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taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
s of every kind that are scheduled described during the year 2019, as well as other significant discoveries and events related to
paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
of archosaurs that are scheduled to occur in the year 2019.


General research

* A study on patterns of evolutionary integration among regions of the archosaur skull, based on data from extant and fossil taxa, is published by Felice ''et al.'' (2019). * A review of the
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
history of
crocodyliforms Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
,
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
dinosaurs, nonavian
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaurs and mammals from the Mesozoic of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
is published by
Krause Krause (German for ''ruffle'') is a common German surname. Geographical distribution As of 2014, 64.9% of all known bearers of the surname ''Krause'' were residents of Germany (frequency 1:531), 20.6% of the United States (1:7,541), 3.5% of Brazil ...
''et al.'' (2019). * A study on the biogeography of Cretaceous terrestrial tetrapods, including terrestrial crocodyliforms, non-avian dinosaurs, birds and pterosaurs, is published by Kubo (2019). * A study on size and shape differences between brains and
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
s of extant
American alligator The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the g ...
and
domestic chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
, and on its implications for inferring whether endocasts are a reliable proxy for brain
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
in archosaurs in general, is published by Watanabe ''et al.'' (2019). * A study comparing the mechanical properties of teeth of ''
Suchomimus ''Suchomimus'' (meaning "crocodile mimic") is a genus of spinosaurid dinosaur that lived between 125 and 112 million years ago in what is now Niger, during the Aptian to early Albian stages of the Early Cretaceous period. It was named and ...
tenerensis'' and ''
Sarcosuchus ''Sarcosuchus'' (; ) is an extinct genus of crocodyliform and distant relative of living crocodilians that lived during the Early Cretaceous, from the late Hauterivian to the early Albian, 133 to 112 million years ago of what is now Africa and ...
imperator'' is published by Kundanati ''et al.'' (2019). * A study on the distribution of
medullary bone The medullary cavity (''medulla'', innermost part) is the central cavity of bone shafts where red bone marrow and/or yellow bone marrow (adipose tissue) is stored; hence, the medullary cavity is also known as the marrow cavity. Located in the ma ...
in the skeletons of living birds, aiming to refine the set of criteria used to evaluate purported records of medullary bone tissue in fossil
avemetatarsalia Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinos ...
ns, is published by Canoville,
Schweitzer Schweitzer is a surname. Notable people with the surname include: * Albert Schweitzer, German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize * Anton Schweitzer, opera composer * Brian Schweitzer, forme ...
& Zanno (2019). * A study comparing the anatomy of hindlimbs of
cursorial A cursorial organism is one that is adapted specifically to run. An animal can be considered cursorial if it has the ability to run fast (e.g. cheetah) or if it can keep a constant speed for a long distance (high endurance). "Cursorial" is often u ...
birds, non-avian theropod dinosaurs and other cursorial animals, aiming to determine whether cursorial birds are good kinematic model for reconstructions of theropod dinosaur locomotion, is published by Grossi ''et al.'' (2019). * A study on the microstructure of eggshells in birds and non-avian
maniraptora Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Deinonychosauria, Oviraptoros ...
n dinosaurs is published by Choi, Han & Lee (2019). * Hu ''et al.'' (2019) reconstruct the
vomer The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...
of ''
Sapeornis ''Sapeornis'' is a monotypic genus of avialan which lived during the early Cretaceous period ( late Barremian to early Aptian, roughly 125-120 mya). ''Sapeornis'' contains only one species, ''Sapeornis chaoyangensis''. Description ''Sapeornis' ...
'' and ''
Sinovenator ''Sinovenator'' (meaning "Chinese hunter") is a genus of troodontid dinosaur from China. It is from the early Cretaceous Period (geology), Period. Discovery and naming Two specimens of a troodontidae, troodontid were described in 2002 in pale ...
'', and evaluate their implications for the knowledge of the evolution of the skull of paravians. * A study on the anatomy of skull
fenestra A fenestra (fenestration; plural fenestrae or fenestrations) is any small opening or pore, commonly used as a term in the biological sciences. It is the Latin word for "window", and is used in various fields to describe a pore in an anatomical st ...
e in
sauropsids Sauropsida ("lizard faces") is a clade of amniotes, broadly equivalent to the class Reptilia. Sauropsida is the sister taxon to Synapsida, the other clade of amniotes which includes mammals as its only modern representatives. Although early synap ...
, and on its implications for reconstructions of dinosaur soft tissues, is published online by Holliday ''et al.'' (2019). * A study aiming to determine the likely
karyotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
of the dinosaur and the early
diapsid Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago ...
ancestor of birds is published by Griffin, Larkin & O’Connor (2019). * A study on the evolution of bipedality in archosaurs is published by Grinham, VanBuren &
Norman Norman or Normans may refer to: Ethnic and cultural identity * The Normans, a people partly descended from Norse Vikings who settled in the territory of Normandy in France in the 10th and 11th centuries ** People or things connected with the Norm ...
(2019). * A study on the evolution of the brain of bird-line archosaurs is published by Beyrand ''et al.'' (2019). * A review of the progress in the field of archosaur paleo
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
, focusing in particular on the study of the dinosaurs, is published by Bailleul, O’Connor &
Schweitzer Schweitzer is a surname. Notable people with the surname include: * Albert Schweitzer, German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize * Anton Schweitzer, opera composer * Brian Schweitzer, forme ...
(2019). * A study on the phylogenetic distribution of the
hyposphene-hypantrum articulation The hyposphene-hypantrum articulation is an accessory joint found in the vertebrae of several fossil reptiles of the group Archosauromorpha. It consists of a process on the backside of the vertebrae, the hyposphene, that fits in a depression in the ...
in the vertebrae of archosaurs is published by Stefanic & Nesbitt (2019). * A study comparing growth patterns of the
American alligator The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the g ...
,
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
and ''
Tenontosaurus ''Tenontosaurus'' ( ; ) is a genus of medium- to large-sized ornithopod dinosaur. It was a relatively medium sized ornithopod, reaching in length and in body mass. It had an unusually long, broad tail, which like its back was stiffened with a n ...
tilletti'' is published by Brunner ''et al.'' (2019). * A study comparing the position, size and number of pneumatic foramina in the vertebrae of pterosaurs and birds is published by Buchmann, Avilla & Rodrigues (2019). * A study on non-avian dinosaur and bird tracks (representing some of the oldest known bird tracks) preserved in slabs used as building stones at the
Chengde Mountain Resort Chengde Mountain Resort in Chengde (; Manchu: ''Halhūn be jailara gurung''), is a large complex of imperial palaces and gardens situated in the Shuangqiao District of Chengde in northeastern Hebei province, northern China, about 225 km northea ...
