extending to the last

thoracic vertebrae In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebra (anatomy), vertebrae and they are intermediate in size b ...

(colloquially includes the loin or "coupling," though technically incorrect usage) * Barrel: the body of the horse, enclosing the

rib cage The rib cage, as an enclosure that comprises the ribs, vertebral column and sternum in the thorax of most vertebrates, protects vital organs such as the heart, lungs and great vessels. The sternum, together known as the thoracic cage, is a semi- ...

and the major

internal organs In biology, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to act together in a ...

* Buttock: the part of the hindquarters behind the thighs and below the root of the tail * Cannon or

cannon bone Good conformation in the limbs leads to improved movement and decreased likelihood of injuries. Large differences in bone structure and size can be found in horses used for different activities, but correct conformation remains relatively simil ...

: the area between the knee or hock and the fetlock joint, sometimes called the "shin" of the horse, though technically it is the

third metacarpal The third metacarpal bone (metacarpal bone of the middle finger) is a little smaller than the second. The dorsal aspect of its base presents on its radial side a pyramidal eminence, the styloid process, which extends upward behind the capitate; ...

Chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrelat ...

: a

callosity A callosity is another name for callus, a piece of skin that has become thickened as a result of repeated contact and friction. Primates All Old World monkeys, gibbons, and some chimpanzees have pads on their rears known as ischial callositie ...

on the inside of each leg * Chin groove: the part of the horse's head behind the lower lip and chin, the area that dips down slightly on the lower jaw; area where the

curb chain A curb chain, or curb strap, is a piece of horse tack required for proper use on any type of curb bit. It is a flat linked chain or flat strap that runs under the chin groove of the horse, between the bit shank's purchase arms. It has a buckl ...

of certain bits is fastened * Coupling: ''see'' "Loin" ''below'' *

Coronet A coronet is a small crown consisting of ornaments fixed on a metal ring. A coronet differs from other kinds of crowns in that a coronet never has arches, and from a tiara in that a coronet completely encircles the head, while a tiara does ...

or coronary band: the ring of soft tissue just above the horny hoof that blends into the skin of the leg * Crest: the upper portion of the neck where the mane grows *

Croup Croup, also known as laryngotracheobronchitis, is a type of respiratory infection that is usually caused by a virus. The infection leads to swelling inside the trachea, which interferes with normal breathing and produces the classic symptoms o ...

: the topline of the hindquarters, beginning at the hip, extending proximate to the

sacral vertebrae The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...

and stopping at the dock of the tail (where the

coccygeal vertebrae The coccyx ( : coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other ...

begin); sometimes called "rump" *

Dock A dock (from Dutch language, Dutch ''dok'') is the area of water between or next to one or a group of human-made structures that are involved in the handling of boats or ships (usually on or near a shore) or such structures themselves. The ex ...

: the living part of the tail, consisting of the coccygeal vertebrae,

muscles Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other types of muscle ...

and

ligaments A ligament is the fibrous connective tissue that connects bones to other bones. It is also known as ''articular ligament'', ''articular larua'', ''fibrous ligament'', or ''true ligament''. Other ligaments in the body include the: * Peritoneal li ...

. Sometimes used colloquially to refer to the root of the tail, below. * Elbow: The joint of the front leg at the point where the belly of the horse meets the leg. Homologous to the elbow in humans *

Ergot Ergot ( ) or ergot fungi refers to a group of fungi of the genus ''Claviceps''. The most prominent member of this group is ''Claviceps purpurea'' ("rye ergot fungus"). This fungus grows on rye and related plants, and produces alkaloids that ca ...

: a callosity on the back of the fetlock * Face: the area between the forehead and the tip of the upper lip *

Fetlock Fetlock is the common name in horses, large animals, and sometimes dogs for the metacarpophalangeal and metatarsophalangeal joints (MCPJ and MTPJ). Although it somewhat resembles the human ankle in appearance, the joint is homologous to the ba ...

: sometimes called the "

ankle The ankle, or the talocrural region, or the jumping bone (informal) is the area where the foot and the leg meet. The ankle includes three joints: the ankle joint proper or talocrural joint, the subtalar joint, and the inferior tibiofibular join ...

" of the horse, though it is not the same skeletal structure as an ankle in humans; known to anatomists as the

metacarpophalangeal The metacarpophalangeal joints (MCP) are situated between the metacarpal bones and the proximal phalanges of the fingers. These joints are of the condyloid kind, formed by the reception of the rounded heads of the metacarpal bones into shallow cav ...

(front) or

metatarsophalangeal The metatarsophalangeal joints (MTP joints), also informally known as toe knuckles, are the joints between the metatarsal bones of the foot and the proximal bones (proximal phalanges) of the toes. They are condyloid joints, meaning that an ellipt ...

