CRuMs
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CRuMS
CRuMs is a proposed clade of microbial eukaryotes, whose name is an acronym of the following constituent groups: i) collodictyonids also known as diphylleids, ii) rigifilids and iii) mantamonadids as sister of the Amorphea. It more or less supersedes Varisulca, as Ancyromonadida Ancyromonadida or Planomonadida is a small group of biflagellated protists found in the soil and in aquatic habitats, where they feed on bacteria.Cavalier-Smith, T. (2013)Early evolution of eukaryote feeding modes, cell structural diversity, and ... are inferred not to be specifically related to the orders Collodictyonida, Rigifilida and Mantamonadida. Phylogeny References {{Taxonbar, from=Q59153571 Eukaryote subphyla Podiata ...
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Varisulca
Varisulca was a proposed basal Podiate taxon. It encompassed several lineages of heterotrophic protists, most notably the ancyromonads (planomonads), collodictyonids (diphylleids), rigifilids (''Rigifila'', ''Micronuclearia'') and mantamonadids. Recent evidence suggests that the latter three are closely related to each other, forming a clade called CRuMs, but that this is unlikely to be specifically related to ancyromonads Cavalier-Smith had proposed the new subphylum Varisulca which consists of the classes Hilomonadea, Diphyllatea and Glissodiscea. The validity of this proposed taxonomy has yet to ruled upon by the Society of Protistologists. It is unlikely to be widely accepted, since Varisulca appears to be paraphyletic or polyphyletic (ancyromonads are not inferred to be the sister group to CRuMs). Glissodiscea and Multirhiza (a taxon encompassing Diphyllatea and Glissodiscea) also appear paraphyletic or polyphyletic, since ''Mantamonas'' belongs to CRuMs but ancyrom ...
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Mantamonadida
The Mantamonadidae are of Motility, free-living heterotrophic flagellates that move primarily by gliding on surfaces (rather than swimming). There is one genus, ''Mantamonas''. It has been suggested previously that the Mantamonadidae be classified in Apusozoa as sister of the Apusomonadida, Apusmonadida on the basis of Ribosomal RNA, rRNA analyses. However, mantamonads are currently placed in CRuMs on the basis of phylogenomic analyses that identify their closest relatives as the collodictyonids (=diphylleids) and ''Rigifila''. Taxonomy * Order Mantamonadida Cavalier-Smith Glücksman et al. 2011 ** Family Mantamonadidae Cavalier-Smith Glücksman et al. 2011 *** Genus ''Mantamonas'' Cavalier-Smith Glücksman et al. 2011 **** Species ''Mantamonas plastica'' Cavalier-Smith & Glücksman 2011 Phylogeny References

Podiata orders {{eukaryote-stub ...
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Mantamonadids
The Mantamonadidae are of free-living heterotrophic flagellates that move primarily by gliding on surfaces (rather than swimming). There is one genus, '' Mantamonas''. It has been suggested previously that the Mantamonadidae be classified in Apusozoa as sister of the Apusmonadida on the basis of rRNA analyses. However, mantamonads are currently placed in CRuMs CRuMs is a proposed clade of microbial eukaryotes, whose name is an acronym of the following constituent groups: i) collodictyonids also known as diphylleids, ii) rigifilids and iii) mantamonadids as sister of the Amorphea. It more or less s ... on the basis of phylogenomic analyses that identify their closest relatives as the collodictyonids (=diphylleids) and ''Rigifila''. Taxonomy * Order Mantamonadida Cavalier-Smith Glücksman et al. 2011 ** Family Mantamonadidae Cavalier-Smith Glücksman et al. 2011 *** Genus '' Mantamonas'' Cavalier-Smith Glücksman et al. 2011 **** Species '' Mantamonas plastica'' Cavalier- ...
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Rigifilids
Rigifilida is a clade of non-ciliate phagotrophic eukaryotes. It consists of two genera: ''Micronuclearia'' and '' Rigifila''. Characteristics Cells of rigifilids are covered with either a single or a double-layered submembrane pellicular lamina that makes them rigid in consistence. Slender branching filopodia emanate from a ventral aperture of the cell and are employed to collect bacteria upon which they feed and to attach the organism to the substratum. Around this aperture, the pellicle is reflexed around forming a peristomial collar. Other notable features are flat and irregular shaped mitocondrial cristae, a single dorsal nucleus and the lack of centrioles and cilia. Phylogeny Taxonomy Rigifilida is currently placed in CRuMs. *Order Rigifilida Cavalier-Smith 2012 icronucleariida Cavalier-Smith 2008** Family Rigifilidae Yabuki & Cavalier-Smith 2012 *** Genus '' Rigifila'' Yabuki & Cavalier-Smith 2012 **** Species ''Rigifila ramosa'' Yabuki & Cavalier-Smith 2012 ** Fami ...
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Rigifilida
Rigifilida is a clade of non-ciliate phagotrophic eukaryotes. It consists of two genera: ''Micronuclearia'' and '' Rigifila''. Characteristics Cells of rigifilids are covered with either a single or a double-layered submembrane pellicular lamina that makes them rigid in consistence. Slender branching filopodia emanate from a ventral aperture of the cell and are employed to collect bacteria upon which they feed and to attach the organism to the substratum. Around this aperture, the pellicle is reflexed around forming a peristomial collar. Other notable features are flat and irregular shaped mitocondrial cristae, a single dorsal nucleus and the lack of centrioles and cilia. Phylogeny Taxonomy Rigifilida is currently placed in CRuMs. *Order Rigifilida Cavalier-Smith 2012 icronucleariida Cavalier-Smith 2008** Family Rigifilidae Yabuki & Cavalier-Smith 2012 *** Genus '' Rigifila'' Yabuki & Cavalier-Smith 2012 **** Species '' Rigifila ramosa'' Yabuki & Cavalier-Smith 2012 ** Fam ...
