Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organism
s. It is a polyphyletic
grouping that includes species from multiple distinct clade
s. Included organisms range from unicellular microalgae
, such as ''Chlorella
and the diatom
s, to multicellular
forms, such as the giant kelp
, a large brown alga
which may grow up to in length. Most are aquatic and autotrophic
and lack many of the distinct cell and tissue types, such as stoma
, which are found in land plants
. The largest and most complex marine algae are called seaweed
s, while the most complex freshwater forms are the Charophyta
, a division
of green algae which includes, for example, ''Spirogyra
'' and stonewort
No definition of algae is generally accepted. One definition is that algae "have chlorophyll
as their primary photosynthetic pigment and lack a sterile covering of cells around their reproductive cells". Likewise, the colorless Prototheca
are all devoid of any chlorophyll. Although cyanobacteria
are often referred to as "blue-green algae", most authorities exclude all prokaryotes
from the definition of algae.
Algae constitute a polyphyletic
since they do not include a common ancestor, and although their plastid
s seem to have a single origin, from cyanobacteria,
they were acquired in different ways. Green algae
are examples of algae that have primary chloroplast
s derived from endosymbiotic
s and brown algae are examples of algae with secondary chloroplasts derived from an endosymbiotic red alga
. Algae exhibit a wide range of reproductive strategies, from simple asexual
cell division to complex forms of sexual reproduction
Algae lack the various structures that characterize land plants, such as the phyllids (leaf-like structures) of bryophyte
s in nonvascular plants
, and the root
, and other organs
found in tracheophyte
s (vascular plants
). Most are phototroph
ic, although some are mixotroph
ic, deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy
, or phagotrophy
. Some unicellular species of green algae
, many golden algae
s, and other algae have become heterotroph
s (also called colorless or apochlorotic algae), sometimes parasitic, relying entirely on external energy sources and have limited or no photosynthetic apparatus. Some other heterotrophic organisms, such as the apicomplexans
, are also derived from cells whose ancestors possessed plastids, but are not traditionally considered as algae. Algae have photosynthetic machinery ultimately derived from cyanobacteria
that produce oxygen
as a by-product of photosynthesis, unlike other photosynthetic bacteria such as purple
and green sulfur bacteria
. Fossilized filamentous algae from the Vindhya
basin have been dated back to 1.6 to 1.7 billion years ago.
Because of the wide range of types of algae, they have increasing different industrial and traditional applications in human society. Traditional seaweed farming
practices have existed for 1000s of years and have strong traditions in East Asia food cultures. More modern algaculture
applications extend the food traditions
for other applications include cattle feed, using algae for bioremediation
or pollution control, transforming sunlight into algae fuel
s or other chemicals used in industrial processes, and in medical and scientific applications. A 2020 review, found that these applications of algae could play an important role in carbon sequestration
in order to mitigate climate change
while providing valuable value-add products for global economies.
Etymology and study
The singular is the Latin word for 'seaweed' and retains that meaning in English. The etymology
is obscure. Although some speculate that it is related to Latin , 'be cold', no reason is known to associate seaweed with temperature. A more likely source is , 'binding, entwining'.
The Ancient Greek
word for 'seaweed' was (), which could mean either the seaweed (probably red algae) or a red dye derived from it. The Latinization, , meant primarily the cosmetic rouge. The etymology is uncertain, but a strong candidate has long been some word related to the Biblical (), 'paint' (if not that word itself), a cosmetic eye-shadow used by the ancient Egyptians and other inhabitants of the eastern Mediterranean. It could be any color: black, red, green, or blue.
Accordingly, the modern study of marine and freshwater algae is called either phycology
or algology, depending on whether the Greek or Latin root is used. The name ''fucus'' appears in a number of taxa
The committee on the International Code of Botanical Nomenclature has recommended certain suffixes for use in the classification of algae. These are -phyta for division, ''-phyceae'' for class, ''-phycideae'' for subclass, ''-ales'' for order, ''-inales'' for suborder, ''-aceae'' for family, ''-oidease'' for subfamily, a Greek-based name for genus, and a Latin-based name for species.
Algal characteristics basic to primary classification
The primary classification of algae is based on certain morphological features. The chief among these are (a) pigment constitution of the cell, (b) chemical nature of stored food materials, (c) kind, number, point of insertion and relative length of the flagella on the motile cell, (d) chemical composition of cell wall and (e) presence or absence of a definitely organized nucleus in the cell or any other significant details of cell structure.