(hind) joint; homologous to the "

ball A ball is a round object (usually spherical, but can sometimes be ovoid) with several uses. It is used in ball games, where the play of the game follows the state of the ball as it is hit, kicked or thrown by players. Balls can also be used f ...

" of the foot or the metacarpophalangeal joints of the

fingers A finger is a limb of the body and a type of digit, an organ of manipulation and sensation found in the hands of most of the Tetrapods, so also with humans and other primates. Most land vertebrates have five fingers (Pentadactyly). Chambers 1 ...

in humans *

Flank Flank may refer to: * Flank (anatomy), part of the abdomen ** Flank steak, a cut of beef ** Part of the external anatomy of a horse * Flank speed, a nautical term * Flank opening, a chess opening * A term in Australian rules football * The si ...

: where the hind legs and the barrel meet, specifically the area right behind the rib cage and in front of the stifle joint * Forearm: the area of the front leg between the knee and elbow, consisting of the fused

radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...

and

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

, and all the tissue around these bones; anatomically, the antebrachium. * Forehead: the area between the poll, the eyes and the arch of the nose *

Forelock The forelock or foretop is a part of a horse's mane, that grows from the animal's poll and falls forward between the ears and onto the forehead. Some breeds, particularly pony breeds, have a naturally thick forelock, while other breeds, such as ...

: the continuation of the mane, which hangs from between the ears down onto the forehead of the horse *

Frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

: the highly elastic wedge-shaped mass on the underside of the hoof, which normally makes contact with the ground every stride, and supports both the locomotion and circulation of the horse * Gaskin: the large muscle on the hind leg, just above the hock, below the stifle, homologous to the calf of a human * Girth or heartgirth: the area right behind the elbow of the horse, where the

girth Girth may refer to: ;Mathematics * Girth (functional analysis), the length of the shortest centrally symmetric simple closed curve on the unit sphere of a Banach space * Girth (geometry), the perimeter of a parallel projection of a shape * Girth ...

of the saddle would go; this area should be where the barrel is at its greatest diameter in a properly-conditioned horse that is not pregnant or obese * Hindquarters: the large, muscular area of the hind legs, above the stifle and behind the barrel. Can also be used to refer to the back end of a horse. *

Hock Hock may refer to: Common meanings: * Hock (wine), a type of wine * Hock (anatomy), part of an animal's leg * To leave an item with a pawnbroker People: * Hock (surname) * Richard "Hock" Walsh (1948-1999), Canadian blues singer Other uses: * A t ...

: the tarsus of the horse (hindlimb equivalent to the human ankle and heel), the large joint on the hind leg *

Hoof The hoof (plural: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits, yet the rumin ...

: the foot of the horse; the hoof wall is the tough outside covering of the hoof that comes into contact with the ground and is, in many respects, a much larger and stronger version of the human

fingernail A nail is a claw-like plate found at the tip of the fingers and toes on most primates. Nails correspond to the claws found in other animals. Fingernails and toenails are made of a tough protective protein called alpha-keratin, which is a polymer ...

* Jugular groove: the line of indentation on the lower portion of the neck, can be seen from either side, just above the

windpipe The trachea, also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all air-breathing animals with lungs. The trachea extends from the la ...

; beneath this area run the

jugular vein The jugular veins are veins that take deoxygenated blood from the head back to the heart via the superior vena cava. The internal jugular vein descends next to the internal carotid artery and continues posteriorly to the sternocleidomastoid ...

, the

carotid artery Carotid artery may refer to: * Common carotid artery, often "carotids" or "carotid", an artery on each side of the neck which divides into the external carotid artery and internal carotid artery * External carotid artery, an artery on each side of t ...

and part of the

sympathetic trunk The sympathetic trunks (sympathetic chain, gangliated cord) are a paired bundle of nerve fibers that run from the base of the skull to the coccyx. They are a major component of the sympathetic nervous system. Structure The sympathetic trunk lies j ...

* Knee: the

carpus In human anatomy, the wrist is variously defined as (1) the carpus or carpal bones, the complex of eight bones forming the proximal skeletal segment of the hand; "The wrist contains eight bones, roughly aligned in two rows, known as the carpal ...

of the horse (equivalent to the human wrist), the large joint in the front legs, above the cannon bone * Loin: the area right behind the saddle, going from the last rib to the croup, anatomically approximate to the

lumbar spine The lumbar vertebrae are, in human anatomy, the five vertebrae between the rib cage and the pelvis. They are the largest segments of the vertebral column and are characterized by the absence of the foramen transversarium within the transverse p ...

