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Pulmonata or pulmonates, is an informal group (previously an
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
, and before that a subclass) of
snails A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastrop ...
and
slug Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a smal ...
s characterized by the ability to breathe air, by virtue of having a pallial lung instead of a
gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
, or gills. The group includes many land and freshwater families, and several marine families. The
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
Pulmonata as traditionally defined was found to be polyphyletic in a molecular study per Jörger ''et al.'', dating from 2010. Pulmonata are known from the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
period to the present. Pulmonates have a single atrium and kidney, and a concentrated symmetrical nervous system. The mantle cavity is on the right side of the body, and lacks gills, instead being converted into a vascularised lung. Most species have a shell, but no operculum, although the group does also include several shell-less slugs. Pulmonates are hermaphroditic, and some groups possess love darts.


Linnean taxonomy

The taxonomy of this group according to the
taxonomy of the Gastropoda (Ponder & Lindberg, 1997) The taxonomy of the Gastropoda, as revised by Winston Ponder and David R. Lindberg in 1997, is an older taxonomy of the class Gastropoda, the class of molluscs consisting of all snails and slugs. The full name of the work in which this taxonomy wa ...
was as follows: Order Pulmonata
Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in nat ...
in Blainville, 1814 - pulmonates *Suborder
Systellommatophora The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005). They are marine and terrestrial pulmonate gastropods within the Heterobranchi ...
Pilsbry, 1948 ***Superfamily Onchidioidea Rafinesque, 1815 ***Superfamily
Otinoidea Otinoidea is a superfamily of very small, air-breathing sea snails and sea slugs, marine pulmonate gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known ...
H. & A. Adams, 1855 ***Superfamily
Rathouisioidea Veronicelloidea is a superfamily of air-breathing land slugs. They are terrestrial pulmonate gastropod molluscs in the clade Systellommatophora. Taxonomy The following two families were recognized in the taxonomy of Bouchet & Rocroi (2005): * ...
Sarasin, 1889 *Suborder
Basommatophora Basommatophora was a term that was previously used as a taxonomic informal group, a group of snails within the informal group Pulmonata, the air-breathing slugs and snails. According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), w ...
Keferstein in Bronn, 1864 - freshwater pulmonates, pond snails ***Superfamily Acroloxoidea Thiele, 1931 ***Superfamily
Amphiboloidea Amphiboloidea is a Taxonomy (biology), taxonomic superfamily of air-breathing land snails. Distribution Amphibolids are found in Indo-Pacific intertidal mangrove, saltmarsh and estuarine mudflat habitats. Taxonomy 2005 taxonomy According ...
J.E. Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for ...
, 1840 ***Superfamily
Chilinoidea Chilinoidea is a family (biology), superfamily of air-breathing freshwater snails, pulmonate gastropod mollusks in the clade Hygrophila (Gastropoda), Hygrophila. References