, originating from the
Tuchengzi Formation The Tuchengzi Formation (formerly known as Tucheng Conglomerate, from ) is a geological formation in China whose strata span the Tithonian (Late Jurassic) to Berriasian ( Early Cretaceous) ages.Liu Y.-Q. Kuang H.-W., Jiang X.-J., Peng N., Xu H. & ...
(
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) and dating to the Jurassic-Cretaceous boundary, is published online by Xing ''et al.'' (2019). * Description of non-avian dinosaur and bird tracks from the Upper Cretaceous Chignik Formation (southwestern
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
), evaluating their implications for the knowledge of habitat preferences of northern high-latitude dinosaurs, is published by Fiorillo ''et al.'' (2019). * An assemblage of non-avian dinosaur and bird feathers is described from the Lower Cretaceous Koonwarra Fossil Bed (
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
) by Kundrát ''et al.'' (2019). * A study on
barb Barb or the BARBs or ''variation'' may refer to: People * Barb (name), a list of people and fictional characters with the given name or surname * Barb, a term used by fans of Nicki Minaj to refer to themselves * The Barbs, a band Places * Barb, ...
angles in birds and non-avian dinosaurs, evaluating their implications for the knowledge of feather shape evolution and the utility of barb angles for determination of flight abilities of fossil taxa, is published by Wang, Tang &
Clarke Clarke is a surname which means "clerk". The surname is of English and Irish origin and comes from the Latin . Variants include Clerk and Clark. Clarke is also uncommonly chosen as a given name. Irish surname origin Clarke is a popular surname i ...
(2019).


Pseudosuchians


Research

* A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of '' Coahomasuchus chathamensis'', and on its implications for inferring
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
and growth strategy of this species, is published by Hoffman, Heckert & Zanno (2019). * Tolchard ''et al.'' (2019) revise fragmentary archosaurian remains from the latest
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
lower
Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological Stratigraphic unit, group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern ...
(
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
), interpreting them as fossils of at least two distinct taxa of "
rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia ...
ns", thus representing the southernmost palaeolatitudes that these animals are known to have occurred, their first definitive remains from southern Africa, and some of the most recent records of members of this
grade Grade most commonly refers to: * Grade (education), a measurement of a student's performance * Grade, the number of the year a student has reached in a given educational stage * Grade (slope), the steepness of a slope Grade or grading may also ref ...
. * A study on the anatomy of the skeleton of ''
Poposaurus ''Poposaurus'' (" Popo Agie reptile") is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-b ...
gracilis'' is published online by Schachner ''et al.'' (2019). * A study on the age of sandstones of the Badong Formation preserving fossils of ''
Lotosaurus ''Lotosaurus'' is an extinct genus of sail-backed poposauroid Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual ...
adentus'' is published by Wang ''et al.'' (2019). * Description of the anatomy of the skull of a new specimen of ''Prestosuchus chiniquensis'' from the ''Dinodontosaurus'' Assemblage Zone of the Pinheiros-Chiniquá Sequence, Santa Maria Super sequence (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) is published by Mastrantonio ''et al.'' (2019), who also present the first description of a
rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia ...
n cranial
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
. * A study on habitat shifts during the evolutionary history of
Crocodylomorpha Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cro ...
is published by Wilberg, Turner & Brochu (2019). * A study on patterns of body size evolution of crocodylomorphs is published by Godoy ''et al.'' (2019). * A study on the quality of the fossil record of non‐marine crocodylomorphs is published by Mannion ''et al.'' (2019). * A study on the evolution of skull shape in crocodylomorphs is published online by Godoy (2019). * A study on the diversity of feeding ecologies of Mesozoic
crocodyliforms Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
is published by Melstrom & Irmis (2019). * A study on patterns of crocodyliform snout shape, on their inferred diet and on the relationship between form and function of crocodyliform skull shape throughout the evolutionary history of this group is published online by Drumheller & Wilberg (2020). * New fossil material (an isolated left
dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
) of ''
Orthosuchus ''Orthosuchus'' (meaning "straight crocodile") is an extinct genus of protosuchian crocodyliform that lived during the Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series ( ...
stormbergi'' is described from the Upper
Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological Stratigraphic unit, group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern ...
(
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
) by Dollman, Viglietti & Choiniere (2019), who also examine the
stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostrati ...
positions of all valid
crocodylomorph Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cro ...
specimens from the main
Karoo Basin The Karoo Supergroup is the most widespread stratigraphy, stratigraphic unit in Africa south of the Kalahari Desert. The group (stratigraphy), supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Pennsylvan ...
. *
Teleosaurid Teleosauridae is a family of extinct typically marine crocodylomorphs similar to the modern gharial that lived during the Jurassic period. Teleosaurids were thalattosuchians closely related to the fully aquatic metriorhynchoids, but were less ad ...
and
metriorhynchid Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period (Bajocian to early Aptian) of Europe, North America and South America. The name Metriorhynchidae ...
teeth are described from, respectively, the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
(
Aalenian The Aalenian () is a subdivision of the Middle Jurassic Epoch/Series of the geologic timescale that extends from about 174.1 Ma to about 170.3 Ma (million years ago). It was preceded by the Toarcian and succeeded by the Bajocian. Stratigraphic ...
) and
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
(
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
) of
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
by Čerňanský ''et al.'' (2019), representing the first record of members of both families from the country. * Partial skeleton of a
teleosauroid Teleosauroidea is an extinct superfamily of thalattosuchian crocodyliforms living from the Early Jurassic to the Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier o ...
crocodylomorph, representing the most recent record of a definitive non- machimosaurin teleosauroid in Africa reported so far, is described from the
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
of
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
by Dridi & Johnson (2019). * Fossils of a member of Teleosauroidea with an estimated body length of 9.6 m, representing the most recent definitive record of Teleosauroidea reported so far, are described from the Lower Cretaceous (upper
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
)
Paja Formation Paja may refer to: * Paja, Iran, a village in Sari County * Paja (given name), a Serbian masculine name * Paja (surname) * Paja Formation, geologic formation in Colombia * Paja Brava :''Paja Brava is also used in Spanish speaking countries to ...