* Mane: long and relatively coarse hair growing from the dorsal ridge of the neck * Muzzle: the chin, mouth, and nostrils of the face *

Pastern The is a part of the leg of a horse between the fetlock and the top of the hoof. It incorporates the long pastern bone (proximal phalanx) and the short pastern bone (middle phalanx), which are held together by two sets of paired ligaments to fo ...

: the connection between the coronet and the fetlock, made up of the middle and proximal phalanx *

Poll Poll, polled, or polling may refer to: Figurative head counts * Poll, a formal election ** Election verification exit poll, a survey taken to verify election counts ** Polling, voting to make decisions or determine opinions ** Polling places o ...

: commonly refers to the poll joint at the beginning of the neck, immediately behind the ears, a slight depression at the joint where the atlas (C1) meets the occipital crest; anatomically, the occipital crest itself is the "poll" * Root of the tail or root of the dock: the point where the tail is "set on" (attached) to the rump; p.38 Sometimes also called the "dock" * Shoulder: made up of the

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...

and associated muscles, runs from the withers to the point of shoulder (the joint at the front of the chest, i.e. the

glenoid The glenoid fossa of the scapula or the glenoid cavity is a bone part of the shoulder. The word ''glenoid'' is pronounced or (both are common) and is from el, gléne, "socket", reflecting the shoulder joint's ball-and-socket form. It is a sha ...

); the angle of the shoulder has a great effect on the horse's movement and jumping ability, and is an important aspect of

equine conformation Equine conformation evaluates a horse's bone structure, musculature, and its body proportions in relation to each other. Undesirable conformation can limit the ability to perform a specific task. Although there are several faults with universa ...

* Splints: bones found on each of the legs, on either side of the cannon bone (8 total); partially

vestigial Vestigiality is the retention, during the process of evolution, of genetically determined structures or attributes that have lost some or all of the ancestral function in a given species. Assessment of the vestigiality must generally rely on co ...

, these bones support the corresponding carpal bones in the forelimb, and the corresponding tarsal bones in the hindlimb; anatomically referred to as

Metacarpal In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ...

Metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...

II (on the medial aspect (inside)) and IV (on the lateral aspect (outside)) *

Stifle joint The stifle joint (often simply stifle) is a complex joint in the hind limbs of quadruped mammals such as the sheep, horse or dog. It is the equivalent of the human knee and is often the largest synovial joint in the animal's body. The stifle joint ...

: corresponds to the knee of a human, consists of the articulation between femur and tibia, as well as the articulation between patella and femur *

Tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...

: the long hairs which grow from the dock; may also include the dock *Throatlatch (also, throttle, throatlash, throat): the point at which the windpipe meets the head at the underside of the jaw, corresponding to where the eponymous part of a

bridle A bridle is a piece of equipment used to direct a horse. As defined in the ''Oxford English Dictionary'', the "bridle" includes both the that holds a bit that goes in the mouth of a horse, and the reins that are attached to the bit. Headgear w ...

goes. *

Withers The withers is the ridge between the shoulder blades of an animal, typically a quadruped. In many species, it is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, cattle ar ...

: the highest point of the thoracic vertebrae, the point just above the tops of the shoulder blades, seen best with horse standing square and head slightly lowered; the height of the horse is measured at the withers.

Digestive system

Topography of Viscera of Horse Left Deep View

Technique of dehydration applied in a foal

Horses and other equids evolved as

grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...

animals, adapted to eating small amounts of the same kind of food all day long. In the wild, the horse adapted to eating

prairie Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...

grasses in semi-arid regions and traveling significant distances each day in order to obtain adequate nutrition. Therefore, the digestive system of a horse is about long, and most of this is intestines.

Mouth

Digestion begins in the

mouth In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on ...

, which is also called the "oral cavity." It is made up of the teeth, the hard palate, the soft palate, the tongue and related muscles, the cheeks and the lips. Horses also have three pairs of salivary glands, the parotoid (largest salivary gland and located near the poll), mandibular (located in the jaw), and sublingual (located under the tongue). Horses select pieces of

forage Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...

and pick up finer foods, such as

grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...

, with their sensitive,

prehensile Prehensility is the quality of an appendage or organ (anatomy), organ that has Adaptation (biology), adapted for grasping or holding. The word is derived from the Latin term ''prehendere'', meaning "to grasp". The ability to grasp is likely der ...

lip The lips are the visible body part at the mouth of many animals, including humans. Lips are soft, movable, and serve as the opening for food intake and in the articulation of sound and speech. Human lips are a tactile sensory organ, and can be ...

s. The front

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

of the horse, called

incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...

s, clip

, and food is then pushed back in the mouth by the

tongue The tongue is a muscular organ (anatomy), organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive system, digestive process, and is the primary organ of taste. The tongue's upper surfa ...

