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Caryophyllales
Caryophyllales ( ) is a diverse and heterogeneous order of flowering plants that includes the cacti, carnations, amaranths, ice plants, beets, and many carnivorous plants. Many members are succulent, having fleshy stems or leaves. The betalain pigments are unique in plants of this order and occur in all its families with the exception of Caryophyllaceae and Molluginaceae. Description The members of Caryophyllales include about 6% of eudicot species. This order is part of the core eudicots. Currently, the Caryophyllales contains 37 families, 749 genera, and 11,620 species The monophyly of the Caryophyllales has been supported by DNA sequences, cytochrome c sequence data and heritable characters such as anther wall development and vessel-elements with simple perforations. Circumscription As with all taxa, the circumscription of Caryophyllales has changed within various classification systems. All systems recognize a core of families with centrospermous ovules and seeds. Mor ...
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Caryophyllineae
Caryophyllineae is a suborder of flowering plants. Systematics Caryophyllales is separated into 2 sub-orders: Caryophyllineae and Polygonineae. Caryophyllineae contains 21 families and 8,600 species and major families include Aizoaceae, Basellaceae, Caryophyllaceae, Didiereaceae, Phytolaccaceae (including Petiveriaceae), Nyctaginaceae, Molluginaceae, Amaranthaceae (sometimes including Chenopodiaceae), Cactaceae, and Portulacaceae.Judd et al. (2008). Plant Systematics: A Phylogenetic Approach, Third Edition. Sinauer Associates, Inc. Sunderland, MAM This roughly constitutes the Caryophyllales as defined before the Angiosperm Phylogeny Group expanded it to include the plants which can be classified in the Polygonales or Polygonineae (depending on whether they are considered an order or a suborder of Caryophyllales). The Caryophyllineae is sometimes called "core Caryophyllales" and the Polygonineae is sometimes called the "non-core Caryophyllales". The core Caryophyllineae sub-ord ...
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Centrospermae
''Centrospermae'' is a descriptive botanical name, published in 1878 by Eichler, meaning "with the seed in the center", referring to the free (central) placentation. It was used in the Engler system and the Wettstein system) for an order of flowering plants. In its circumscription, Centrospermae corresponds fairly closely to the order Caryophyllales in the system of Cronquist. In the APG III system, and in later versions of the Kubitzki system, Caryophyllales is defined to include much more than Centrospermae. Molecular phylogenetic studies have shown that Centrospermae is monophyletic. It is equivalent to a clade known as the core Caryophyllales, whenever the latter is defined to exclude the families Rhabdodendraceae, Simmondsiaceae, Physenaceae and Asteropeiaceae. Samuel F. Brockington, Roolse Alexandre, Jeremy Ramdial, Michael J. Moore, Sunny Crawley, Amit Dhingra, Khidir Hilu, Douglas E. Soltis, and Pamela S. Soltis. 2009. "Phylogeny of the Caryophyllales sensu la ...
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Portulacineae
Portulacineae is a suborder of flowering plants in the order Caryophyllales comprising the families Anacampserotaceae, Basellaceae, Cactaceae (cacti), Didiereaceae, Halophytaceae, Montiaceae, Portulacaceae, and Talinaceae. All three major kinds of succulent plant — stem succulents, leaf succulents, and caudiciform A caudex (plural: caudices) of a plant is a stem, but the term is also used to mean a rootstock and particularly a basal stem structure from which new growth arises.pages 456 and 695 In the strict sense of the term, meaning a stem, "caudex" is m ... plants — are represented within this suborder. References * * Caryophyllales {{caryophyllales-stub ...
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Betalain
Betalains are a class of red and yellow tyrosine-derived pigments found in plants of the order Caryophyllales, where they replace anthocyanin pigments. Betalains also occur in some higher order fungi. They are most often noticeable in the petals of flowers, but may color the fruits, leaves, stems, and roots of plants that contain them. They include pigments such as those found in beets. Description The name "betalain" comes from the Latin name of the common beet (''Beta vulgaris''), from which betalains were first extracted. The deep red color of beets, bougainvillea, amaranth, and many cacti results from the presence of betalain pigments. The particular shades of red to purple are distinctive and unlike that of anthocyanin pigments found in most plants. There are two categories of betalains: * Betacyanins include the reddish to violet betalain pigments. Among the betacyanins present in plants include betanin, isobetanin, probetanin, and neobetanin. * Betaxanthins are those be ...
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Carnivorous Plant
Carnivorous plants are plants that derive some or most of their nutrients from trapping and consuming animals or protozoans Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic tissues and debris. Histo ..., typically insects and other arthropods. Carnivorous plants still generate some of their energy from photosynthesis. Carnivorous plants have adapted to grow in places where the soil is thin or poor in soil nutrient, nutrients, especially nitrogen, such as acidic bogs. They can be found on all continents except Antarctica, as well as many Pacific islands. In 1875 Charles Darwin published ''Insectivorous Plants (book), Insectivorous Plants'', the first treatise to recognize the significance of carnivory in plants, describing years of painstaking research. True carnivory is believed to have convergent evoluti ...