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Collodictyonidae
Collodictyonidae (also Diphylleidae) is a group of aquatic, unicellular eukaryotic organisms with two to four terminal flagella. They feed by phagocytosis, ingesting other unicellular organisms like algae and bacteria. The most remarkable fact of this clade is its uncertain position in the tree of life. Recent molecular analyses place Collodictyonids (e.g. Collodictyon) in a clade also containing Rigifilida and Mantamonadidae. This clade has been named CRuMs and is sister to Amorphea. Phylogeny Taxonomy * Class Diphyllatea Cavalier-Smith 2003 nisomonadea; Diphyllatia Cavalier-Smith 2003** Order Diphylleida Cavalier-Smith 1993 ollodictyonida *** Family Sulcomonadidae Cavalier-Smith 2012 **** Genus '' Sulcomonas'' Brugerolle 2006 ***** Species ''S. lacustris'' Brugerolle 2006 *** Family Diphylleidae Cavalier-Smith 1993 ollodictyonidae Brugerolle et al. 2002**** Genus ''Diphylleia'' Massart 1920 non Michaux 1803 'Aulacomonas'' Skuja 1939">Aulacomonas.html" ;"title="'Aula ...
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Collodictyonids
Collodictyonidae (also Diphylleidae) is a group of aquatic, unicellular eukaryotic organisms with two to four terminal flagella. They feed by phagocytosis, ingesting other unicellular organisms like algae and bacteria. The most remarkable fact of this clade is its uncertain position in the tree of life. Recent molecular analyses place Collodictyonids (e.g. Collodictyon) in a clade also containing Rigifilida and Mantamonadidae. This clade has been named CRuMs and is sister to Amorphea. Phylogeny Taxonomy * Class Diphyllatea Cavalier-Smith 2003 nisomonadea; Diphyllatia Cavalier-Smith 2003** Order Diphylleida Cavalier-Smith 1993 ollodictyonida *** Family Sulcomonadidae Cavalier-Smith 2012 **** Genus '' Sulcomonas'' Brugerolle 2006 ***** Species ''S. lacustris'' Brugerolle 2006 *** Family Diphylleidae Cavalier-Smith 1993 ollodictyonidae Brugerolle et al. 2002**** Genus ''Diphylleia'' Massart 1920 non Michaux 1803 'Aulacomonas'' Skuja 1939">Aulacomonas.html" ;"title="'Aulacomo ...
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Eukaryote
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Euka ...
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Eukaryote Subphyla
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Eukaryo ...
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Ancyromonadida
Ancyromonadida or Planomonadida is a small group of biflagellated protists found in the soil and in aquatic habitats, where they feed on bacteria.Cavalier-Smith, T. (2013)Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa European journal of protistology, 49(2), 115-178. Includes freshwater or marine organisms, benthic, dorsoventrally compressed and with two unequal flagellae, each emerging from a separate pocket. The apical anterior flagellum can be very thin or end in the cell membrane, while the posterior flagellum is long and is inserted ventrally or laterally. The cell membrane is supported by a thin single layer teak and the mitochondrial crests are discoidal / flat. The group's placement is doubtful, as it seems to fall outside the five supergroups of eukaryotes. Cavalier-Smith considers that they constitute a basal group to Amoebozoa and Opisthokonta and places it together w ...
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Opimoda
The Scotokaryotes (Cavalier-Smith) is a proposed basal Neokaryote clade as sister of the Diaphoretickes. Basal Scotokaryote groupings are the Metamonads, the Malawimonas and the Podiata. In this phylogeny the Discoba are sometimes seen as paraphyletic and basal Eukaryotes. An alternative to the Unikont–Bikont division was suggested by Derelle ''et al.'' in 2015, where they proposed the acronyms Opimoda–Diphoda respectively, as substitutes to the older terms. The name Opimoda is formed from the letters (shown in capitals) of OPIsthokonta and aMOebozoa. In this phylogeny Discoba belongs to the Diphoda clade. Taxonomy A proposed cladogram is See also *Diphoda A bikont ("two flagella") is any of the eukaryotic organisms classified in the group Bikonta. Many single-celled members of the group, and the presumed ancestor, have two flagella. Enzymes Another shared trait of bikonts is the fusion of two ge ... References {{Taxonbar, from=Q21399998 Eukaryote unranked ...
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Malawimonadea
Malawimonadidae is a group of unicellular eukaryotes of outsize importance in understanding eukaryote phylogeny. Phylogeny Taxonomy * Phylum Neolouka Cavalier-Smith 2013 ** Class Malawimonadea Cavalier-Smith 2003 *** Order Malawimonadida Cavalier-Smith 2003 **** Family Malawimonadidae O’Kelly & Nerad 1999 ***** Genus '' Gefionella'' Heiss, Ekelund & Simpson 2018 ****** Species '' G. okellyi'' Heiss, Ekelund & Simpson 2018 ***** Genus ''Malawimonas ''Malawimonas'' is a Loukozoa genus, possible sister of the Podiata. History of the discovery of Malawimonads In 1993, Charles J O’ Kelly studied the jakobid groups flagellates and implications for the early diversification of eukaryotes and ...'' O’Kelly & Nerad 1999 ****** Species '' M. californiana'' ****** Species '' M. jakobiformis'' O’Kelly & Nerad 1999 References Excavata families {{Excavata-stub ...
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