History of classification of algae
Although Carolus Linnaeus
(1754) included algae along with lichens in his 25th class Cryptogamia, he did not elaborate further on the classification of algae.
Jean Pierre Étienne Vaucher
(1803) was perhaps the first to propose a system of classification of algae, and he recognized three groups, Conferves, Ulves, and Tremelles. While Johann Heinrich Friedrich Link
(1820) classified algae on the basis of the colour of the pigment and structure, William Henry Harvey
(1836) proposed a system of classification on the basis of the habitat and the pigment. J. G. Agardh
(1849–1898) divided algae into six orders: Diatomaceae, Nostochineae, Confervoideae, Ulvaceae, Floriadeae and Fucoideae. Around 1880, algae along with fungi were grouped under Thallophyta, a division created by Eichler (1836). Encouraged by this, Adolf Engler
and Karl A. E. Prantl
(1912) proposed a revised scheme of classification of algae and included fungi in algae as they were of opinion that fungi have been derived from algae. The scheme proposed by Engler and Prantl is summarised as follows:
# Eumycetes (Fungi)
The algae contain chloroplast
s that are similar in structure to cyanobacteria. Chloroplasts contain circular DNA
like that in cyanobacteria and are interpreted as representing reduced endosymbiotic cyanobacteria. However, the exact origin of the chloroplasts is different among separate lineages of algae, reflecting their acquisition during different endosymbiotic events. The table below describes the composition of the three major groups of algae. Their lineage relationships are shown in the figure in the upper right. Many of these groups contain some members that are no longer photosynthetic. Some retain plastids, but not chloroplasts, while others have lost plastids entirely.
based on plastid
not nucleocytoplasmic genealogy:
, in ''Species Plantarum
'' (1753), the starting point for modern botanical nomenclature
, recognized 14 genera of algae, of which only four are currently considered among algae. In ''Systema Naturae
'', Linnaeus described the genera ''Volvox
'' and ''Corallina
'', and a species of ''Acetabularia
'' (as ''Madrepora
''), among the animals.
In 1768, Samuel Gottlieb Gmelin
(1744–1774) published the ''Historia Fucorum'', the first work dedicated to marine algae and the first book on marine biology
to use the then new binomial nomenclature of Linnaeus. It included elaborate illustrations of seaweed and marine algae on folded leaves.
W. H. Harvey
(1811–1866) and Lamouroux
were the first to divide macroscopic algae into four divisions based on their pigmentation. This is the first use of a biochemical criterion in plant systematics. Harvey's four divisions are: red algae (Rhodospermae), brown algae (Melanospermae), green algae (Chlorospermae), and Diatomaceae.
At this time, microscopic algae were discovered and reported by a different group of workers (e.g., O. F. Müller
) studying the Infusoria
(microscopic organisms). Unlike macroalgae
, which were clearly viewed as plants, microalgae
were frequently considered animals because they are often motile.
Even the nonmotile (coccoid) microalgae were sometimes merely seen as stages of the lifecycle of plants, macroalgae, or animals.
Although used as a taxonomic category in some pre-Darwinian classifications, e.g., Linnaeus (1753), de Jussieu (1789), Horaninow (1843), Agassiz (1859), Wilson & Cassin (1864), in further classifications, the "algae" are seen as an artificial, polyphyletic group.
Throughout the 20th century, most classifications treated the following groups as divisions or classes of algae: cyanophyte
s, and chlorophyte
s. Later, many new groups were discovered (e.g., Bolidophyceae
), and others were splintered from older groups: charophyte
s and glaucophyte
s (from chlorophytes), many heterokontophyte
s (e.g., synurophytes
from chrysophytes, or eustigmatophyte
s from xanthophytes), haptophyte
s (from chrysophytes), and chlorarachniophyte
s (from xanthophytes).