Panpulmonata {{Pulmonata-stub ...
H. & A. Adams, 1855 ***Superfamily
Glacidorboidea The Glacidorbidae is a taxonomic family of freshwater snails. Taxonomy 2005 taxonomy Glacidorbidae is the only family in the superfamily Glacidorboidea. Glacidorboidea has been classified within the informal group Lower Heterobranchia in the ...
Ponder, 1986 ***Superfamily
Lymnaeoidea Lymnaeoidea, common name the pond snails, is a taxonomic superfamily of small to large air-breathing freshwater snails, aquatic pulmonate gastropod mollusks, that belong to the superorder Hygrophila.MolluscaBase eds. (2020). MolluscaBase. Lymna ...
Rafinesque, 1815 ***Superfamily
Planorboidea Planorboidea is a superfamily of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks. All of the gastropods in this superfamily are sinistral in shell coiling. The monophyly of Planorboidea was confirmed by Albrecht et al. (2 ...
Rafinesque, 1815 ***Superfamily
Siphonarioidea Siphonarioidea is a taxonomic superfamily of air-breathing sea snails or false limpets, marine gastropod mollusks in the clade Panpulmonata. Distribution The Siphonarioidea are broadly distributed across the globe, however they are known to be ...
J.E. Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for ...
, 1840 *Suborder
Eupulmonata Eupulmonata is a taxonomic clade of air-breathing snails. The great majority of this group are land snails and slugs, but some are marine and some are saltmarsh snails that can tolerate salty conditions. Linnean taxonomy *Suborder Eupulmonat ...
Haszprunar & Huber, 1990 **Infraorder Acteophila Dall, 1885 = formerly Archaeopulmonata ***Superfamily Melampoidea Stimpson, 1851 **Infraorder Trimusculiformes Minichev & Starobogatov, 1975 ***Superfamily
Trimusculoidea ''Trimusculus'' is a genus of medium-sized air-breathing sea snails or false limpets, marine pulmonate gastropod molluscs in the family Trimusculidae. ''Trimusculus'' is the only genus in the family Trimusculidae. Trimusculidae, the button sna ...
Zilch, 1959 **Infraorder
Stylommatophora Stylommatophora is an orderPhilippe Bouchet, Jean-Pierre Rocroi, Bernhard Hausdorf, Andrzej Kaim, Yasunori Kano, Alexander Nützel, Pavel Parkhaev, Michael Schrödl and Ellen E. Strong. 2017. Revised Classification, Nomenclator and Typification ...
A. Schmidt, 1856 - land snails ***Subinfraorder
Orthurethra Orthurethra is a clade of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora. In the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005, Orthurethra is treated as an informal group and subcla ...
***Superfamily Achatinelloidea Gulick, 1873 ***Superfamily Cochlicopoidea Pilsbry, 1900 ***Superfamily Partuloidea Pilsbry, 1900 ***Superfamily Pupilloidea Turton, 1831 ***Subinfraorder
Sigmurethra Sigmurethra is a taxonomic category of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This is an informal group which includes most land snails and slugs. The two strong synapomorphies of Sigmurethra are a ...
***Superfamily
Acavoidea The Acavoidea are a taxonomic superfamily of air-breathing land snails and slugs, terrestrial pulmonate gastropod mollusks in the informal group Sigmurethra. This taxonomy was based on the study by Nordsieck, published in 1986. Taxonomy ...
Pilsbry, 1895 ***Superfamily
Achatinoidea The Achatinoidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod mollusks in the informal group Sigmurethra. 2005 taxonomy According to taxonomy of the Gastropoda by Bouchet & Rocroi, 2005 there are four fami ...
Swainson, 1840 ***Superfamily
Aillyoidea Aillyidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the informal group Sigmurethra. Aillyidae is the only family in the superfamily Aillyoidea. This family and the genus is named in honor of Swedish m ...
Baker, 1960 ***Superfamily
Arionoidea Arionoidea is a taxonomic group, superfamily of air-breathing land slugs, shell-less terrestrial pulmonate gastropod mollusks. Families Families within the superfamily Arionoidea include: * Arionidae * Anadenidae * Ariolimacidae * Bin ...
J.E. Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for ...
in Turnton, 1840 ***Superfamily Buliminoidea Clessin, 1879 ***Superfamily Camaenoidea Pilsbry, 1895 ***Superfamily
Clausilioidea The Clausilioidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod mollusks in the informal group Sigmurethra. This classification was based on different studies dealing with the Clausiliidae, published by H. Nords ...
Mörch, 1864 ***Superfamily Dyakioidea Gude & Woodward, 1921 ***Superfamily
Gastrodontoidea Gastrodontoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the limacoid clade. Taxonomy According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), families in this superfamily ...
Tryon, 1866 ***Superfamily
Helicoidea Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora. Taxonomy 2005 taxonomy There are 19 families within the superfamily Helicoidea according to the ta ...
Rafinesque, 1815 ***Superfamily
Helixarionoidea Helicarionoidea is a superfamily of air-breathing land snails and semi-slugs, terrestrial pulmonate gastropod mollusks in the infraorder Limacoidei. Families There are three families within the superfamily Helicarionoidea: * Helicarionidae * Ar ...
Bourguignat, 1877 ***Superfamily
Limacoidea Limacoidea is a taxonomic superfamily of medium-sized to large, air-breathing land slugs and snails. They are terrestrial pulmonate gastropod mollusks in the infraorder Stylommatophora (according to the taxonomy of the Gastropoda by Bouchet ...
Rafinesque, 1815 ***Superfamily
Oleacinoidea The Oleacinoidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod mollusks in the suborder Helicina of the order Stylommatophora. MolluscaBase eds. (2020). MolluscaBase. Oleacinoidea H. Adams & A. Adams, 1855. Acces ...
H. & A. Adams, 1855 ***Superfamily Orthalicoidea Albers-Martens, 1860 ***Superfamily Plectopylidoidea Moellendorf, 1900 ***Superfamily Polygyroidea Pilsbry, 1894 ***Superfamily Punctoidea Morse, 1864 ***Superfamily Rhytidoidea Pilsbry, 1893 ***Superfamily Sagdidoidera Pilsbry, 1895 ***Superfamily Staffordioidea Thiele, 1931 ***Superfamily
Streptaxoidea Streptaxoidea is a Taxonomic rank, superfamily of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the suborder Achatinina of the order Stylommatophora. Taxonomy 2005 taxonomy There have been reco ...
J.E. Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for ...
, 1806 ***Superfamily Strophocheiloidea Thiele, 1926 ***Superfamily Trigonochlamydoidea Hese, 1882 ***Superfamily Zonitoidea Mörch, 1864 ***? Superfamily
Athoracophoroidea Athoracophoridae, common name the leaf-veined slugs, are a family (biology), family of air-breathing land slugs, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the infraorder Stylommatophora, the stalk-eyed snails and slugs. Ma ...
P. Fischer, 1883 = Tracheopulmonata ***? Superfamily
Succineoidea Succineoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the suborder Helicina. MolluscaBase eds. (2021). MolluscaBase. Succineoidea Beck, 1837. Accessed through: World Register of Marine ...
Beck, 1837 = Heterurethra