(
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
) by Cortes ''et al.'' (2019). * Redescription of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimens of ''
Mystriosaurus ''Mystriosaurus'' is an extinct genus of teleosaurid crocodyliform from the Early Jurassic (Toarcian). Fossil specimens have been found in the Whitby Mudstone of England and Posidonia Shale of Germany. The only known species, ''M. laurillardi'', ...
laurillardi'' and ''"
Steneosaurus ''Steneosaurus'' (from el, στενός , 'narrow' and el, σαῦρος , 'lizard') is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic (Callovian or early Oxfordian) of France. The genus has been used as a was ...
" brevior'' and a study on the taxonomic validity and phylogenetic relationships of these species is published by Sachs ''et al.'' (2019). * A study on the anatomy and phylogenetic relationships of metriorhynchoids from the Jurassic Rosso Ammonitico Veronese Formation (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
) is published by Cau (2019), who provides a revised diagnosis of '' Neptunidraco ammoniticus''. * A three-dimensionally preserved
occiput The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone overlies the occipital lobes of the cereb ...
of a member of the genus '' Torvoneustes'', indicating that members of this genus reached larger body sizes than previously supposed, is described from the Upper Jurassic
Kimmeridge Clay The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for North ...
Formation (
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
) by Young ''et al.'' (2019). * A study on teeth morphology and
tooth enamel Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The ...
microstructure in ''
Mariliasuchus ''Mariliasuchus'' ("Marilia crocodile") is an extinct genus of Late Cretaceous notosuchian mesoeucrocodylian found near Marilia, Brazil. The first bone remains were found and collected in 1995 by Brazilian paleontologist William Nava, in red r ...
amarali'' is published by Augusta & Zaher (2019). * A study on the arrangement and morphology of the
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...
s of
baurusuchids Baurusuchidae is a Gondwanan family of mesoeucrocodylians that lived during the Late Cretaceous. It is a group of terrestrial hypercarnivorous crocodilians from South America (Argentina and Brazil) and possibly Pakistan. Baurusuchidae has b ...
is published by Montefeltro (2019). * A study on the anatomy of the
pterygoid Pterygoid, from the Greek for 'winglike', may refer to: * Pterygoid bone, a bone of the palate of many vertebrates * Pterygoid processes of the sphenoid bone ** Lateral pterygoid plate ** Medial pterygoid plate * Lateral pterygoid muscle * Medial ...
region and skull airways of ''
Caipirasuchus ''Caipirasuchus'' is an extinct genus of sphagesaurid notosuchians known from the Late Cretaceous of northern São Paulo State, southeastern Brazil. The type species, ''C. paulistanus'', was named in 2011. A second species, ''C. montealtensis' ...
paulistanus'' and ''C. montealtensis'' is published online by Dias ''et al.'' (2019), who report possible anatomical evidence of vocal capacity of ''C. montealtensis''. * Description of fossils and possible
gastrolith A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In othe ...
s of a large-bodied sphagesaurid from the Upper Cretaceous Adamantina Formation (Brazil) is published online by Cunha ''et al.'' (2019). * Description of new fossil material of ''
Pepesuchus ''Pepesuchus'' is an extinct genus of carnivorous metasuchian from the Late Cretaceous period (geology), period. It is a Peirosauridae, peirosaurid which lived during the Campanian and Maastrichtian stages of the Late Cretaceous in what is now st ...
'' from the
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Adamantina Formation The Adamantina Formation is a geological formation in the Bauru Basin of western São Paulo state, in southeastern Brazil. Its strata date back to the Late Cretaceous epoch of the Cretaceous Period, during the Mesozoic Era. The formation is p ...
(
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) and a study on the phylogenetic relationships of this taxon is published by Geroto & Bertini (2019). * A study on the
diagenesis Diagenesis () is the process that describes physical and chemical changes in sediments first caused by water-rock interactions, microbial activity, and compaction after their deposition. Increased pressure and temperature only start to play a ...
of fossils of '' Montealtosuchus arrudacamposi'' from the Upper Cretaceous Adamantina Formation is published by Marchetti ''et al.'' (2019). * A study on the phylogenetic relationships of members of
Neosuchia Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to ''Crocodylus niloticus' ...
and on the evolution of longirostry in this group is published online by Groh ''et al.'' (2019). * A study on the taxonomic status and phylogenetic relationships of ''
Sarcosuchus ''Sarcosuchus'' (; ) is an extinct genus of crocodyliform and distant relative of living crocodilians that lived during the Early Cretaceous, from the late Hauterivian to the early Albian, 133 to 112 million years ago of what is now Africa and ...
hartti'' is published online by Souza ''et al.'' (2019). * Partial
dyrosaurid Dyrosauridae is a family of extinct neosuchian crocodyliforms that lived from the Late Cretaceous (Maastrichtian) to the Eocene. Dyrosaurid fossils are globally distributed, having been found in Africa, Asia, Europe, North America and South Amer ...
skeleton discovered in the 1930s in
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
(
Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretaceous ...
) strata along the Atlantic coast of
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
is described by Martin, Sarr & Hautier (2019). * Description of new dyrosaurid specimens from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
–early
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
of
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...
(
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), and a study on their implications for the validity of the species ''
Hyposaurus ''Hyposaurus'' is a genus of extinct marine dyrosaurid crocodyliform. Fossils have been found in Paleocene aged rocks of the Iullemmeden Basin in West Africa, Campanian–Maastrichtian (Late Cretaceous) Shendi Formation of Sudan and Maastrichtian ...
rogersii'', is published online by Souza ''et al.'' (2019). * Revision of the large-sized
neosuchia Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to ''Crocodylus niloticus' ...
ns '' Kansajsuchus'' and "'' Turanosuchus''" from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
of Central Asia is published by Kuzmin ''et al.'' (2019), who interpret ''Kansajsuchus'' as a member of
Paralligatoridae Paralligatoridae is an extinct family of neosuchian crocodyliforms that existed during the Jurassic and Cretaceous periods. It includes the genera ''Paralligator'', '' Brillanceausuchus'', '' Kansajsuchus'', ''Shamosuchus'', ''Scolomastax'', ''Sa ...