, and ground up for swallowing by the

premolar The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mouth ...

s and

molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...

Esophagus

The

esophagus The esophagus (American English) or oesophagus (British English; both ), non-technically known also as the food pipe or gullet, is an organ in vertebrates through which food passes, aided by peristaltic contractions, from the pharynx to the ...

is about in length, and carries food to the stomach. A muscular ring, called the cardiac sphincter, connects the stomach to the esophagus. This sphincter is very well developed in horses. This and the oblique angle at which the esophagus connects to the stomach explains why horses cannot

vomit Vomiting (also known as emesis and throwing up) is the involuntary, forceful expulsion of the contents of one's stomach through the mouth and sometimes the nose. Vomiting can be the result of ailments like food poisoning, gastroenteriti ...

. The esophagus is also the area of the digestive tract where horses may suffer from choke.

Stomach

Horses have a relatively small

stomach The stomach is a muscular, hollow organ in the gastrointestinal tract of humans and many other animals, including several invertebrates. The stomach has a dilated structure and functions as a vital organ in the digestive system. The stomach i ...

for their size, and this limits the amount of feed a horse can take in at one time. The average sized horse () has a stomach with a capacity of around , and works best when it contains about . Because the stomach empties when full, whether stomach enzymes have completed their processing of the food or not, and doing so prevents full digestion and proper utilization of feed, continuous foraging or several small feedings per day are preferable to one or two large ones. The horse stomach consists of a non-glandular proximal region (saccus cecus), divided by a distinct border, the margo plicatus, from the glandular distal stomach. In the stomach, assorted

acids In computer science, ACID ( atomicity, consistency, isolation, durability) is a set of properties of database transactions intended to guarantee data validity despite errors, power failures, and other mishaps. In the context of databases, a sequ ...

and the enzyme

pepsin Pepsin is an endopeptidase that breaks down proteins into smaller peptides. It is produced in the gastric chief cells of the stomach lining and is one of the main digestive enzymes in the digestive systems of humans and many other animals, w ...

break down food. Pepsin allows for the further breakdown of proteins into

amino acid Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although hundreds of amino acids exist in nature, by far the most important are the alpha-amino acids, which comprise proteins. Only 22 alpha am ...

chains. Other enzymes include

resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...

and

lipase Lipase ( ) is a family of enzymes that catalyzes the hydrolysis of fats. Some lipases display broad substrate scope including esters of cholesterol, phospholipids, and of lipid-soluble vitamins and sphingomyelinases; however, these are usually tr ...

. Additionally, the stomach absorbs some water, as well as ions and lipid-soluble compounds.

Small intestine

The horse's

small intestine The small intestine or small bowel is an organ in the gastrointestinal tract where most of the absorption of nutrients from food takes place. It lies between the stomach and large intestine, and receives bile and pancreatic juice through the p ...

is long and holds . This is the major digestive organ, and where most nutrients are absorbed."Horse Nutrition - The Horse's Digestive System." Bulletin 762-00, Ohio State University.
Web site accessed 9 February 2007. It has three parts, the

duodenum The duodenum is the first section of the small intestine in most higher vertebrates, including mammals, reptiles, and birds. In fish, the divisions of the small intestine are not as clear, and the terms anterior intestine or proximal intestine m ...

jejunum The jejunum is the second part of the small intestine in humans and most higher vertebrates, including mammals, reptiles, and birds. Its lining is specialised for the absorption by enterocytes of small nutrient molecules which have been previous ...

and

ileum The ileum () is the final section of the small intestine in most higher vertebrates, including mammals, reptiles, and birds. In fish, the divisions of the small intestine are not as clear and the terms posterior intestine or distal intestine ma ...

. The majority of digestion occurs in the duodenum while the majority of absorption occurs in the jejunum.

Bile Bile (from Latin ''bilis''), or gall, is a dark-green-to-yellowish-brown fluid produced by the liver of most vertebrates that aids the digestion of lipids in the small intestine. In humans, bile is produced continuously by the liver (liver bile ...

from the liver aids in digesting fats in the duodenum combined with

enzyme Enzymes () are proteins that act as biological catalysts by accelerating chemical reactions. The molecules upon which enzymes may act are called substrates, and the enzyme converts the substrates into different molecules known as products. A ...

s from the

pancreas The pancreas is an organ of the digestive system and endocrine system of vertebrates. In humans, it is located in the abdomen behind the stomach and functions as a gland. The pancreas is a mixed or heterocrine gland, i.e. it has both an end ...

and small intestine. Horses, in common with mammals such as

camels A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...