2005 taxonomy

The taxonomy of this group according to the
taxonomy of the Gastropoda (Bouchet & Rocroi, 2005) The taxonomy of the Gastropoda as it was revised in 2005 by Philippe Bouchet and Jean-Pierre Rocroi is a system for the scientific classification of gastropod mollusks. (Gastropods are a taxonomic class of animals which consists of snails and ...
was as follows:


Informal Group Pulmonata

Contains the informal group Basommatophora and the clade Eupulmonata


Informal Group Basommatophora

Contains the clade Hygrophila *Superfamily
Amphiboloidea Amphiboloidea is a Taxonomy (biology), taxonomic superfamily of air-breathing land snails. Distribution Amphibolids are found in Indo-Pacific intertidal mangrove, saltmarsh and estuarine mudflat habitats. Taxonomy 2005 taxonomy According ...
**Family
Amphibolidae Amphibolidae is a family of air-breathing snails with opercula, terrestrial pulmonate gastropod molluscs. This family of pulmonate gastropods, for breathe air, but also have opercula and at least some species go through a free-swimming velig ...
*Superfamily
Siphonarioidea Siphonarioidea is a taxonomic superfamily of air-breathing sea snails or false limpets, marine gastropod mollusks in the clade Panpulmonata. Distribution The Siphonarioidea are broadly distributed across the globe, however they are known to be ...
**Family
Siphonariidae Siphonariidae, also known as false limpets, are a taxonomic family of small to medium-sized air-breathing sea snails, marine and brackish water pulmonate gastropod molluscs. __TOC__ Shell description Empty siphonariid limpet shells can b ...
** † Family
Acroreiidae † Acroreiidae is an extinct family of air-breathing snails, pulmonate gastropod mollusks in the superfamily Siphonariidae. The living siphonariids are all marine pulmonates, false sea limpets that breathe air. Taxonomy The family Acroreiidae ...


= Clade Hygrophila

= *Superfamily
Chilinoidea Chilinoidea is a family (biology), superfamily of air-breathing freshwater snails, pulmonate gastropod mollusks in the clade Hygrophila (Gastropoda), Hygrophila. References

Panpulmonata {{Pulmonata-stub ...
**Family
Chilinidae ''Chilina'' is a genus of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks in the superfamily Chilinoidea. ''Chilina'' is the only genus in the family Chilinidae. Distribution Chilinidae occupies the temperate and c ...
**Family
Latiidae ''Latia'' is a genus of very small, air-breathing freshwater snails or limpets, aquatic pulmonate gastropod molluscs in the superfamily Chilinoidea. ''Latia'' is the only genus in the family Latiidae. Species in this genus are the only ...
*Superfamily Acroloxoidea **Family
Acroloxidae Acroloxidae, commonly known as river limpets, are a taxonomic family of very small, freshwater snails, aquatic pulmonate gastropod limpet-like mollusks with a simple flattened conical shell in the clade Hygrophila. Acroloxidae is the only fam ...
*Superfamily
Lymnaeoidea Lymnaeoidea, common name the pond snails, is a taxonomic superfamily of small to large air-breathing freshwater snails, aquatic pulmonate gastropod mollusks, that belong to the superorder Hygrophila.MolluscaBase eds. (2020). MolluscaBase. Lymna ...
**Family
Lymnaeidae Lymnaeidae, common name the pond snails, is a taxonomic family of small to large air-breathing freshwater snails, aquatic pulmonate gastropod mollusks, that belong to the clade Hygrophila. Lymnaeidae is the only family within the superfamily ...
*Superfamily
Planorboidea Planorboidea is a superfamily of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks. All of the gastropods in this superfamily are sinistral in shell coiling. The monophyly of Planorboidea was confirmed by Albrecht et al. (2 ...
**Family Planorbidae **Family
Physidae Physidae, common name the bladder snails, is a monophyletic taxonomic family of small air-breathing freshwater snails, aquatic pulmonate gastropod molluscs in the superfamily Lymnaeoidea .MolluscaBase eds. (2021). MolluscaBase. Physidae Fitzin ...