, and consider ''Turanosuchus aralensis'' to be a member of the genus ''Kansajsuchus'' belonging or related to the species ''K. extensus''. * A study on the inner cavities of the skull of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''
Lohuecosuchus ''Lohuecosuchus'' (meaning "Lo Hueco crocodile") is an extinct genus of Allodaposuchidae, allodaposuchid eusuchian Crocodylomorpha, crocodylomorph that lived during the Late Cretaceous (late Campanian to early Maastrichtian) in what is now Spain ...
megadontos'' is published by Serrano-Martínez ''et al.'' (2019). * Revision of the fossil material of ''
Allodaposuchus ''Allodaposuchus'' is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages of the Late Cretaceous. Although generally classified as a non-crocodylian eusuchian crocodylomorph, ...
precedens'' from Vălioara (
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
) is published online by Narváez ''et al.'' (2019), who emend the diagnosis for this species. * A study on palaeodiversity of
eusuchia Eusuchia is a clade of crocodylomorphs that first appeared in the Early Cretaceous with ''Hylaeochampsa''. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades d ...
ns over time is published online by De Celis, Narváez & Ortega (2019). * A tooth of a juvenile specimen of ''
Deinosuchus ''Deinosuchus'' () is an extinct genus of alligatoroid crocodilian, related to modern alligators and caimans, that lived 82 to 73 million years ago (Ma), during the late Cretaceous period. The name translates as "terrible crocodile" and is ...
'', providing new information on the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
of this reptile, is described by Brownstein (2019). * A well-preserved braincase of ''
Diplocynodon ''Diplocynodon'' is an extinct genus of alligatoroid that lived during the Paleocene to Middle Miocene in Europe. It looked very similar to the modern caiman in that it was small and had bony armour scutes covering its neck, back, belly, and tail ...
tormis'' is described from the middle
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
site of ‘Teso de la Flecha’ (
Salamanca Salamanca () is a city in western Spain and is the capital of the Province of Salamanca in the autonomous community of Castile and León. The city lies on several rolling hills by the Tormes River. Its Old City was declared a UNESCO World Heritag ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) by Serrano-Martínez ''et al.'' (2019). * A study on the anatomy and phylogenetic relationships of ''Diplocynodon hantoniensis'' is published online by Rio ''et al.'' (2020). * Chroust, Mazuch & Luján (2019) describe new fossil material of ''Diplocynodon'' from four sites in the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
dating to Eocene–Oligocene transition, and evaluate the implications of these fossils for the knowledge of the course of the Eocene–Oligocene cooling event in Central Europe. * New crocodylian fossils, documenting the presence of four previously unrecognised alligatoroids, are described from the Lower
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Castillo Formation (
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
) by Solórzano ''et al.'' (2019). * A taxonomic and phylogenetic revision of ''
Necrosuchus ''Necrosuchus'' is an extinct genus of caiman from modern day Argentina that lived during the Paleocene epoch (Selandian age, about 60 million years ago). It inhabited the fluvio-lacustrine environment of the Patagonian Salamanca Formation. Hist ...
ionensis'' is published online by Cidade, Fortier & Hsiou (2019). * Ten late Miocene specimens of ''
Mourasuchus ''Mourasuchus'' is an extinct genus of giant, aberrant caiman from the Miocene of South America. Its skull has been described as duck-like, being broad, flat, and very elongate, superficially resembling ''Stomatosuchus'' from the Late Cretaceou ...
'', tentatively assigned to the species ''M. arendsi'', are described from
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
and from the
Solimões Formation The Pebas Formation is a lithostratigraphic unit of Miocene age, found in western Amazonia. The formation extends over , including parts of Brazil, Peru, Ecuador and Colombia.Wesselingh et al., 2006 It is interpreted as representing the deposits ...
of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
by Cidade ''et al.'' (2019), who also discuss the morphology of ''Mourasuchus'' and paleogeographic distribution of this genus in the Miocene of South America. * A study on the anatomy of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
of ''Mourasuchus amazonensis'' and on the taxonomic status of species belonging to the genus ''Mourasuchus'' is published by Cidade ''et al.'' (2019). * A study on the feeding habits of ''Mourasuchus'' is published by Cidade, Riff & Hsiou (2019). * A study on the structure of the
vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordata, ...
of ''
Purussaurus ''Purussaurus'' is an extinct genus of giant caiman that lived in South America during the Miocene epoch, from the Friasian to the Huayquerian in the SALMA classification. It is known from skull material found in the Brazilian and Peruvian Amazon ...
mirandai'', providing evidence of a deviation from the vertebral count present in all extant crocodilians, is published by Scheyer ''et al.'' (2019). * A study on the taxonomic status of ''
Balanerodus ''Balanerodus'' is an extinct monospecific genus of alligatorid crocodylian. Fossils have been found from the Fitzcarrald Arch in the Peruvian Amazon and the La Victoria Formation of the Honda Group in Colombia and date back to the Friasian and ...
logimus'' and '' Caiman venezuelensis'' is published by Cidade ''et al.'' (2019). * Fossils of a specimen of ''
Asiatosuchus ''Asiatosuchus'' is an extinct genus of crocodyloid crocodilians that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to ''Asiatosuchus'' since the genus was named in 1940. These specie ...
depressifrons'' from the late
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
of Mont de
Berru :''Berrú leads here. For Max Berrú Carrión (1942-2018), the Ecuadorian and Chilean musician, see Max Berrú'' Berru () is a commune in the Marne department in northeastern France. Berru, along with the neighboring commune of Cernay-lès-Reims, ...
(
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
), representing the oldest European
crocodyloid Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to th ...
remains reported so far, are described by Delfino ''et al.'' (2019). * A study on geographical origin, historical
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
and evolution of traits aiding dispersal of members of the genus ''
Crocodylus ''Crocodylus'' is a genus of true crocodiles in the family Crocodylidae. Taxonomy The generic name, ''Crocodylus'', was proposed by Josephus Nicolaus Laurenti in 1768. ''Crocodylus'' contains 13–14 extant (living) species and 5 extinct species ...
'' is published by Nicolaï & Matzke (2019). * Evidence of gavialine‐specific
atavistic In biology, an atavism is a modification of a biological structure whereby an ancestral genetic trait reappears after having been lost through evolutionary change in previous generations. Atavisms can occur in several ways; one of which is when ...
characters in the skeletons of fossil tomistomines ''
Penghusuchus ''Penghusuchus'' is an extinct genus of gavialid crocodylian. It is known from a skeleton found in Upper Miocene rocks of Penghu Island, off Taiwan. It may be related to two other fossil Asian gavialids: '' Maomingosuchus petrolica'' of southe ...
pani'' and ''
Toyotamaphimeia ''Toyotamaphimeia machikanensis'' (Toyotama-hime from Mountain Machikane ( :ja:待兼山)) is an extinct gavialid crocodylian which lived in Japan during the Pleistocene. A specimen recovered in 1964 at Osaka University during the construction of ...
machikanensis'' is presented by Iijima & Kobayashi (2019). * Skull and
mandibular In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
elements of a tomistomine (probably belonging to the genus ''
Maomingosuchus ''Maomingosuchus'' is an extinct genus of gavialoid crocodylian from Late Eocene of Southeast Asia. It was discovered in Priabonian-aged deposits of China and possibly also Thailand. The type species, originally ''Tomistoma petrolica,'' was named ...