, do not have a

gall bladder In vertebrates, the gallbladder, also known as the cholecyst, is a small hollow organ where bile is stored and concentrated before it is released into the small intestine. In humans, the pear-shaped gallbladder lies beneath the liver, although ...

, meaning bile flows constantly. Most food is digested and absorbed into the bloodstream from the small intestine, including proteins, simple carbohydrate, fats, and vitamins A, D, . Any remaining liquids and roughage move into the large intestine.

Large intestine

Cecum

The

cecum The cecum or caecum is a pouch within the peritoneum that is considered to be the beginning of the large intestine. It is typically located on the right side of the body (the same side of the body as the appendix (anatomy), appendix, to which i ...

is the first section of the

large intestine The large intestine, also known as the large bowel, is the last part of the gastrointestinal tract and of the digestive system in tetrapods. Water is absorbed here and the remaining waste material is stored in the rectum as feces before being r ...

. It is also known as the "water gut" or "hind gut". It is a cul-de-sac pouch, about long that holds . It contains

bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...

that digest

cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...

plant

fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...

through

fermentation Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food ...

. These bacteria feed upon chyme digestive, and also produce certain fat-soluble vitamins which are absorbed by the horse. The reason horses must have their diets changed slowly is so the bacteria in the cecum are able to modify and adapt to the different chemical structure of new feedstuffs. Too abrupt a change in diet can cause

colic Colic or cholic () is a form of pain that starts and stops abruptly. It occurs due to muscular contractions of a hollow tube ( small and large intestine, gall bladder, ureter, etc.) in an attempt to relieve an obstruction by forcing content out ...

, as the new food is not properly digested.

Other section of the large intestine

The large colon, small colon, and

rectum The rectum is the final straight portion of the large intestine in humans and some other mammals, and the Gastrointestinal tract, gut in others. The adult human rectum is about long, and begins at the rectosigmoid junction (the end of the s ...

make up the remainder of the

. The large colon is long and holds up to of semi-liquid matter. It is made up of the right

ventral Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Th ...

(lower) colon, the left ventral colon, the left

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...

(upper) colon, the right dorsal colon, and the transverse colon, in that order. Three

flexure A flexure is a flexible element (or combination of elements) engineered to be compliant in specific degrees of freedom. Flexures are a design feature used by design engineers (usually mechanical engineers) for providing adjustment or compliance ...

s are also named; the sternal flexure, between right and left ventral colon; the pelvic flexure, between left ventral and left dorsal colon; the diaphragmatic flexure, between left dorsal and right dorsal colon. The main purpose of the large colon is to absorb carbohydrates, which were broken down from cellulose in the cecum. Due to its many twists and turns, it is a common place for a type of

horse colic Colic in horses is defined as abdominal pain, but it is a clinical symptom rather than a diagnosis. The term colic can encompass all forms of gastrointestinal conditions which cause pain as well as other causes of abdominal pain not involving the g ...

called an impaction. The small colon is in length and holds only of material. It is the area where the majority of water in the horse's diet is absorbed, and is the place where

fecal Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...

lumps are formed. The

is about long, and acts as a holding chamber for waste matter, which is then expelled from the body via the

anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, d ...

Reproductive system

Mare

The mare's reproductive system is responsible for controlling gestation, birth, and lactation, as well as her estrous cycle and mating behavior. It lies ventral to the 4th or 5th lumbar vertebrae, although its position within the mare can vary depending on the movement of the intestines and distention of the bladder. The mare has two ovaries, usually in length and thick, that generally tend to decrease in size as the mare ages. In equine ovaries, unlike in humans, the vascular tissue is cortical to follicular tissue, so ovulation can only occur at an ovulation fossa near the infundibulum. The ovaries connect to the fallopian tubes (oviducts), which serve to move the ovum from the ovary to the uterus. To do so, the oviducts are lined with a layer of

cilia The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projecti ...

, which produce a current that flows toward the uterus. Each oviduct attaches to one of the two horns of the uterus, which are approximately in length. These horns attach to the body of the uterus ( long). The equine uterus is bipartite, meaning the two

uterine horns The uterine horns (cornua of uterus) are the points in the upper uterus where the fallopian tubes exit to meet the ovaries. They are one of the points of attachment for the round ligament of uterus (the other being the mons pubis). They also prov ...

fuse into a relatively large uterine body (resembling a shortened bicornuate uterus or a stretched

simplex In geometry, a simplex (plural: simplexes or simplices) is a generalization of the notion of a triangle or tetrahedron to arbitrary dimensions. The simplex is so-named because it represents the simplest possible polytope in any given dimension. ...

uterus). Caudal to the uterus is the cervix, about long, which separates the uterus from the vagina. Usually in diameter with longitudinal folds on the interior surface, it can expand to allow the passage of the foal. The vagina of the mare is long, and is quite elastic, allowing it to expand. The vulva is the external opening of the vagina, and consists of the clitoris and two labia. It lies ventral to the rectum. The mare has two mammary glands, which are smaller in maiden mares. They have two ducts each, which open externally.