Clade Eupulmonata

Contains the clades Systellommatophora and Stylommatophora *Superfamily
Trimusculoidea ''Trimusculus'' is a genus of medium-sized air-breathing sea snails or false limpets, marine pulmonate gastropod molluscs in the family Trimusculidae. ''Trimusculus'' is the only genus in the family Trimusculidae. Trimusculidae, the button sna ...
**Family
Trimusculidae ''Trimusculus'' is a genus of medium-sized air-breathing sea snails or false limpets, marine pulmonate gastropod molluscs in the family Trimusculidae. ''Trimusculus'' is the only genus in the family Trimusculidae. Trimusculidae, the button s ...
*Superfamily
Otinoidea Otinoidea is a superfamily of very small, air-breathing sea snails and sea slugs, marine pulmonate gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known ...
**Family Otinidae **Family
Smeagolidae ''Smeagol'' is a genus of small air-breathing sea slugs of the upper intertidal zone.Marshall, B.; Rosenberg, G. (2016). Smeagol Climo, 1980. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecie ...
*Superfamily
Ellobioidea Ellobiidae, common name the hollow-shelled snails, is a family of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Eupulmonata.Bouchet, P. (2012). Ellobiidae. Accessed through: World Register of Marine Sp ...
**Family Ellobiidae


= Clade

Systellommatophora The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005). They are marine and terrestrial pulmonate gastropods within the Heterobranchi ...
( =
Gymnomorpha The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005). They are marine and terrestrial pulmonate gastropods within the Heterobranchia. Th ...
)

= *Superfamily Onchidioidea **Family
Onchidiidae Onchidiidae are a family of small, air-breathing sea (and land) slugs. They are shell-less marine (except for 2 species)preview pulmonate gastropod molluscs. Onchidiidae is the only family within the superfamily Onchidioidea. These animals are ...
*Superfamily
Veronicelloidea Veronicelloidea is a superfamily of air-breathing land slugs. They are terrestrial pulmonate gastropod molluscs in the clade Systellommatophora. Taxonomy The following two families were recognized in the taxonomy of Bouchet & Rocroi (20 ...
**Family
Veronicellidae The Veronicellidae, also known by their common name the leatherleaf slugs, are a family of pulmonate terrestrial slugs. The herbivorous molluscs occur mainly in the tropical and subtropical areas of America, Asia and Africa. They act as intermed ...
**Family
Rathouisiidae Rathouisiidae is a family of carnivorous air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Veronicelloidea, the leatherleaf slugs and their allies. This family has no subfamilies according to the ''Taxonomy of ...


= Clade

Stylommatophora Stylommatophora is an orderPhilippe Bouchet, Jean-Pierre Rocroi, Bernhard Hausdorf, Andrzej Kaim, Yasunori Kano, Alexander Nützel, Pavel Parkhaev, Michael Schrödl and Ellen E. Strong. 2017. Revised Classification, Nomenclator and Typification ...

= Contains the subclades Elasmognatha, Orthurethra and the informal group Sigmurethra


= Subclade Elasmognatha

= *Superfamily
Succineoidea Succineoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the suborder Helicina. MolluscaBase eds. (2021). MolluscaBase. Succineoidea Beck, 1837. Accessed through: World Register of Marine ...
**Family
Succineidae Succineidae are a family of small to medium-sized, air-breathing land snails (and slugs), terrestrial pulmonate gastropod molluscs in the superfamily Succineoidea.MolluscaBase eds. (2021). MolluscaBase. Succineidae Beck, 1837. Accessed thr ...
*Superfamily
Athoracophoroidea Athoracophoridae, common name the leaf-veined slugs, are a family (biology), family of air-breathing land slugs, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the infraorder Stylommatophora, the stalk-eyed snails and slugs. Ma ...
**Family Athoracophoridae


= Subclade

Orthurethra Orthurethra is a clade of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora. In the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005, Orthurethra is treated as an informal group and subcla ...