With the abandonment of plant-animal dichotomous classification, most groups of algae (sometimes all) were included in Protist
a, later also abandoned in favour of Eukaryota
. However, as a legacy of the older plant life scheme, some groups that were also treated as protozoa
ns in the past still have duplicated classifications (see ambiregnal protist
Some parasitic algae (e.g., the green algae ''Prototheca
'' and ''Helicosporidium
'', parasites of metazoans, or ''Cephaleuros
'', parasites of plants) were originally classified as fungi
s, or protist
ans of ''incertae sedis
'', while others (e.g., the green algae ''Phyllosiphon
'' and ''Rhodochytrium
'', parasites of plants, or the red algae ''Pterocladiophila
'' and ''Gelidiocolax mammillatus
'', parasites of other red algae, or the dinoflagellates ''Oodinium
'', parasites of fish) had their relationship with algae conjectured early. In other cases, some groups were originally characterized as parasitic algae (e.g., ''Chlorochytrium
''), but later were seen as endophytic
algae. Some filamentous bacteria (e.g., ''Beggiatoa
'') were originally seen as algae. Furthermore, groups like the apicomplexan
s are also parasites derived from ancestors that possessed plastids, but are not included in any group traditionally seen as algae.
Relationship to land plants
The first land plants probably evolved from shallow freshwater charophyte algae much like ''Chara
'' almost 500 million years ago. These probably had an isomorphic alternation of generations
and were probably filamentous. Fossils of isolated land plant spores suggest land plants may have been around as long as 475 million years ago.
A range of algal morphologies
is exhibited, and convergence
of features in unrelated groups is common. The only groups to exhibit three-dimensional multicellular thalli
are the reds
, and some chlorophytes
Apical growth is constrained to subsets of these groups: the florideophyte
reds, various browns, and the charophytes.
The form of charophytes is quite different from those of reds and browns, because they have distinct nodes, separated by internode 'stems'; whorls of branches reminiscent of the horsetail
s occur at the nodes.
s are another polyphyletic
trait; they appear in the coralline algae
and the Hildenbrandiales
, as well as the browns.
Most of the simpler algae are unicellular flagellate
s or amoeboid
s, but colonial and nonmotile forms have developed independently among several of the groups. Some of the more common organizational levels, more than one of which may occur in the lifecycle
of a species, are
: small, regular groups of motile cells
* Capsoid: individual non-motile cells embedded in mucilage
* Coccoid: individual non-motile cells with cell walls
* Palmelloid: nonmotile cells embedded in mucilage
* Filamentous: a string of nonmotile cells connected together, sometimes branching
* Parenchymatous: cells forming a thallus with partial differentiation of tissues
In three lines, even higher levels of organization have been reached, with full tissue differentiation. These are the brown algae,—some of which may reach 50 m in length (kelp
s)—the red algae, and the green algae. The most complex forms are found among the charophyte algae (see Charales
), in a lineage that eventually led to the higher land plants. The innovation that defines these nonalgal plants is the presence of female reproductive organs with protective cell layers that protect the zygote and developing embryo. Hence, the land plants are referred to as the Embryophyte
The term algal turf is commonly used but poorly defined. Algal turfs are thick, carpet-like beds of seaweed that retain sediment and compete with foundation species like corals
, and they are usually less than 15 cm tall. Such a turf may consist of one or more species, and will generally cover an area in the order of a square metre or more. Some common characteristics are listed:
* Algae that form aggregations that have been described as turfs include diatoms, cyanobacteria, chlorophytes, phaeophytes and rhodophytes. Turfs are often composed of numerous species at a wide range of spatial scales, but monospecific turfs are frequently reported.
* Turfs can be morphologically highly variable over geographic scales and even within species on local scales and can be difficult to identify in terms of the constituent species.
* Turfs have been defined as short algae, but this has been used to describe height ranges from less than 0.5 cm to more than 10 cm. In some regions, the descriptions approached heights which might be described as canopies (20 to 30 cm).
Many algae, particularly members of the Characeae
species, have served as model experimental organisms to understand the mechanisms of the water permeability of membranes, osmoregulation
, turgor regulation
, salt tolerance
, cytoplasmic streaming
, and the generation of action potentials
s are found not only in higher plants, but in algae, too.