Stallion

The stallion's reproductive system is responsible for his

sexual behavior Human sexual activity, human sexual practice or human sexual behaviour is the manner in which humans experience and express their sexuality. People engage in a variety of sexual acts, ranging from activities done alone (e.g., masturbation) ...

and

secondary sex characteristics Secondary sex characteristics are features that appear during puberty in humans, and at sexual maturity in other animals. These characteristics are particularly evident in the sexually dimorphic phenotypic traits that distinguish the sexes of a sp ...

(such as a large crest). The external genitalia include the

urethra The urethra (from Greek οὐρήθρα – ''ourḗthrā'') is a tube that connects the urinary bladder to the urinary meatus for the removal of urine from the body of both females and males. In human females and other primates, the urethra con ...

; the

testes A testicle or testis (plural testes) is the male reproductive gland or gonad in all bilaterians, including humans. It is homologous to the female ovary. The functions of the testes are to produce both sperm and androgens, primarily testoster ...

, which average long; the

penis A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males do n ...

, which, when housed within the prepuce, is long and in diameter with the distal end and when erect, increases by 3 to . The internal genitalia

accessory sex glands Male accessory glands (MAG) in humans are the seminal vesicles, prostate gland, and the bulbourethral glands (also called Cowper's glands). In insects, male accessory glands produce products that mix with the sperm to protect and preserve them, in ...

are the vesicular glands,

prostate gland The prostate is both an accessory gland of the male reproductive system and a muscle-driven mechanical switch between urination and ejaculation. It is found only in some mammals. It differs between species anatomically, chemically, and physio ...

, and

bulbourethral glands The bulbourethral glands or Cowper's glands (named for English anatomist William Cowper) are two small exocrine glands in the reproductive system of many male mammals (of all domesticated animals, they are absent only in dogs). They are homologo ...

, which contribute fluid to the

semen Semen, also known as seminal fluid, is an organic bodily fluid created to contain spermatozoa. It is secreted by the gonads (sexual glands) and other sexual organs of male or hermaphroditic animals and can fertilize the female ovum. Semen i ...

ejaculation Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the male reproductory tract as a result of an orgasm. It is the final stage and natural objective of male sexual stimulation, and an essential componen ...

, but are not strictly necessary for fertility.

Teeth

A horse's teeth include

s, molars, and sometimes canine teeth. A horse's incisors, premolars, and molars, once fully developed, continue to erupt throughout its lifetime as the grinding surface is worn down through chewing. Because of this pattern of wear, a rough estimate of a horse's age can be made from an examination of the teeth. Abnormal wear of the teeth, caused by conformational defects, abnormal behaviors, or improper diets, can cause serious health issues and can even result in the death of the horse.

Feet/hooves

The

hoof The hoof (plural: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits, yet the rumin ...

of the horse encases the second and third

phalanx The phalanx ( grc, φάλαγξ; plural phalanxes or phalanges, , ) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar pole weapons. The term is particularly use ...

of the lower limbs, analogous to the fingertip or toe tip of a human. In essence, a horse travels on its "tiptoes". The hoof wall is a much larger, thicker and stronger version of the human

or toenail, made up of similar materials, primarily

keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, ho ...

, a very strong

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...

molecule. The horse's hoof contains a high proportion of

sulfur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...

-containing

amino acids Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although hundreds of amino acids exist in nature, by far the most important are the alpha-amino acids, which comprise proteins. Only 22 alpha am ...

which contribute to its resilience and toughness. Vascular fold-like structures called laminae suspend the distal phalanx from the hoof wall.

Skeletal system

The

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

of the horse has three major functions in the body. It protects vital organs, provides framework, and supports soft parts of the body. Horses have , which are divided into the appendicular skeleton (the legs) and the axial skeleton (the

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordata, ...

sternum The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Sh ...

, and

ribs The rib cage, as an enclosure that comprises the ribs, vertebral column and sternum in the thorax of most vertebrates, protects vital organs such as the heart, lungs and great vessels. The sternum, together known as the thoracic cage, is a semi- ...

). Both pelvic and thoracic limbs contain the same number of bones, per limb. Bones are connected to muscles via tendons and other bones via ligaments. Bones are also used to store minerals, and are the site of

red blood cell Red blood cells (RBCs), also referred to as red cells, red blood corpuscles (in humans or other animals not having nucleus in red blood cells), haematids, erythroid cells or erythrocytes (from Greek ''erythros'' for "red" and ''kytos'' for "holl ...

formation. *The Appendicular system includes the

limbs of the horse Good conformation in the limbs leads to improved movement and decreased likelihood of injuries. Large differences in bone structure and size can be found in horses used for different activities, but correct conformation remains relatively simil ...

