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Caryophyllineae
Caryophyllineae is a suborder of flowering plants. Systematics Caryophyllales is separated into 2 sub-orders: Caryophyllineae and Polygonineae. Caryophyllineae contains 21 families and 8,600 species and major families include Aizoaceae, Basellaceae, Caryophyllaceae, Didiereaceae, Phytolaccaceae (including Petiveriaceae), Nyctaginaceae, Molluginaceae, Amaranthaceae (sometimes including Chenopodiaceae), Cactaceae, and Portulacaceae.Judd et al. (2008). Plant Systematics: A Phylogenetic Approach, Third Edition. Sinauer Associates, Inc. Sunderland, MAM This roughly constitutes the Caryophyllales as defined before the Angiosperm Phylogeny Group expanded it to include the plants which can be classified in the Polygonales or Polygonineae (depending on whether they are considered an order or a suborder of Caryophyllales). The Caryophyllineae is sometimes called "core Caryophyllales" and the Polygonineae is sometimes called the "non-core Caryophyllales". The core Caryophyllineae sub-ord ...
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Centrospermae
''Centrospermae'' is a descriptive botanical name, published in 1878 by Eichler, meaning "with the seed in the center", referring to the free (central) placentation. It was used in the Engler system and the Wettstein system) for an order of flowering plants. In its circumscription, Centrospermae corresponds fairly closely to the order Caryophyllales in the system of Cronquist. In the APG III system, and in later versions of the Kubitzki system, Caryophyllales is defined to include much more than Centrospermae. Molecular phylogenetic studies have shown that Centrospermae is monophyletic. It is equivalent to a clade known as the core Caryophyllales, whenever the latter is defined to exclude the families Rhabdodendraceae, Simmondsiaceae, Physenaceae and Asteropeiaceae. Samuel F. Brockington, Roolse Alexandre, Jeremy Ramdial, Michael J. Moore, Sunny Crawley, Amit Dhingra, Khidir Hilu, Douglas E. Soltis, and Pamela S. Soltis. 2009. "Phylogeny of the Caryophyllales sensu la ...
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Caryophyllaceae
Caryophyllaceae, commonly called the pink family or carnation family, is a family of flowering plants. It is included in the dicotyledon order Caryophyllales in the APG III system, alongside 33 other families, including Amaranthaceae, Cactaceae, and Polygonaceae. It is a large family, with 81 genera and about 2,625 known species. This cosmopolitan family of mostly herbaceous plants is best represented in temperate climates, with a few species growing on tropical mountains. Some of the more commonly known members include pinks and carnations (''Dianthus''), and firepink and campions ('' Lychnis'' and ''Silene''). Many species are grown as ornamental plants, and some species are widespread weeds. Most species grow in the Mediterranean and bordering regions of Europe and Asia. The number of genera and species in the Southern Hemisphere is rather small, although the family does contain Antarctic pearlwort (''Colobanthus quitensis''), the world's southernmost dicot, which is one ...
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Portulacaceae
The Portulacaceae are a family of flowering plants, comprising 115 species in a single genus ''Portulaca''. Formerly some 20 genera with about 500 species, were placed there, but it is now restricted to encompass only one genus, the other genera being placed elsewhere. The family has been recognised by most taxonomists, and is also known as the purslane family. It has a cosmopolitan distribution, with the highest diversity in semiarid regions of the Southern Hemisphere in Africa, Australia, and South America, but with a few species also extending north into Arctic regions. The family is very similar to the Caryophyllaceae, differing in the calyx, which has only two sepals. The APG II system (2003; unchanged from the APG system of 1998) assigns it to the order Caryophyllales in the clade core eudicots. In the APG III system, several genera were moved to the Montiaceae, Didiereaceae, Anacampserotaceae and Talinaceae, thus making the family monotypic and only containing the genus ' ...
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Perisperm
In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the female gametophyte (formed from a haploid megaspore) in its center. The female gametophyte — specifically termed a ''megagametophyte''— is also called the ''embryo sac'' in angiosperms. The megagametophyte produces an egg cell for the purpose of fertilization. The ovule is a small structure present in the ovary. It is attached to the placenta by a stalk called a funicle. The funicle provides nourishment to the ovule. Location within the plant In flowering plants, the ovule is located inside the portion of the flower called the gynoecium. The ovary of the gynoecium produces one or more ovules and ultimately becomes the fruit wall. Ovules are attached to the placenta in the ovary through a stalk-like structure known as a ''funiculus'' (p ...
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Embryo
An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm cell. The resulting fusion of these two cells produces a single-celled zygote that undergoes many cell divisions that produce cells known as blastomeres. The blastomeres are arranged as a solid ball that when reaching a certain size, called a morula, takes in fluid to create a cavity called a blastocoel. The structure is then termed a blastula, or a blastocyst in mammals. The mammalian blastocyst hatches before implantating into the endometrial lining of the womb. Once implanted the embryo will continue its development through the next stages of gastrulation, neurulation, and organogenesis. Gastrulation is the formation of the three germ layers that will form all of the different parts of the body. Neurulation forms the nervous ...