= *Superfamily Partuloidea **Family
Partulidae Partulidae is a family of air-breathing land snails, terrestrial gastropod mollusks in the superfamily Pupilloidea.MolluscaBase eds. (2021). MolluscaBase. Partulidae Pilsbry, 1900. Accessed through: World Register of Marine Species at: http:// ...
**Family
Draparnaudiidae ''Draparnaudia'' is a genus of air-breathing land snails, Terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Partuloidea. ''Draparnaudia'' is the only genus in the family Draparnaudiidae. Both the family name an ...
*Superfamily Achatinelloidea **Family Achatinellidae *Superfamily Cochlicopoidea **Family
Cochlicopidae Cochlicopidae is a taxonomic family of small, air-breathing, land snails, terrestrial pulmonate gastropod mollusks in the superfamily Cochlicopoidea. Anatomy In this family, the number of haploid chromosome A chromosome is a long DN ...
**Family
Amastridae Amastridae is a taxonomic family of small, air-breathing, land snails, terrestrial pulmonate gastropod mollusks in the superfamily Pupilloidea. MolluscaBase eds. (2021). MolluscaBase. Amastridae Pilsbry, 1910. Accessed through: World Register ...
*Superfamily Pupilloidea **Family Pupillidae **Family
Argnidae Argnidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks. Distribution Distribution of Argnidae include Europe: southern and eastern Alps and Carpathian Mountains.
**Family
Chondrinidae Chondrinidae is a family of mostly minute air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Pulmonata. Anatomy In this family, the number of haploid chromosomes lies between 26 and 30 (according to the values i ...
** † Family
Cylindrellinidae †Cylindrellinidae is an extinct family (biology), family of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks.MolluscaBase eds. (2022). MolluscaBase. Cylindrellinidae Zilch, 1959 †. Accessed through: Wo ...
**Family
Lauriidae Lauriidae is a family of land snails. It is classified within the informal group Orthurethra, itself belonging to the clade Stylommatophora within the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005) ...
**Family
Orculidae Orculidae is a family of mostly minute, air-breathing, land snails; terrestrial pulmonate gastropod mollusks in the superfamily Pupilloidea. MolluscaBase eds. (2021). MolluscaBase. Orculidae Pilsbry, 1918. Accessed through: World Register of Mar ...
**Family
Pleurodiscidae Pleurodiscus is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Pupilloidea. ''Pleurodiscus'' is the only genus in the family Pleurodiscidae. Species Species within the genus ''Pleurodiscus'' ...
**Family
Pyramidulidae Pyramidulidae is a family (biology), family of small air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Pupilloidea. MolluscaBase eds. (2022). MolluscaBase. Pyramidulidae Kennard & B. B. W ...
**Family Spelaeodiscidae **Family Strobilopsidae **Family
Valloniidae Valloniidae is a taxonomic family of small and minute, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Pupilloidea. MolluscaBase eds. (2020). MolluscaBase. Valloniidae Morse, 1864. Accessed through: World R ...
**Family Vertiginidae *Superfamily
Enoidea Pupilloidea is a superfamily of small and very small air-breathing land snails, terrestrial gastropod mollusks in the infraorder '' Pupilloidei'' . Taxonomy This superfamily contains the following families: * Achatinellidae Gulick, 1873 * Agard ...
**Family
Enidae Enidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks. MolluscaBase eds. (2020). MolluscaBase. Enidae B. B. Woodward, 1903 (1880). Accessed through: World Register of Marine Species at: http://www.marinespecies ...
**Family
Cerastidae Cerastidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Stylommatophora.MolluscaBase eds. (2020). MolluscaBase. Cerastidae Wenz, 1923. Accessed through: World Register of Marine Species at: http ...


= Informal Group

Sigmurethra Sigmurethra is a taxonomic category of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This is an informal group which includes most land snails and slugs. The two strong synapomorphies of Sigmurethra are a ...