; *The Axial system is composed of the spine, ribs and skull; The bones of the horse are the same as those of other domestic species, but the third metacarpal and metatarsal are much more developed and the second and fourth are undeveloped, having the first and fifth metacarpal and metatarsal.Riegal, Ronald J. DVM, and Susan E. Hakola RN. Illustrated Atlas of Clinical Equine Anatomy and Common Disorders of the Horse Vol. II. Equistar Publication, Limited. Marysville, OH. Copyright 2000.

Ligaments and tendons

Ligaments

Ligament A ligament is the fibrous connective tissue that connects bones to other bones. It is also known as ''articular ligament'', ''articular larua'', ''fibrous ligament'', or ''true ligament''. Other ligaments in the body include the: * Peritoneal li ...

s attach bone to bone or bone to tendon, and are vital in stabilizing joints as well as supporting structures. They are made up of fibrous material that is generally quite strong. Due to their relatively poor blood supply, ligament injuries generally take a long time to heal.

Tendons

Tendon A tendon or sinew is a tough, high-tensile-strength band of dense fibrous connective tissue that connects muscle to bone. It is able to transmit the mechanical forces of muscle contraction to the skeletal system without sacrificing its ability ...

s are cords of connective tissue attaching muscle to bone, cartilage or other tendons. They are a major contributor to shock absorption, are necessary for support of the horse's body, and translate the force generated by muscles into movement. Tendons are classified as flexors (flex a joint) or extensors (extend a joint). However, some tendons will flex multiple joints while extending another (the flexor tendons of the hind limb, for example, will flex the

fetlock Fetlock is the common name in horses, large animals, and sometimes dogs for the metacarpophalangeal and metatarsophalangeal joints (MCPJ and MTPJ). Although it somewhat resembles the human ankle in appearance, the joint is homologous to the ba ...

, pastern, and coffin joint, but extend the hock joint). In this case, the tendons (and associated muscles) are named for their most distal action (digital flexion). Tendons form in the embryo from fibroblasts which become more tightly packed as the tendon grows. As tendons develop they lay down

collagen Collagen () is the main structural protein in the extracellular matrix found in the body's various connective tissues. As the main component of connective tissue, it is the most abundant protein in mammals, making up from 25% to 35% of the whole ...

, which is the main structural protein of connective tissue. As tendons pass near bony prominences, they are protected by a fluid filled synovial structure, either a

tendon sheath A tendon sheath is a layer of synovial membrane around a tendon. It permits the tendon to stretch and not adhere to the surrounding fascia. It has two layers: * synovial sheath A synovial sheath is one of the two membranes of a tendon sheath wh ...

or a sac called a

bursa ( grc-gre, Προῦσα, Proûsa, Latin: Prusa, ota, بورسه, Arabic:بورصة) is a city in northwestern Turkey and the administrative center of Bursa Province. The fourth-most populous city in Turkey and second-most populous in the ...

. Tendons are easily damaged if placed under too much strain, which can result in a painful, and possibly career-ending, injury.

Tendinitis Tendinopathy, a type of tendon disorder that results in pain, swelling, and impaired function. The pain is typically worse with movement. It most commonly occurs around the shoulder (rotator cuff tendinitis, biceps tendinitis), elbow (tennis elb ...

is most commonly seen in high performance horses that gallop or jump. When a tendon is damaged the healing process is slow because tendons have a poor blood supply, reducing the availability of nutrients and oxygen to the tendon. Once a tendon is damaged the tendon will always be weaker, because the collagen fibres tend to line up in random arrangements instead of the stronger linear pattern. Scar tissue within the tendon decreases the overall elasticity in the damaged section of the tendon as well, causing an increase in strain on adjacent uninjured tissue.

Muscular system

Leisering's Atlas of the Anatomy of the Horse Muscles

When a muscle contracts, it pulls a tendon, which acts on the horse's bones to move them. Muscles are commonly arranged in pairs so that they oppose each other (they are "antagonists"), with one flexing the joint (a flexor muscle) and the other extending it (extensor muscle). Therefore, one muscle of the pair must be relaxed in order for the other muscle in the pair to contract and bend the joint properly. A muscle is made up of several muscle bundles, which in turn are made up of muscle fibers. Muscle fibers have

myofibril A myofibril (also known as a muscle fibril or sarcostyle) is a basic rod-like organelle of a muscle cell. Skeletal muscles are composed of long, tubular cells known as muscle fibers, and these cells contain many chains of myofibrils. Each myofib ...

s, which are able to contract due to

actin Actin is a family of globular multi-functional proteins that form microfilaments in the cytoskeleton, and the thin filaments in muscle fibrils. It is found in essentially all eukaryotic cells, where it may be present at a concentration of over ...

and

myosin Myosins () are a superfamily of motor proteins best known for their roles in muscle contraction and in a wide range of other motility processes in eukaryotes. They are ATP-dependent and responsible for actin-based motility. The first myosin ...