Some species of algae form symbiotic relationships
with other organisms. In these symbioses, the algae supply photosynthates (organic substances) to the host organism providing protection to the algal cells. The host organism derives some or all of its energy requirements from the algae. Examples are:
s are defined by the International Association for Lichenology
to be "an association of a fungus and a photosynthetic symbiont
resulting in a stable vegetative body having a specific structure". The fungi, or mycobionts, are mainly from the Ascomycota
with a few from the Basidiomycota
. In nature they do not occur separate from lichens. It is unknown when they began to associate. One mycobiont associates with the same phycobiont species, rarely two, from the green algae, except that alternatively, the mycobiont may associate with a species of cyanobacteria (hence "photobiont" is the more accurate term). A photobiont may be associated with many different mycobionts or may live independently; accordingly, lichens are named and classified as fungal species. The association is termed a morphogenesis because the lichen has a form and capabilities not possessed by the symbiont species alone (they can be experimentally isolated). The photobiont possibly triggers otherwise latent genes in the mycobiont.
is an example of a common green alga genus worldwide that can grow on its own or be lichenised. Lichen thus share some of the habitat and often similar appearance with specialized species of algae (''aerophyte
s'') growing on exposed surfaces such as tree trunks and rocks and sometimes discoloring them.
s are accumulated from the calcareous exoskeleton
s of marine invertebrate
s of the order Scleractinia
s). These animals metabolize sugar
and oxygen to obtain energy for their cell-building processes, including secretion
of the exoskeleton, with water and carbon dioxide
as byproducts. Dinoflagellates (algal protists) are often endosymbiont
s in the cells of the coral-forming marine invertebrates, where they accelerate host-cell metabolism by generating sugar and oxygen immediately available through photosynthesis using incident light and the carbon dioxide produced by the host. Reef-building stony corals (hermatypic coral
s) require endosymbiotic algae from the genus ''Symbiodinium
'' to be in a healthy condition. The loss of ''Symbiodinium'' from the host is known as coral bleaching
, a condition which leads to the deterioration of a reef.
ic green algae live close to the surface of some sponges, for example, breadcrumb sponges (''Halichondria panicea
''). The alga is thus protected from predators; the sponge is provided with oxygen and sugars which can account for 50 to 80% of sponge growth in some species.
, and Heterokontophyta
, the three main algal divisions
, have lifecycles which show considerable variation and complexity. In general, an asexual phase exists where the seaweed's cells are diploid
, a sexual phase where the cells are haploid
, followed by fusion of the male and female gamete
s. Asexual reproduction permits efficient population increases, but less variation is possible. Commonly, in sexual reproduction of unicellular and colonial algae, two specialized, sexually compatible, haploid gametes make physical contact and fuse to form a zygote
. To ensure a successful mating, the development and release of gametes is highly synchronized and regulated; pheromones may play a key role in these processes. Sexual reproduction allows for more variation and provides the benefit of efficient recombinational repair of DNA damages during meiosis
, a key stage of the sexual cycle. However, sexual reproduction is more costly than asexual reproduction. Meiosis has been shown to occur in many different species of algae.
The ''Algal Collection of the US National Herbarium'' (located in the National Museum of Natural History
) consists of approximately 320,500 dried specimens, which, although not exhaustive (no exhaustive collection exists), gives an idea of the order of magnitude of the number of algal species (that number remains unknown). Estimates vary widely. For example, according to one standard textbook, in the British Isles
the ''UK Biodiversity Steering Group Report'' estimated there to be 20,000 algal species in the UK. Another checklist reports only about 5,000 species. Regarding the difference of about 15,000 species, the text concludes: "It will require many detailed field surveys before it is possible to provide a reliable estimate of the total number of species ..."
Regional and group estimates have been made, as well:
* 5,000–5,500 species of red algae worldwide
* "some 1,300 in Australian Seas"
* 400 seaweed species for the western coastline of South Africa, and 212 species from the coast of KwaZulu-Natal. Some of these are duplicates, as the range extends across both coasts, and the total recorded is probably about 500 species. Most of these are listed in List of seaweeds of South Africa
. These exclude phytoplankton
and crustose corallines.
* 669 marine species from California (US)
* 642 in the check-list of Britain and Ireland
and so on, but lacking any scientific basis or reliable sources, these numbers have no more credibility than the British ones mentioned above. Most estimates also omit microscopic algae, such as phytoplankton.
The most recent estimate suggests 72,500 algal species worldwide.
The distribution of algal species has been fairly well studied since the founding of phytogeography
in the mid-19th century.
Algae spread mainly by the dispersal of spore
s analogously to the dispersal of Plantae
. This dispersal can be accomplished by air, water, or other organisms. Due to this, spores can be found in a variety of environments: fresh and marine waters, air, soil, and in or on other organisms.