= *Superfamily
Clausilioidea The Clausilioidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod mollusks in the informal group Sigmurethra. This classification was based on different studies dealing with the Clausiliidae, published by H. Nords ...
**Family Clausiliidae ** † Family
Anadromidae Anadromidae is an extinct family of gastropods in the clade Eupulmonata (according to taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Taxonomy The family Anadromidae is classified within the informal group Sigmurethra, itself belonging ...
** † Family
Filholiidae †Filholiidae is an extinct family of fossil air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Clausilioidea The Clausilioidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod ...
** † Family
Palaeostoidae Palaeostoidae is an extinct family of air-breathing land snails, terrestrial gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks ...
*Superfamily Orthalicoidea **Family
Orthalicidae Orthalicidae (orthalicid land snails) are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the subfamily Orthalicoidea of the order Stylommatophora. MolluscaBase eds. (2021). MolluscaBase ...
**Family
Cerionidae Cerionidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea. Pre-2008 taxonomy According to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005), the family Cerionidae is ...
**Family Coelociontidae ** † Family
Grangerellidae Grangerellidae is an extinct family of gastropods in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). This family has no subfamilies. Genera Genera within the family Grangerellidae include: ...
**Family
Megaspiridae Megaspiridae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).Breure A. S. H. & Romero P. (2012). "Supp ...
**Family
Placostylidae Bothriembryontidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Orthalicoidea. Taxonomy This family has been named Placostylidae and had no subfamilies (according to the taxonomy of th ...
**Family
Urocoptidae Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea. Taxonomy 2005 taxonomy The family Urocoptidae was classified in the superfamily Orthalicoidea (according to ...
*Superfamily
Achatinoidea The Achatinoidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod mollusks in the informal group Sigmurethra. 2005 taxonomy According to taxonomy of the Gastropoda by Bouchet & Rocroi, 2005 there are four fami ...
**Family Achatinidae **Family
Ferussaciidae Ferussaciidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Achatinoidea. MolluscaBase eds. (2020). MolluscaBase. Ferussaciidae Bourguignat, 1883. Accessed through: World Register of Marine ...
**Family
Micractaeonidae ''Micractaeon koptawelilensis'' is a species of land snail, terrestrial gastropod mollusks in the superfamily Achatinoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). ''Micractaeon koptawelilensis'' is the only s ...
**Family
Subulinidae Subulininae is a subfamily of small tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Achatinidae. Distribution Worldwide.
*Superfamily
Aillyoidea Aillyidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the informal group Sigmurethra. Aillyidae is the only family in the superfamily Aillyoidea. This family and the genus is named in honor of Swedish m ...
**Family
Aillyidae Aillyidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the informal group Sigmurethra. Aillyidae is the only family in the superfamily Aillyoidea. This family and the genus is named in honor of Swedish m ...
*Superfamily
Testacelloidea The Testacelloidea are a superfamily of air-breathing land snails and slugs, terrestrial gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as mollusc ...
**Family Testacellidae **Family
Oleacinidae Oleacinidae is a Taxonomy (biology), taxonomic family (biology), family of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the clade Eupulmonata (according to the Taxonomy of the Gastropoda (Bouchet & ...
**Family
Spiraxidae Spiraxidae is a family of predatory air-breathing land snails, terrestrial pulmonate gastropods in the superfamily Testacelloidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Distribution The distribution of th ...
*Superfamily
Papillodermatoidea ''Papilloderma altonagai'' is a species of air-breathing land slug, a terrestrial pulmonate gastropod mollusk in the informal group Sigmurethra. ''Papilloderma altonagai'' is the only species in the genus ''Papilloderma'', which is the only genu ...
**Family
Papillodermatidae ''Papilloderma altonagai'' is a species of air-breathing land slug, a terrestrial molluscs, terrestrial pulmonate gastropod mollusk in the informal group Sigmurethra. ''Papilloderma altonagai'' is the only species in the genus ''Papilloderma'', ...
*Superfamily
Streptaxoidea Streptaxoidea is a Taxonomic rank, superfamily of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the suborder Achatinina of the order Stylommatophora. Taxonomy 2005 taxonomy There have been reco ...
**Family
Streptaxidae Streptaxidae is a family of carnivorous air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora. Six Streptaxidae subfamilies are accepted in the 2005 taxonomy of the Gastropoda by Bouchet & Rocroi. Stre ...
*Superfamily Rhytidoidea **Family Rhytididae **Family
Chlamydephoridae ''Chlamydephorus'' is a genus of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the family Chlamydephoridae. It is the only genus within the family Chlamydephoridae.Herbert D. G. (1997). "The terrestrial slugs of KwaZulu- ...
**Family Haplotrematidae **Family
Scolodontidae Scolodontidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or ...
*Superfamily
Acavoidea The Acavoidea are a taxonomic superfamily of air-breathing land snails and slugs, terrestrial pulmonate gastropod mollusks in the informal group Sigmurethra. This taxonomy was based on the study by Nordsieck, published in 1986. Taxonomy ...
**Family Acavidae **Family
Caryodidae Caryodidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Acavoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). This family is endemic to eastern ...
**Family
Dorcasiidae Dorcasiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Acavoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). The family Dorcasiidae has no subfamilies. Di ...
**Family
Macrocyclidae Macrocyclidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Acavoidea The Acavoidea are a taxonomic superfamily of air-breathing land snails and slugs, terrestrial pulmonate gastrop ...
**Family
Megomphicidae Megomphicidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Acavoidea The Acavoidea are a taxonomic superfamily of air-breathing land snails and slugs, terrestrial pulmonat ...
**Family
Strophocheilidae Strophocheilidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Acavoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Fossil record The genus '' E ...
*Superfamily Punctoidea **Family
Punctidae Punctidae, common name the "dot snails" or "pinhead snails", is a family of minute air-breathing land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the informal group Sigmurethra (according to the taxonomy of the Gastropo ...
** † Family Anastomopsidae **Family Charopidae **Family
Cystopeltidae Cystopeltidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Punctoidea Punctoidea is a superfamily of air-breathing land snails and slugs, terrestrial pulmonate gastropod mollusks in t ...
**Family
Discidae Discidae is a taxonomic family of small air-breathing land snails, terrestrial gastropod mollusks in the superfamily Punctoidea. Genera The family Discidae has no subfamilies. Genera within the family Discidae include: * ''Anguispira'' Morse, ...
**Family Endodontidae **Family
Helicodiscidae Helicodiscidae is a family of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Punctoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Genera Genera within the family i ...
**Family
Oreohelicidae Oreohelicidae is a family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Punctoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Anatomy In this famil ...
**Family Thyrophorellidae *Superfamily
Sagdoidea Sagdoidea is a superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the infraorder Helicoidei of the suborder Helicina. MolluscaBase eds. (2021). MolluscaBase. Sagdoidea Pilsbry, 1895. Accessed through: World Re ...
**Family
Sagdidae Sagdidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks with highest diversity in the Greater Antilles. It has been classified in its own superfamily Sagdoidea and as a member of the superfamily Helicoidea. So ...


=

limacoid clade The Limacoidei is a taxonomic infraorder of air-breathing land snails, semislugs and slugs, terrestrial pulmonate gastropod molluscs in the suborder Helicina Distribution The original ancestral area of limacoid families is thought to be the ...