'') are described from the late
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
lignite Lignite, often referred to as brown coal, is a soft, brown, combustible, sedimentary rock formed from naturally compressed peat. It has a carbon content around 25–35%, and is considered the lowest rank of coal due to its relatively low heat ...
seams of Krabi (
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
) by Martin ''et al.'' (2019), providing evidence of tomistomines living in the tropics in the late Eocene. * A revision of members of the genus ''
Gavialis ''Gavialis'' is a genus of crocodylians that includes the living gharial ''Gavialis gangeticus'' and one known extinct species, '' Gavialis bengawanicus.'' ''G. gangeticus'' comes from the Indian Subcontinent, while ''G. bengawanicus'' is known ...
'' described on the basis of fossils from the
Sivalik Hills The Sivalik Hills, also known as the Shivalik Hills and Churia Hills, are a mountain range of the outer Himalayas that stretches over about from the Indus River eastwards close to the Brahmaputra River, spanning the northern parts of the Indi ...
of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
and
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
is published by Martin (2019). * A study on the systematics of crocodilians known from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
fossil locality of Monteviale (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
) is published by Macaluso ''et al.'' (2019). * A revision of fossil record of
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
crocodilians from
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
(Italy) is published by Zoboli ''et al.'' (2019). * A review of the fossil crocodylomorph fauna of the Cenozoic of South America is published by Cidade, Fortier & Hsiou (2019). * A method for the quantification of size- and shape-
heterodont In anatomy, a heterodont (from Greek, meaning 'different teeth') is an animal which possesses more than a single tooth morphology. In vertebrates, heterodont pertains to animals where teeth are differentiated into different forms. For example, ...
y in members of Crocodylia is presented by D’Amore ''et al.'' (2019), who apply their method to extant and fossil crocodylomorphs. * A study on the global diversification dynamics of crocodylians since the Cretaceous is published online by Solórzano ''et al.'' (2019). * A study testing whether the bone ornamentation may play a role in terms of load-bearing capacity and mechanical strength of pseudosuchian
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...
s, based on data from five osteoderms of crocodylomorphs (representing four species: '' Caiman crocodilus'', ''
Osteolaemus tetraspis The dwarf crocodile (''Osteolaemus tetraspis''), also known as the African dwarf crocodile, broad-snouted crocodile (a name more often used for the Asian mugger crocodile) or bony crocodile, is an African crocodile that is also the smallest exta ...
'', ''
Hyposaurus ''Hyposaurus'' is a genus of extinct marine dyrosaurid crocodyliform. Fossils have been found in Paleocene aged rocks of the Iullemmeden Basin in West Africa, Campanian–Maastrichtian (Late Cretaceous) Shendi Formation of Sudan and Maastrichtian ...
rogersii'', ''
Sarcosuchus ''Sarcosuchus'' (; ) is an extinct genus of crocodyliform and distant relative of living crocodilians that lived during the Early Cretaceous, from the late Hauterivian to the early Albian, 133 to 112 million years ago of what is now Africa and ...
imperator'') and one aetosaur osteoderm (''
Aetosaurus ''Aetosaurus'' is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: ''A. ferratus'', the type species from Germany a ...
'' sp.), is published by Clarac ''et al.'' (2019). * A study on the utility of head width as a body size proxy in extant crocodylians, and on its implications for estimates of body size of extinct
crocodyliforms Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
, is published by O’Brien ''et al.'' (2019). * A study comparing skull anatomy and inferred head musculature, stress distribution in skulls and feeding mechanisms in members of the genera ''
Pelagosaurus ''Pelagosaurus'' (meaning "lizard of the open sea") is an extinct genus of thalattosuchian crocodyliform that lived during the Toarcian stage of the Lower Jurassic, around 183 Ma to 176 Ma (million years ago), in shallow epicontinental seas that ...
'' and ''
Gavialis ''Gavialis'' is a genus of crocodylians that includes the living gharial ''Gavialis gangeticus'' and one known extinct species, '' Gavialis bengawanicus.'' ''G. gangeticus'' comes from the Indian Subcontinent, while ''G. bengawanicus'' is known ...
'', and evaluating changes in
mandibular In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
function and feeding through time in the macroevolution of Crocodylomorpha, is published by Ballell ''et al.'' (2019). * Description of fossils of longirostrine crocodylians from the
Bartonian The Bartonian is, in the ICS's geologic time scale, a stage or age in the middle Eocene Epoch or Series. The Bartonian Age spans the time between . It is preceded by the Lutetian and is followed by the Priabonian Age. Stratigraphic definitio ...
of southern
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
is published by Jouve, Khalloufi & Zouhri (2019), who also discuss the implications of these fossils for the knowledge of the evolution of crocodylians through the Eocene–Oligocene transition. * A study on the diversity of Late Jurassic crocodylomorph teeth from Valmitão (
Lourinhã Formation The Lourinhã Formation () is a fossil rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the ...
,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
), and on the ecological niches and feeding behaviours of crocodylomorphs from this assemblage, is published online by Guillaume ''et al.'' (2019). * Rivera-Sylva ''et al.'' (2019) report the first crocodyliform remains from La Parrita locality (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
Cerro del Pueblo Formation The Cerro del Pueblo Formation is a geological Formation (geology), formation in Coahuila, Mexico whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel et al., ...
,
Coahuila Coahuila (), formally Coahuila de Zaragoza (), officially the Free and Sovereign State of Coahuila de Zaragoza ( es, Estado Libre y Soberano de Coahuila de Zaragoza), is one of the 32 states of Mexico. Coahuila borders the Mexican states of N ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
). * Description of an isolated crocodyliform tooth from the upper Eocene Ergilin Dzo Formation (
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
) and a study on the implications of this fossil for the knowledge of the regional paleoclimate of the area of Mongolia during the late Eocene is published by Iijima ''et al.'' (2019). * A study on the morphological diversity and phylogenetic affinities of crocodylomorph teeth from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
Tremp Formation The Tremp Formation ( es, Formación de Tremp, ca, Formació de Tremp), alternatively described as Tremp Group ( es, Grupo Tremp), is a geological formation in the comarca Pallars Jussà, Lleida, Spain. The formation is restricted to the Trem ...