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Eudicot
The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the later evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots. Numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Some common and familiar eudicots include sunflower, dandelion, forget-me-not, cabbage, apple, buttercup, maple, and macadamia. Most leafy trees of midlatitudes also belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, and ''Ginkgo biloba'', which is not an angiosperm. Description The close relationships among flowering plants with tricolpate po ...
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Aizoaceae
The Aizoaceae, or fig-marigold family, is a large family of dicotyledonous flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...s containing 135 genus, genera and about 1800 species. They are commonly known as ice plants or carpet weeds. They are often called vygies in South Africa and New Zealand. Highly Succulent plant, succulent species that resemble stones are sometimes called mesembs. Description The family Aizoaceae is widely recognised by taxonomists. It once went by the botanical name "Ficoidaceae", now disallowed. The APG II system of 2003 (unchanged from the APG system of 1998) also recognizes the family, and assigns it to the order Caryophyllales in the clade core eudicots. The APG II system also classes the former families Mesembryanthemaceae Fenzl, S ...
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Molluginaceae
The Molluginaceae are a family of flowering plants recognized by several taxonomists. It was previously included in the larger family Aizoaceae. The APG III system of 2009 made no change in the status of the family as compared to the APG II system of 2003 and the APG system of 1998, apart from a reassignment of several genera, such as the placement of ''Corrigiola'' and '' Telephium'' into Caryophyllaceae, '' Corbichonia'' in Lophiocarpaceae, '' Microtea'' into Microteaceae and '' Limeum'' in Limeaceae, because the family was found to be widely polyphyletic in Caryophyllales. In addition ''Macarthuria'' was found not to be related to '' Limeum'' as previously thought and thus it was placed in Macarthuriaceae, and similarly species formerly placed in ''Hypertelis'', apart from type species ''Hypertelis spergulacea'', a true Molluginaceae, were found to belong elsewhere and were described as '' Kewa'' in the family Kewaceae, named for the Royal Botanic Gardens Kew. Molluginaceae i ...
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Core Eudicots
The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the later evolutionary divergence of tricolpate dicotyledon, dicots from earlier, less specialized, dicots. Numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Some common and familiar eudicots include sunflower, dandelion, forget-me-not, cabbage, apple, buttercup, maple, and macadamia. Most leafy trees of midlatitudes also belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, and ''Ginkgo biloba'', which is not an angiosperm. Description The close relationships among flowering plants with poll ...
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Caryophyllaceae
Caryophyllaceae, commonly called the pink family or carnation family, is a family of flowering plants. It is included in the dicotyledon order Caryophyllales in the APG III system, alongside 33 other families, including Amaranthaceae, Cactaceae, and Polygonaceae. It is a large family, with 81 genera and about 2,625 known species. This cosmopolitan family of mostly herbaceous plants is best represented in temperate climates, with a few species growing on tropical mountains. Some of the more commonly known members include pinks and carnations (''Dianthus''), and firepink and campions ('' Lychnis'' and ''Silene''). Many species are grown as ornamental plants, and some species are widespread weeds. Most species grow in the Mediterranean and bordering regions of Europe and Asia. The number of genera and species in the Southern Hemisphere is rather small, although the family does contain Antarctic pearlwort (''Colobanthus quitensis''), the world's southernmost dicot, which is one ...
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Kewaceae
''Kewa'' is a genus of flowering plants, consisting of eight species of succulent sub-woody plants, native to eastern and southern Africa, including Saint Helena and Madagascar. These are small shrubs or herbs that form cushions and have edible, acid-tasting leaves. ''Kewa'' is the only genus in the family Kewaceae. The species were formerly included in the genus ''Hypertelis'' of the family Molluginaceae, but molecular studies have shown that most species did not belong there, but were rather distantly related to Molluginaceae, being placed in a clade comprising Aizoaceae, Gisekiaceae and Barbeuiaceae. Only the type species ''Hypertelis spergulacea'' remains in Molluginaceae; all others are transferred to the genus ''Kewa'', which was named for Kew, where the Royal Botanic Gardens, Kew are situated. Species Species transferred to ''Kewa'' from ''Hypertelis'' in 2014 were: *''Kewa acida'' (Hook.f.) Christenh. – Saint Helena *''Kewa angrae-pequenae'' (Friedrich) Christenh. – N ...
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Cactus
A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Greek word (''káktos''), a name originally used by Theophrastus for a spiny plant whose identity is now not certain. Cacti occur in a wide range of shapes and sizes. Although some species live in quite humid environments, most cacti live in habitats subject to at least some drought. Many live in extremely dry environments, even being found in the Atacama Desert, one of the driest places on Earth. Because of this, cacti show many adaptations to conserve water. For example, almost all cacti are succulents, meaning they have thickened, fleshy parts adapted to store water. Unlike many other succulents, the stem is the only part of most cacti where this vital process takes place. Most species of cacti have lost true leaves, retaining only spines, ...
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