= *Superfamily Staffordioidea **Family
Staffordiidae Staffordiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea. MolluscaBase eds. (2021). MolluscaBase. Staffordiidae Thiele, 1931. Accessed through: World Register of Mar ...
*Superfamily Dyakioidea **Family
Dyakiidae Dyakiidae is a family of air-breathing land snails terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Some of the species in this family are sin ...
*Superfamily
Gastrodontoidea Gastrodontoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the limacoid clade. Taxonomy According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), families in this superfamily ...
**Family
Gastrodontidae Gastrodontidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Gastrodontoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). MolluscaBase eds. (2020). MolluscaBase ...
**Family
Chronidae Chronidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea within the superorder Eupulmonata. MolluscaBase eds. (2021). MolluscaBase. Chronidae Thiele, 1931. Accessed through: ...
**Family
Euconulidae Euconulidae is a taxonomic family of minute, air-breathing land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the superfamily Trochomorphoidea. This land snail family is closely allied to the Zonitidae, the glass snails. ...
**Family
Oxychilidae Oxychilidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Gastrodontoidea. Distribution The distribution of Oxychilidae includes the Nearctic, western-Palearctic, eastern-Palear ...
**Family
Pristilomatidae Pristilomatidae is a taxonomic family of air-breathing land snails, terrestrial gastropod mollusks in the superfamily Gastrodontoidea. According to the 2005 taxonomy of the Gastropoda by Bouchet & Rocroi these snails belong to the "limacoid ...
**Family
Trochomorphidae Trochomorphidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Gastrodontoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Since 2017, its classification has bee ...
**Fossil taxa probably belonging to the Gastrodontoidea ***Subfamily † Archaeozonitinae ***Subfamily † Grandipatulinae ***Subfamily † Palaeoxestininae *Superfamily
Parmacelloidea Parmacelloidea is a superfamily of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora and the informal group Pulmonata. These are limacoid or keelback slugs. Families Families within the superfamily ...
**Family
Parmacellidae Parmacellidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks within the superfamily Parmacelloidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). This family has no subfamilies (ac ...
**Family
Milacidae Milacidae is a family of air-breathing, keeled, land slugs. These are shell-less terrestrial gastropod mollusks in the superfamily Parmacelloidea. This family has no subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, ...
**Family
Trigonochlamydidae Trigonochlamydidae is a family of air-breathing land slugs, terrestrial gastropod molluscs in the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Distribution The native distribution of Trigonochla ...
*Superfamily Zonitoidea **Family
Zonitidae Zonitidae, common name the true glass snails, are a family of mostly rather small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Zonitoidea. Zonitidae is the only family in the superfamily Zonitoidea. Th ...
*Superfamily
Helicarionoidea Helicarionoidea is a superfamily of air-breathing land snails and semi-slugs, terrestrial pulmonate gastropod mollusks in the infraorder Limacoidei. Families There are three families within the superfamily Helicarionoidea: * Helicarionidae ...
**Family
Helicarionidae Helicarionidae is a family of air-breathing land snails or semi-slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea. Distribution The distribution of Helicarionidae includes the eastern Palearctic, Malagasy, Ind ...
**Family Ariophantidae **Family
Urocyclidae Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Some spec ...
*Superfamily
Limacoidea Limacoidea is a taxonomic superfamily of medium-sized to large, air-breathing land slugs and snails. They are terrestrial pulmonate gastropod mollusks in the infraorder Stylommatophora (according to the taxonomy of the Gastropoda by Bouchet ...
**Family
Limacidae Limacidae, also known by their common name the keelback slugs, are a taxonomic family of medium-sized to very large, air-breathing land slugs, terrestrial pulmonate gastropod molluscs in the superfamily Limacoidea. Distribution The distr ...
**Family
Agriolimacidae ''Agriolimacidae'' is a family of small and medium-sized land slugs, or shell-less snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusc, mollusks. Distribution Distribution of Limacidae is Holarctic, this include: Nearctic, wes ...
**Family
Boettgerillidae ''Boettgerilla'' is a genus of air-breathing land slugs, terrestrial pulmonate gastropod molluscs in the family Boettgerillidae. The generic name ''Boettgerilla'' is named after the German malacologist Oskar Boettger. Taxonomy ''Boettger ...
**Family
Vitrinidae Vitrinidae is a family of small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Limacoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). MolluscaBase eds. (2021). MolluscaBa ...