Whether a spore is to grow into an organism depends on the combination of the species and the environmental conditions where the spore lands.
The spores of freshwater algae are dispersed mainly by running water and wind, as well as by living carriers.
However, not all bodies of water can carry all species of algae, as the chemical composition of certain water bodies limits the algae that can survive within them.
Marine spores are often spread by ocean currents. Ocean water presents many vastly different habitats based on temperature and nutrient availability, resulting in phytogeographic zones, regions, and provinces.
To some degree, the distribution of algae is subject to floristic discontinuities caused by geographical features, such as Antarctica
, long distances of ocean or general land masses. It is, therefore, possible to identify species occurring by locality, such as "Pacific
algae" or "North Sea
algae". When they occur out of their localities, hypothesizing a transport mechanism is usually possible, such as the hulls of ships. For example, ''Ulva reticulata
'' and ''U. fasciata
'' travelled from the mainland to Hawaii
in this manner.
Mapping is possible for select species only: "there are many valid examples of confined distribution patterns." For example, ''Clathromorphum
'' is an arctic genus and is not mapped far south of there. However, scientists regard the overall data as insufficient due to the "difficulties of undertaking such studies."
Algae are prominent in bodies of water, common in terrestrial environments, and are found in unusual environments, such as on snow
. Seaweeds grow mostly in shallow marine waters, under deep; however, some such as ''Navicula
pennata'' have been recorded to a depth of . A type of algae, ''Ancylonema nordenskioeldii'', was found in Greenland
in areas known as the 'Dark Zone', which caused an increase in the rate of melting ice sheet. Same algae was found in the Italian Alps
, after pink ice appeared on parts of the Presena glacier.
The various sorts of algae play significant roles in aquatic ecology. Microscopic forms that live suspended in the water column (phytoplankton
) provide the food base for most marine food chain
s. In very high densities (algal bloom
s), these algae may discolor the water and outcompete, poison, or asphyxiate
other life forms.
Algae can be used as indicator organism
s to monitor pollution in various aquatic systems.
In many cases, algal metabolism is sensitive to various pollutants. Due to this, the species composition of algal populations may shift in the presence of chemical pollutants.
To detect these changes, algae can be sampled from the environment and maintained in laboratories with relative ease.
On the basis of their habitat, algae can be categorized as: aquatic
, endozoic), parasitic
In classical Chinese
, the word is used both for "algae" and (in the modest tradition of the imperial scholars
) for "literary talent". The third island in Kunming Lake
beside the Summer Palace
in Beijing is known as the Zaojian Tang Dao, which thus simultaneously means "Island of the Algae-Viewing Hall" and "Island of the Hall for Reflecting on Literary Talent".
, a gelatin
ous substance derived from red algae, has a number of commercial uses. It is a good medium on which to grow bacteria and fungi, as most microorganisms cannot digest agar.
, or alginate, is extracted from brown algae
. Its uses range from gelling agents in food, to medical dressings. Alginic acid also has been used in the field of biotechnology
as a biocompatible medium
for cell encapsulation and cell immobilization. Molecular cuisine
is also a user of the substance for its gelling properties, by which it becomes a delivery vehicle for flavours.
Between 100,000 and 170,000 wet tons of ''Macrocystis
'' are harvested annually in New Mexico
extraction and abalone
To be competitive and independent from fluctuating support from (local) policy on the long run, biofuels should equal or beat the cost level of fossil fuels. Here, algae-based fuels hold great promise, directly related to the potential to produce more biomass per unit area in a year than any other form of biomass. The break-even point for algae-based biofuels is estimated to occur by 2025.
For centuries, seaweed has been used as a fertilizer; George Owen of Henllys
writing in the 16th century referring to drift weed in South Wales
Today, algae are used by humans in many ways; for example, as fertilizer
s, soil conditioner
s, and livestock feed. Aquatic and microscopic species are cultured in clear tanks or ponds and are either harvested or used to treat effluents pumped through the ponds. Algaculture
on a large scale is an important type of aquaculture
in some places. Maerl
is commonly used as a soil conditioner.