(north-eastern
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) is published online by Blanco ''et al.'' (2019).


New taxa


Non-avian dinosaurs


Research


General

* A study aiming to identify the most likely area for the geographic origin of dinosaurs is published by Lee ''et al.'' (2019). * A study evaluating the impact of new fossil discoveries and changing phylogenetic hypotheses on
biogeographical Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, i ...
scenarios for dinosaur origins is published by Marsola ''et al.'' (2019). * A study aiming to determine the degree of differences of dinosaur phylogenies inferred from skull and postcranial characters is published online by Li,
Ruta ''Ruta'' (commonly known as rue) is a genus of strongly scented evergreen subshrubs, 20–60 cm tall, in the family Rutaceae, native to the Mediterranean region, Macaronesia and southwest Asia. About ten species are accepted in the genus. T ...
& Wills (2019). * A study on the
chronostratigraphic Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time. The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geological ...
position of the uppermost Cretaceous dinosaur localities from south-western Europe, and on their implications for inferring the course of the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
dinosaur turnover, is published by Fondevilla ''et al.'' (2019). * A study aiming to quantify the habitat of latest Cretaceous North American dinosaurs, based on data from fossil occurrences and climatic and environmental modelling, and evaluating its implications for inferring whether dinosaur diversity was in decline prior to the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
, is published by Chiarenza ''et al.'' (2019). * A study on factors determining the community richness of large herbivorous dinosaurs from the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) is published by Mallon (2019). * A review of the fossil record of Late Cretaceous non-avian dinosaurs from the James Ross Basin (
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
) is published by Lamanna ''et al.'' (2019), who also describe fragmentary new
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
and
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world ...
material from the Cape Lamb Member of the
Snow Hill Island Formation The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
and the Sandwich Bluff Member of the
Lopez de Bertodano Formation The Lopez de Bertodano Formation is a geological formation in the James Ross archipelago of the Antarctic Peninsula. The strata date from the end of the Late Cretaceous (upper-lower Maastrichtian stage) to the Danian stage of the lower Paleocene ...
. * A review and evaluation of studies on molecular data from Mesozoic dinosaur fossils is published by
Schweitzer Schweitzer is a surname. Notable people with the surname include: * Albert Schweitzer, German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize * Anton Schweitzer, opera composer * Brian Schweitzer, forme ...
''et al.'' (2019). * A study on the nature of putative remains of ancient proteins, blood vessels, and cells preserved with dinosaur fossils, based on data from fossils of ''
Centrosaurus ''Centrosaurus'' ( ; ) is a genus of herbivorous ceratopsian dinosaur from the Late Cretaceous of Canada. Their remains have been found in the Dinosaur Park Formation, dating from 76.5 to 75.5 million years ago. Discovery and naming The firs ...
apertus'' from the
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
), is published by Saitta ''et al.'' (2019). * A study on the
olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a grey matter, neural structure of the vertebrate forebrain involved in olfaction, the sense of odor, smell. It sends olfactory information to be further processed in the amygdala, the orbitof ...
ratio (the size of the olfactory bulb relative to the cerebral hemisphere) in dinosaurs, and on its implication for inferring olfactory acuity of dinosaurs, is published by Hughes & Finarelli (2019). * A study on vascular correlates in dinosaur skulls, evaluating their implications for the knowledge of thermoregulatory strategies of dinosaurs of different sizes, is published online by Porter & Witmer (2019). * A review of the diversity of the musculature of the skulls of herbivorous dinosaurs is published online by Nabavizadeh (2019). * A study on the evolution of different modes of herbivory in non-avian dinosaurs is published online by Button & Zanno (2019). * A study on the structure of eggshells of eggs produced by ''
Lufengosaurus ''Lufengosaurus'' (, meaning "Lufeng lizard") is a genus of Massospondylidae, massospondylid dinosaur which lived during the Early Jurassic period in what is now southwestern China. Discovery, taxonomy and research During the late 1930s geologi ...
'', ''
Massospondylus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by S ...
'' and ''
Mussaurus ''Mussaurus'' (meaning " mouse lizard") is a genus of herbivorous sauropodomorph dinosaur that lived in southern Argentina during the Early Jurassic, with a maximum age of 192.78 ± 0.14 Ma. It receives its name from the small size of the sk ...
'', representing the oldest confirmed
amniote Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are disti ...
eggshells reported so far, is published by Stein ''et al.'' (2019). * Description of
dinosaur egg Dinosaur eggs are the organic vessels in which a dinosaur embryo develops. When the first scientifically documented remains of non-avian dinosaurs were being described in England during the 1820s, it was presumed that dinosaurs had laid eggs bec ...
fossils from the late
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
Chaochuan Formation (
Zhejiang Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by Jiang ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) is published by Zhang ''et al.'' (2019), who name a new
ootaxon Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains of t ...
'' Multifissoolithus chianensis''. * Dinosaurs eggs assigned to the oofamily
Dendroolithidae ''Dendroolithus'' is an oogenus of Dendroolithidae, Dendroolithid dinosaur egg found in the Cenomanian, late Cenomanian Chichengshan Formation (Tiantai Group), in the Gong-An-Zhai Formation, Gong-An-Zhai and Santonian Majiacun Formations of China ...
are described from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Zhaoying Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by He ''et al.'' (2019), who name a new ootaxon '' Pionoolithus quyuangangensis''. * Dinosaurs eggs assigned to the oofamily
Faveoloolithidae Faveoloolithidae is an oofamily of dinosaur eggs. It contains '' Faveoloolithus'', '' Hemifaveoloolithus'', '' Parafaveoloolithus'', and probably '' Sphaerovum''. However, unlike the other Faveoloolthids, ''Sphaerovum'' has compactituberculate o ...
are described from the Upper Cretaceous (
Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The ...
) siltstones within the Daeri Andesite of the Wido Volcanics (
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
) by Kim ''et al.'' (2019), who name a new ootaxon '' Propagoolithus widoensis''. * Possible
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
eggshells are described from the Upper Cretaceous Wido Volcanics (South Korea) by Choi & Lee (2019), who name a new ootaxon '' Reticuloolithus acicularis''. * Description of an intact dinosaur egg from the Cretaceous
Wayan Formation The Wayan Formation is a geological formation in Idaho whose strata date back to the latest Early Cretaceous and the earliest Late Cretaceous. Dinosaur, other reptile, mammal, and micro and macro-floral remains are among the fossils that have been ...