= other Sigmurethra

= Two superfamilies belongs to clade Sigmurethra, but they are not in the limacoid clade. *Superfamily
Arionoidea Arionoidea is a taxonomic group, superfamily of air-breathing land slugs, shell-less terrestrial pulmonate gastropod mollusks. Families Families within the superfamily Arionoidea include: * Arionidae * Anadenidae * Ariolimacidae * Bin ...
**Family
Arionidae Arionidae, common name the "roundback slugs" or "round back slugs" are a taxonomic family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Arionoidea. Distribution The distribution of this family of slug ...
**Family
Anadenidae Anadenidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). A ...
**Family
Ariolimacidae Ariolimacidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Arionoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Subfamilies and genera The family Ariolimac ...
**Family
Binneyidae Binneyidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Arionoidea Arionoidea is a taxonomic group, superfamily of air-breathing land slugs, shell-less terrestrial pulmonate gastr ...
**Family
Oopeltidae Oopeltidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Arionoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Subfamilies and genera The family Oopeltidae c ...
**Family Philomycidae *Superfamily
Helicoidea Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora. Taxonomy 2005 taxonomy There are 19 families within the superfamily Helicoidea according to the ta ...
**Family Helicidae **Family Bradybaenidae **Family Camaenidae **Family
Cepolidae The bandfishes, family Cepolidae, are 23 species of marine ray-finned fishes, They are native to the East Atlantic and Indo-Pacific wherethey dig burrows in sandy or muddy seabeds and eat zooplankton. Taxonomy The bandfishes belong to the famil ...
**Family
Cochlicellidae Cochlicellidae is a family of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod moll ...
**Family Elonidae **Family
Epiphragmophoridae Epiphragmophorinae is a subfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Xanthonychidae (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). This family has no subfamilies. ...
**Family
Halolimnohelicidae Halolimnohelicidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mol ...
**Family
Helicodontidae Helicodontidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks i ...
**Family Helminthoglyptidae **Family
Humboldtianidae Humboldtianidae is an American family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). Anatomy The anatomy of this fam ...
**Family
Hygromiidae Hygromiidae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea. Helicellidae Ihering, 1909 Anatomy Some snails in genera within this family create ...
**Family
Monadeniidae Monadeniinae is a taxonomic subfamily of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family XanthonychidaeMolluscaBase eds. (2021). MolluscaBase. Monadenia Pilsbry, 1895. Accessed through: Worl ...
**Family Pleurodontidae **Family
Polygyridae Polygyridae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea. MolluscaBase eds. (2021). MolluscaBase. Polygyridae Pilsbry, 1895. Accessed through: World Register of Marine Species a ...
**Family
Sphincterochilidae Sphincterochilidae is a taxonomic family of medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). In the fossil re ...
**Family
Thysanophoridae Thysanophoridae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). These are common woodland snails of Nort ...
**Family
Trissexodontidae Trissexodontidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). This family has no subfamilies. The famil ...
**Family
Xanthonychidae Xanthonychidae is a family (biology), family of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea. MolluscaBase eds. (2021). MolluscaBase. Xanthonychidae Strebel & Pfeffer, 187 ...


2010 taxonomy

Jörger ''et al.'' (2010) analyzed major groups within the Heterobranchia using genetic data and found that Pulmonata as traditionally defined was polyphyletic, for instance some pulmonates were more closely related to Sacoglossa and
Acochlidia Acochlidiacea, common name acochlidians, are a taxonomic clade of very unusual sea snails and sea and freshwater slugs, aquatic gastropod mollusks within the large clade Heterobranchia. Acochlidia is a variant spelling. Description Thes ...
. They proposed the more inclusive taxon Panpulmonata to unite the clades
Siphonarioidea Siphonarioidea is a taxonomic superfamily of air-breathing sea snails or false limpets, marine gastropod mollusks in the clade Panpulmonata. Distribution The Siphonarioidea are broadly distributed across the globe, however they are known to be ...
, Sacoglossa,
Glacidorboidea The Glacidorbidae is a taxonomic family of freshwater snails. Taxonomy 2005 taxonomy Glacidorbidae is the only family in the superfamily Glacidorboidea. Glacidorboidea has been classified within the informal group Lower Heterobranchia in the ...
,
Pyramidelloidea Pyramidelloidea is a superfamily of mostly very small sea snails, marine gastropod mollusks and micromollusks within the clade Panpulmonata. This is a voluminous taxon: above the species level close to 400 named taxa are referred to this gast ...
,
Amphiboloidea Amphiboloidea is a Taxonomy (biology), taxonomic superfamily of air-breathing land snails. Distribution Amphibolids are found in Indo-Pacific intertidal mangrove, saltmarsh and estuarine mudflat habitats. Taxonomy 2005 taxonomy According ...
, Hygrophila,
Acochlidia Acochlidiacea, common name acochlidians, are a taxonomic clade of very unusual sea snails and sea and freshwater slugs, aquatic gastropod mollusks within the large clade Heterobranchia. Acochlidia is a variant spelling. Description Thes ...
and
Eupulmonata Eupulmonata is a taxonomic clade of air-breathing snails. The great majority of this group are land snails and slugs, but some are marine and some are saltmarsh snails that can tolerate salty conditions. Linnean taxonomy *Suborder Eupulmonat ...
.


References


Further reading

* *


External links


Pulmonate snails & slugs in captivity
{{Authority control Obsolete gastropod taxa Carboniferous first appearances Taxa named by Georges Cuvier