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Tepals
A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very similar appearance), as in ''Magnolia'', or because, although it is possible to distinguish an outer whorl of sepals from an inner whorl of petals, the sepals and petals have similar appearance to one another (as in ''Lilium''). The term was first proposed by Augustin Pyramus de Candolle in 1827 and was constructed by analogy with the terms "petal" and "sepal". (De Candolle used the term ''perigonium'' or ''perigone'' for the tepals collectively; today, this term is used as a synonym for ''perianth''.) p. 39. Origin Undifferentiated tepals are believed to be the ancestral condition in flowering plants. For example, '' Amborella'', which is thought to have separated earliest in the evolution of flowering plants, has flowers with undiffer ...
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Betalains
Betalains are a class of red and yellow tyrosine-derived pigments found in plants of the order Caryophyllales, where they replace anthocyanin pigments. Betalains also occur in some higher order fungi. They are most often noticeable in the petals of flowers, but may color the fruits, leaves, stems, and roots of plants that contain them. They include pigments such as those found in beets. Description The name "betalain" comes from the Latin name of the common beet (''Beta vulgaris''), from which betalains were first extracted. The deep red color of beets, bougainvillea, amaranth, and many cacti results from the presence of betalain pigments. The particular shades of red to purple are distinctive and unlike that of anthocyanin pigments found in most plants. There are two categories of betalains: * Betacyanins include the reddish to violet betalain pigments. Among the betacyanins present in plants include betanin, isobetanin, probetanin, and neobetanin. * Betaxanthins are those be ...
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Polygonales
Polygonales was an order of flowering plants recognized by several older systems such as the Wettstein system, last revised in 1935, the Engler system, in its update of 1964, and the Cronquist system, 1981. Its circumscription was typically: * order Polygonales *: family Polygonaceae In these systems the order is placed close to the order Caryophyllales (or its predecessor Centrospermae). Cronquist placed this order in his subclass Caryophyllidae Caryophyllidae is a botanical name at the rank of subclass. At the moment there is no complete consensus about what orders it includes, except that it presumably contains the order Caryophyllales. Note that this is only a naming difficulty: wh ... of three orders. Dahlgren placed the order Polygonales within the superorder Polygoniflorae, alternatively Polygonanae. References {{Reflist Historically recognized angiosperm orders ...
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Angiosperm Phylogeny Group
The Angiosperm Phylogeny Group (APG) is an informal international group of systematic botanists who collaborate to establish a consensus on the taxonomy of flowering plants (angiosperms) that reflects new knowledge about plant relationships discovered through phylogenetic studies. , four incremental versions of a classification system have resulted from this collaboration, published in 1998, 2003, 2009 and 2016. An important motivation for the group was what they considered deficiencies in prior angiosperm classifications since they were not based on monophyletic groups (i.e., groups that include all the descendants of a common ancestor). APG publications are increasingly influential, with a number of major herbaria changing the arrangement of their collections to match the latest APG system. Angiosperm classification and the APG In the past, classification systems were typically produced by an individual botanist or by a small group. The result was a large number of systems ( ...
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Polycarpaea Corymbosa
''Polycarpaea'' is a genus of plants in the family Caryophyllaceae. The genus was named by Jean-Baptiste Lamarck in 1792. Molecular phylogenetic analysis showed that the genus is polyphyletic and needs to be redefined. It falls into three distinct clades. One of these, a group of species related to ''P. corymbosa'', has acquired the photosynthetic pathway. Selected species It contains the following species (but this list may be incomplete): * '' Polycarpaea akkensis'' Maire. * '' Polycarpaea aristata'' Chr.Sm. * '' Polycarpaea balfourii'' Briq. * '' Polycarpaea caespitosa'' Balf.f. * '' Polycarpaea carnosa'' Chr.Sm. * '' Polycarpaea clavifolia'' M.Gilbert. * ''Polycarpaea corymbosa'' (L.) Lam. * ''Polycarpaea divaricata'' Aiton. * ''Polycarpaea filifolia'' Webb ex Christ. * ''Polycarpaea gaudichaudii'' Gagnepain. * ''Polycarpaea gayi'' Webb. * '' Polycarpaea guardafuiensis'' M.Gilbert. * ''Polycarpaea hassalensis'' D.F.Chamb. * ''Polycarpaea hayoides'' D.F.Chamb. * ''Polyca ...
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Aizoaceae
The Aizoaceae, or fig-marigold family, is a large family of dicotyledonous flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...s containing 135 genus, genera and about 1800 species. They are commonly known as ice plants or carpet weeds. They are often called vygies in South Africa and New Zealand. Highly Succulent plant, succulent species that resemble stones are sometimes called mesembs. Description The family Aizoaceae is widely recognised by taxonomists. It once went by the botanical name "Ficoidaceae", now disallowed. The APG II system of 2003 (unchanged from the APG system of 1998) also recognizes the family, and assigns it to the order Caryophyllales in the clade core eudicots. The APG II system also classes the former families Mesembryanthemaceae Fenzl, S ...
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