(
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) assigned to the oogenus ''
Macroelongatoolithus ''Macroelongatoolithus'' is an oogenus of large theropod dinosaur eggs, representing the eggs of giant caenagnathid oviraptorosaurs. They are known from Asia and from North America. Historically, several oospecies have been assigned to ''Macroelo ...
'' is published by Simon ''et al.'' (2019), who interpret this specimen as evidence of presence of a ''
Gigantoraptor ''Gigantoraptor'' () is a genus of large oviraptorosaur dinosaur that lived in Asia during the Late Cretaceous period. It is known from the Iren Dabasu Formation of Inner Mongolia, where the first remains were found in 2005. ''Gigantoraptor'' wa ...
''-sized
oviraptorosaur Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period (geology), Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like s ...
in western North America. * A study on the embryonic metabolism of ''
Troodon ''Troodon'' ( ; ''Troödon'' in older sources) is a wastebasket taxon and a dubious genus of relatively small, bird-like dinosaurs known definitively from the Campanian age of the Late Cretaceous period (about 77  mya). It includes at least ...
formosus'', ''
Protoceratops ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
andrewsi'' and ''
Hypacrosaurus ''Hypacrosaurus'' (meaning "near the highest lizard" reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high because it was almost but not quite as large as ''Tyrannosaurus'') was a genus of duckbill dinosaur similar in appearance to ''Co ...
stebingeri'', and on its implications for the knowledge of the incubation times for dinosaur eggs, is published by Lee (2019). * A new dinosaur nesting site, preserving at least 15 egg clutches probably laid by a non-avian theropod dinosaur, is described from the Upper Cretaceous Javkhlant Formation (
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
) by Tanaka ''et al.'' (2019), who interpret the finding as evidence of colonial nesting in a non-avian dinosaur. * A study aiming to determine possible shifts from quadrupedality to bipedality during
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
in dinosaurs is published online by Chapelle ''et al.'' (2019). * A review of evidence of probable responses of dinosaurs to serious injuries is presented by Hearn & Williams (2019). * A study on the phylogenetic placement of ''
Chilesaurus ''Chilesaurus'' is an extinct genus of herbivorous dinosaur. The type and only known species so far is ''Chilesaurus diegosuarezi''. ''Chilesaurus'' lived about 145 million years ago (Mya) in the Late Jurassic period of Chile. Showing a combinat ...
diegosuarezi'' and its implications for the phylogenetic relationships of major dinosaur groups is published by Müller & Dias-da-Silva (2019).


Saurischians


= Theropods

= * A study on specimen completeness in the fossil record of non-avian theropod dinosaurs is published by Cashmore & Butler (2019). * A study on the distribution of discrete dental characters in theropod dinosaurs, and on the taxonomic value of theropod teeth, is published by Hendrickx ''et al.'' (2019). * A study aiming to evaluate whether the maximum body size of theropod dinosaurs increased across the
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
-
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
boundary is published online by Griffin & Nesbitt (2019). * A revision of theropod dinosaur fossils from the Late Jurassic to mid-Cretaceous of Southeast Asia is published by Samathi, Chanthasit & Sander (2019). * Description of theropod dinosaur teeth from the
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
(
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
-
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
)
Ilek Formation The Ilek Formation is a Lower Cretaceous geologic formation in Western Siberia. Many different fossils have been recovered from the formation. It overlies the Late Jurassic Tyazhin Formation and underlies the Albian Kiya Formation. The formation ...
(West Siberia,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
) is published by Averianov, Ivantsov & Skutschas (2019). * A study re-assessing the evidence for evolutionary
allometric Allometry is the study of the relationship of body size to shape, anatomy, physiology and finally behaviour, first outlined by Otto Snell in 1892, by D'Arcy Thompson in 1917 in ''On Growth and Form'' and by Julian Huxley in 1932. Overview Allom ...
trends in the forelimbs of non-avian theropod dinosaurs is published by Palma Liberona ''et al.'' (2019). * Redescription of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''
Chindesaurus ''Chindesaurus'' ( ) is an extinct genus of basal saurischian dinosaur from the Late Triassic (213-210 million years ago) of the southwestern United States. It is known from a single species, ''C. bryansmalli'', based on a partial skeleton recovere ...
bryansmalli'' and a study on the phylogenetic relationships of this species is published by Marsh ''et al.'' (2019). * Description of two fragmentary
neotheropod Neotheropoda (meaning "new theropods") is a clade that includes coelophysoids and more advanced theropod dinosaurs, and is the only group of theropods that survived the Triassic–Jurassic extinction event. All neotheropods became extinct by the e ...
specimens from the
Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. Th ...
Bull Canyon Formation The Cooper Canyon Formation is a geological formation of Norian age in Texas and New Mexico.Weishampel, David B; et al. (2004).Dinosaur distribution (Late Triassic, North America)" In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (ed ...
(
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), and a study on their implications for the knowledge of body size evolution among early theropods, is published by Griffin (2019). * A study on the anatomy of the braincase, the skull
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
and the inner ear of ''
Zupaysaurus ''Zupaysaurus'' (; "ZOO-pay-SAWR-us") is an extinct genus of early theropod dinosaur living during the Norian stage of the Late Triassic in what is now Argentina. Fossils of the dinosaur were found in the Los Colorados Formation of the Ischigu ...
rougieri'' is published by Paulina-Carabajal, Ezcurra &
Novas A nova (plural novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", which is Latin for "new") that slowly fades over weeks or months. Causes of the dramat ...
(2019). * A study on range of motion and functions of the forelimbs of ''
Dilophosaurus ''Dilophosaurus'' ( ) is a genus of theropod dinosaurs that lived in what is now North America during the Early Jurassic, about 193 million years ago. Three skeletons were discovered in northern Arizona in 1940, and the two best preserved ...
wetherilli'' is published by Senter & Sullivan (2019). * A study on the paleobiology of ''
Cryolophosaurus ''Cryolophosaurus'' ( or ; "CRY-oh-loaf-oh-SAWR-us") is a genus of large theropod dinosaur known from only a single species ''Cryolophosaurus ellioti'', from the early Jurassic of Antarctica. It was one of the largest theropods of the Early Jura ...