Naturally growing seaweeds are an important source of food, especially in Asia leading some to label it as a superfood
. They provide many vitamins including: A, B1
, and C
, and are rich in iodine
, iron, magnesium
, and calcium
. In addition, commercially cultivated microalgae, including both algae and cyanobacteria, are marketed as nutritional supplements, such as spirulina
'' and the vitamin-C supplement from ''Dunaliella
'', high in beta-carotene
Algae are national foods of many nations: China consumes more than 70 species, including ''fat choy
'', a cyanobacterium considered a vegetable; Japan, over 20 species such as ''nori
'' and ''aonori
''; Ireland, dulse
is used to make laver bread in Wales
, where it is known as ; in Korea
, . It is also used along the west coast of North America from California to British Columbia
, in Hawaii and by the Māori
of New Zealand
. Sea lettuce
are salad ingredients in Scotland
, Ireland, Greenland
, and Iceland
. Algae is being considered a potential solution for world hunger problem.
There exist 2 popular forms of Algae which are used in cuisine:
: This form of Algae is found in freshwater and contains photosynthetic
pigments in its chloroplast
. It is high in iron
, vitamin B2
and Omega-3 Fatty acids
Furthermore, it contains all nine of the essential amino acids
the body does not produce on its own
: Known otherwise as cyanobacteria
, this is a form of blue-green algae that is filled with nutrients and contains 10% more protein
than Chlorella as well as more thiamine
The oils from some algae have high levels of unsaturated fatty acid
s. For example, ''Parietochloris incisa
'' is very high in arachidonic acid
, where it reaches up to 47% of the triglyceride pool. Some varieties of algae favored by vegetarianism
contain the long-chain, essential omega-3 fatty acid
s, docosahexaenoic acid
(DHA) and eicosapentaenoic acid
(EPA). Fish oil contains the omega-3 fatty acids, but the original source is algae (microalgae in particular), which are eaten by marine life such as copepod
s and are passed up the food chain. Algae have emerged in recent years as a popular source of omega-3 fatty acids for vegetarians who cannot get long-chain EPA and DHA from other vegetarian sources such as flaxseed oil
, which only contains the short-chain alpha-linolenic acid
* Sewage can be treated with algae, reducing the use of large amounts of toxic chemicals that would otherwise be needed.
* Algae can be used to capture fertilizers in runoff from farms. When subsequently harvested, the enriched algae can be used as fertilizer.
* Aquaria and ponds can be filtered using algae, which absorb nutrients from the water in a device called an algae scrubber
, also known as an algae turf scrubber.
Agricultural Research Service
scientists found that 60–90% of nitrogen runoff and 70–100% of phosphorus runoff can be captured from manure effluents
using a horizontal algae scrubber, also called an algal turf scrubber
(ATS). Scientists developed the ATS, which consists of shallow, 100-foot raceways of nylon netting where algae colonies can form, and studied its efficacy for three years. They found that algae can readily be used to reduce the nutrient runoff from agricultural fields and increase the quality of water flowing into rivers, streams, and oceans. Researchers collected and dried the nutrient-rich algae from the ATS and studied its potential as an organic fertilizer. They found that cucumber and corn seedlings grew just as well using ATS organic fertilizer as they did with commercial fertilizers. Algae scrubbers, using bubbling upflow or vertical waterfall versions, are now also being used to filter aquaria and ponds.
Various polymers can be created from algae, which can be especially useful in the creation of bioplastics. These include hybrid plastics, cellulose based plastics, poly-lactic acid, and bio-polyethylene. Several companies have begun to produce algae polymers commercially, including for use in flip-flops and in surf boards.
The alga ''Stichococcus bacillaris
'' has been seen to colonize silicone resins used at archaeological sites; biodegrading
the synthetic substance.
The natural pigment
s and chlorophyll
s) produced by algae can be used as alternatives to chemical dye
s and coloring agents.
The presence of some individual algal pigments, together with specific pigment concentration ratios, are taxon-specific: analysis of their concentrations with various analytical methods, particularly high-performance liquid chromatography
, can therefore offer deep insight into the taxonomic composition and relative abundance of natural algae populations in sea water samples.
Carrageenan, from the red alga ''Chondrus crispus'', is used as a stabilizer in milk products.
File:Algae bladder 4290.jpg|Algae bladder
* Iron fertilization
Britain and Ireland
* – a database of all algal names including images, nomenclature, taxonomy, distribution, bibliography, uses, extracts