Archeognathus
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Archeognathus
''Archeognathus'' is a fossilized jaw apparatus of a large predatory conodont from the Ordovician period ( Darriwilian to Sandbian stages). Its large size has made classification difficult, and it has historically been compared to conodonts and gnathostomes (jawed fish) since its remains were first discovered in Missouri. Complete articulated jaw apparatus of ''Archeognathus primus'' are common in the Winneshiek Shale The Winneshiek Shale (originally the Winneshiek Lagerstätte) is a Middle Ordovician (Darriwilian-age) geological formation in Iowa. The formation is restricted to the Decorah crater, an impact crater near Decorah, Iowa. Despite only being discove ... lagerstätte of Iowa, allowing its identity as a conodont to be secured. The jaw apparatus of ''Archeognathus primus'' is much more simple than that of most conodonts, with only six elements in its jaw. These include a pair of long, small-toothed, sawblade-like "coleodiform" S elements at the front of the jaw, ...
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Winneshiek Shale
The Winneshiek Shale (originally the Winneshiek Lagerstätte) is a Middle Ordovician (Darriwilian-age) geological formation in Iowa. The formation is restricted to the Decorah crater, an impact crater near Decorah, Iowa. Despite only being discovered in 2005, the Winneshiek Shale is already renowned for the exceptional preservation of its fossils. The shale preserves a unique ecosystem, the Winneshiek biota, which is among the most remarkable Ordovician lagerstätten in the United States. Fossils include the oldest known eurypterid, ''Pentecopterus'', as well as giant conodonts such as ''Iowagnathus'' and ''Archeognathus''. Geology The Winneshiek Shale is a thin and geologically homogeneous package of dark grey to greenish-brown sandy shale. Drill core data has estimated a maximum thickness of 38 meters, though in most areas its thickness is only about 18–27 meters. The shale is replete with pyrite and organic carbon. It lies solely within the Decorah Structure, a 5.6 km ...
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Conodont
Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which are usually found in isolation and are now called conodont elements. Knowledge about soft tissues remains limited. They existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Conodont elements are widely used as index fossils, fossils used to define and identify geological periods. The animals are also called Conodontophora (conodont bearers) to avoid ambiguity. Discovery and understanding of conodonts The teeth-like fossils of the conodont were first discovered by Heinz Christian Pander and the results published in Saint Petersburg, Russia, in 1856. The name ''pander'' is commonly used in scientific names of conodonts. It was only in the early 1980s that the first fossil evidence of ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Darriwilian
The Darriwilian is the upper stage of the Middle Ordovician. It is preceded by the Dapingian and succeeded by the Upper Ordovician Sandbian Stage. The lower boundary of the Darriwilian is defined as the first appearance of the graptolite species ''Undulograptus austrodentatus'' around million years ago. It lasted for about 8.9 million years until the beginning of the Sandbian around million years ago. This stage of the Ordovician was marked by the beginning of the Andean-Saharan glaciation. Naming The name Darriwilian is derived from Darriwil, a parish in County of Grant, Victoria (Australia). The name was proposed in 1899 by Thomas Sergeant Hall. GSSP The GSSP of the Darriwilian is the Huangnitang Section () near the village Huangnitang, 3.5 km southwest of Changshan County Town (Zhejiang, China). It is an outcrop of the Ningkuo Formation, consisting of mainly black shale. The lower boundary of the Darriwilian is defined as the first appearance datum of the graptolite s ...
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Sandbian
The Sandbian is the first stage of the Upper Ordovician. It follows the Darriwilian and is succeeded by the Katian. Its lower boundary is defined as the first appearance datum of the graptolite species '' Nemagraptus gracilis'' around million years ago. The Sandbian lasted for about 5.4 million years until the beginning of the Katian around million years ago. Naming The name Sandbian is derived from the village Södra Sandby (Lund Municipality, Skåne County, Sweden). The name was proposed in 2006. GSSP The GSSP of the Sandbian is the Fågelsång section () at Sularp Brook, east of Lund (Skåne, Sweden). It is an outcrop of shale and mudstone. The lower boundary of the Sandbian is defined as the first appearance datum of graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and ...
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Gnathostomata
Gnathostomata (; from Greek: (') "jaw" + (') "mouth") are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth (a characteristic which has subsequently been lost in some), paired appendages (pectoral and pelvic fins, arms, legs, wings, etc.), the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed a pair of both pectoral and pelvic fins. Until recently these ancestors, know ...
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Missouri
Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee): Iowa to the north, Illinois, Kentucky and Tennessee to the east, Arkansas to the south and Oklahoma, Kansas and Nebraska to the west. In the south are the Ozarks, a forested highland, providing timber, minerals, and recreation. The Missouri River, after which the state is named, flows through the center into the Mississippi River, which makes up the eastern border. With more than six million residents, it is the List of U.S. states and territories by population, 19th-most populous state of the country. The largest urban areas are St. Louis, Kansas City, Missouri, Kansas City, Springfield, Missouri, Springfield and Columbia, Missouri, Columbia; the Capital city, capital is Jefferson City, Missouri, Jefferson City. Humans have inhabited w ...
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Lagerstätte
A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Lagerstätten'' span geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Jurassic Posidonia Shale and Solnhofen Limestone, the Cretaceous Yixian, Santana, and Agua Nueva formations, the Eocene Green River Formation, the Miocene Foulden Maar and Ashfall F ...
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Iowa
Iowa () is a state in the Midwestern region of the United States, bordered by the Mississippi River to the east and the Missouri River and Big Sioux River to the west. It is bordered by six states: Wisconsin to the northeast, Illinois to the east and southeast, Missouri to the south, Nebraska to the west, South Dakota to the northwest, and Minnesota to the north. During the 18th and early 19th centuries, Iowa was a part of French Louisiana and Spanish Louisiana; its state flag is patterned after the flag of France. After the Louisiana Purchase, people laid the foundation for an agriculture-based economy in the heart of the Corn Belt. In the latter half of the 20th century, Iowa's agricultural economy transitioned to a diversified economy of advanced manufacturing, processing, financial services, information technology, biotechnology, and green energy production. Iowa is the 26th most extensive in total area and the 31st most populous of the 50 U.S. states, with a populat ...
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Wastebasket Taxon
Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the sole purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either paraphyletic or polyphyletic, and are therefore not considered valid taxa under strict cladistic rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an evolutionary grade, however. The term was coined in a 1985 essay by Steven Jay Gould. Examples There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups ...
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Enigmatic Prehistoric Animal Genera
Enigmatic is an adjective meaning "mysterious" or "puzzling". It may also refer to: * ''Enigmatic'', a 1970 album by Czesław Niemen * '' Enigmatic: Calling'', a 2005 album by Norwegian progressive metal band Pagan's Mind * Enigmatic scale, musical scale used by Verdi and others * "The Enigmatic", a song by Joe Satriani on the album '' Not of This Earth'' See also * Enigmatic leaf turtle, a species of Asian leaf turtle * Enigmatic moray eel, a species found in the Pacific and Indian Oceans * ''Glaresis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...'', a genus of beetles sometimes called "enigmatic scarab beetles" * Enigma (other) {{disambig ...
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Ordovician Animals
The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian Perio ...
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