. A muscle together with its tendon and bony attachments form an extensor or flexor unit.

Respiratory system and smell

The horse's respiratory system consists of the nostrils, pharynx, larynx, trachea, diaphragm, and lungs. Additionally, the nasolacrimal duct and sinuses are connected to the nasal passage. The horse's respiratory system not only allows the animal to breathe, but also is important in the horse's sense of smell (olfactory ability) as well as in communicating. The soft palate blocks off the

pharynx The pharynx (plural: pharynges) is the part of the throat behind the mouth and nasal cavity, and above the oesophagus and trachea (the tubes going down to the stomach and the lungs). It is found in vertebrates and invertebrates, though its struc ...

from the mouth (oral cavity) of the horse, except when swallowing. This helps prevent the horse from inhaling food, but also means that a horse cannot use its mouth to breathe when in respiratory distress—a horse can only breathe through its nostrils, also called

obligate nasal breathing Obligate nasal breathing describes a physiological necessity to breathe through the nose (or other forms of external nares, depending on the species) as opposed to breathing through the mouth. Definition The term may be misleading, as it implie ...

. For this same reason, horses also cannot

pant Pant may refer to: Clothing * Pants or trousers, an article of outer clothing worn on the lower half of the body * Underpants, an item of underwear Places * Pant, Denbighshire, Wales; a township of Llysfaen *Pant, Merthyr Tydfil, Wales *Pan ...

as a method of

thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...

. The genus '' Equus'' also has a unique part of the respiratory system called the

guttural pouch Guttural pouches are large, auditory-tube diverticula that contain between 300 and 600 ml of air. They are present in odd-toed mammals, some bats, hyraxes, and the American forest mouse. They are paired bilaterally just below the ears, behind the ...

, which is thought to equalize air pressure on the

tympanic membrane In the anatomy of humans and various other tetrapods, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped membrane that separates the external ear from the middle ear. Its function is to transmit sound from the air ...

. Located between the

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

s but below the

occiput The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone overlies the occipital lobes of the cereb ...

, it fills with air when the horse swallows or exhales.

Circulatory system

The horse's circulatory system includes the four-chambered heart, averaging in weight, as well as the blood and blood vessels. Its main purpose is to circulate blood throughout the body to deliver oxygen and nutrients to tissues, and to remove waste from these tissues. The hoof (including the frog - the V shaped part on the bottom of the horses hoof) is a very important part of the circulatory system. As the horse puts weight onto the hoof, the hoof wall is pushed outwards and the frog compressed, driving blood out of the frog, the digital pad, and the laminae of the hoof. When weight is removed from the hoof, the release of pressure pulls blood back down into the foot again. This effectively creates an auxiliary blood-pumping system at the end of each leg. Some of this effect may be lost when a horse is shod (eliminating the expansion and contraction of the hoof wall and raising the frog higher from the ground).

The eye

The horse has one of the largest eyes of all land mammals. Eye size in mammals is significantly correlated to maximum running speed as well as to body size, in accordance with Leuckart's law; animals capable of fast locomotion require large eyes. The eye of the horse is set to the side of its skull, consistent with that of a prey animal. The horse has a wide field of

monocular vision In human species Monocular vision vision is known as seeing and using only one eye in the human species. Depth perception in monocular vision is reduced compared to binocular vision, but still is active primarily due to accommodation of the eye a ...

, as well as good visual acuity. Horses have two-color, or dichromatic vision, which is somewhat like

red-green color blindness Color blindness or color vision deficiency (CVD) is the decreased ability to see color or differences in color. It can impair tasks such as selecting ripe fruit, choosing clothing, and reading traffic lights. Color blindness may make some aca ...

in humans. Because the horse's vision is closely tied to behavior, the horse's visual abilities are often taken into account when handling and training the animal.

Hearing

The hearing of horses is good, superior to that of humans, and the pinna of each ear can rotate up to 180°, giving the potential for 360° hearing without having to move the head.Myers ''Horse Safe'' p.7 Often, the eye of the horse is looking in the same direction as the ear is directed.

References

External links

Veterinary Anatomical lllustrations, University of Wisconsin Digital Collections
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