'' is published by Yun (2019). * A study on tooth formation and replacement rates in ''
Majungasaurus ''Majungasaurus'' (; ) is a genus of abelisaurid theropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period (geology), Period, making it one of the last known non-avian dinosaurs that went ex ...
'', ''
Ceratosaurus ''Ceratosaurus'' (from Ancient Greek, Greek κέρας/κέρατος, ' meaning "horn" and wikt:σαῦρος, σαῦρος ' meaning "lizard") was a carnivorous Theropoda, theropod dinosaur in the Late Jurassic Period (geology), period (Kim ...
'' and ''
Allosaurus ''Allosaurus'' () is a genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch (Kimmeridgian to late Tithonian). The name "''Allosaurus''" means "different lizard" alluding to ...
'' is published by D’Emic ''et al.'' (2019). * A study on the ecology of ''
Ceratosaurus ''Ceratosaurus'' (from Ancient Greek, Greek κέρας/κέρατος, ' meaning "horn" and wikt:σαῦρος, σαῦρος ' meaning "lizard") was a carnivorous Theropoda, theropod dinosaur in the Late Jurassic Period (geology), period (Kim ...
'' is published by Yun (2019). * A study on the phylogenetic relationships of ''
Afromimus ''Afromimus'' (meaning "Africa mimic") is a genus of theropod dinosaur from the Early Cretaceous Elrhaz Formation of Niger. It contains a single species, ''A. tenerensis'', named in 2017 by Paul Sereno from parts of the right leg, vertebrae, and ...
tenerensis'' is published by Cerroni ''et al.'' (2019), who consider this taxon to be more likely an abelisauroid rather than an ornithomimosaur. * Description of isolated neck vertebrae of abelisauroid theropods from the Cretaceous
Kem Kem Beds The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations, ...
(
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
) and a study on the phylogenetic affinities of these fossils is published online by Smyth ''et al.'' (2019). * Partially preserved ilium of an indeterminate
abelisaur Abelisauroidea is typically regarded as a Cretaceous group, though the earliest abelisauridae remains are known from the Middle Jurassic of Argentina (classified as the species Eoabelisaurus mefi) and possibly Madagascar (fragmentary remains o ...
theropod is reported from the
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Kem Kem Beds The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations, ...
(
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
) by Zitouni ''et al.'' (2019); however Smyth ''et al.'' (2019) reinterpret this specimen as a fossil of ''
Spinosaurus ''Spinosaurus'' (; ) is a genus of spinosaurid dinosaur that lived in what now is North Africa during the Cenomanian to upper Turonian stages of the Late Cretaceous period, about 99 to 93.5 million years ago. The genus was known first fro ...
aegyptiacus''. * A study on the anatomy of the brain, braincase and inner ear of ''
Carnotaurus ''Carnotaurus'' (; ) is a genus of theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 71 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a single well-p ...
sastrei'' is published by Cerroni & Paulina-Carabajal (2019). * A study on the phylogenetic affinities of a tooth previously considered to be part of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
of ''
Aerosteon ''Aerosteon'' is a genus of megaraptoran dinosaur from the Late Cretaceous period of Argentina. Its remains were discovered in 1996 in the Anacleto Formation, which is from the late Campanian. The type and only known species is ''A. riocolorade ...
riocoloradensis'' is published online by Hendrickx, Tschopp & Ezcurra (2019), who consider this fossil to be an
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found ...
tooth. *
Megalosaurid Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relative ...
teeth resembling teeth of ''
Torvosaurus ''Torvosaurus'' () is a genus of carnivorous megalosaurid theropod dinosaur that lived approximately 165 to 148 million years ago during the late Middle and Late Jurassic period (Callovian to Tithonian stages) in what is now Colorado, Portuga ...
'' are described from the Upper Jurassic of
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
and
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
by Soto, Toriño & Perea (2019). * Isolated
spinosaurid The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, includi ...
teeth are described from the Lower Cretaceous of Kut Island (
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
) by Buffetaut ''et al.'' (2019). * New spinosaurid specimens are described from the Kem Kem Beds (Morocco) by Arden ''et al.'' (2019), who interpret these specimens as providing evidence of aquatic adaptations in the skulls of spinosaurids, and name a new clade
Spinosaurini The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, includi ...
; the study is subsequently criticized by Hone & Holtz (2019). * New fossil material of juvenile spinosaurids is described from the Kem Kem Beds by Lakin & Longrich (2019). * New theropod fossils, including partial tail vertebra of a member of
Megaraptora Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proporti ...
and an association of tail vertebrae and pelvic elements displaying a combination of characteristics that are present in megaraptorid and
carcharodontosaurid Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae ...
theropods, are described from the early Late Cretaceous
Griman Creek Formation The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian of the Early-Late Cretaceous.Bell et al., 2019 It is most notable being a major s ...
at
Lightning Ridge, New South Wales Lightning Ridge is a small outback town in north-western New South Wales, Australia. Part of Walgett Shire, Lightning Ridge is situated near the southern border of Queensland, about east of the Castlereagh Highway. The Lightning Ridge area is ...
(
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
) by Brougham, Smith & Bell (2019). * Partial postcranial skeleton of a probable
carcharodontosauria Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among t ...
n theropod is described from the
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
(
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
) Freixial Formation (
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
) by Malafaia ''et al.'' (2019). * Description of the anatomy of the
axial skeleton The axial skeleton is the part of the skeleton that consists of the bones of the head and trunk (anatomy), trunk of a vertebrate. In the human skeleton, it consists of 80 bones and is composed of six parts; the human skull, skull (22 bones), als ...
of ''
Concavenator ''Concavenator'' is an extinct genus of theropod dinosaur that lived approximately 130 million years ago during the early Cretaceous period (Barremian age). The type species is ''C. corcovatus''; ''Concavenator corcovatus'' means " Cuenca hunter ...
corcovatus'' is published by Cuesta, Ortega & Sanz (2019). * A study on the anatomy of the brain and inner ear of ''
Giganotosaurus ''Giganotosaurus'' ( ) is a genus of theropod dinosaur that lived in what is now Argentina, during the early Cenomanian age of the Late Cretaceous period, approximately 99.6 to 95 million years ago. The holotype specimen was discovered in the ...
carolinii'' is published online by Paulina-Carabajal & Nieto (2019). * A study on the anatomy of ''
Murusraptor ''Murusraptor'' ("wall thief") is a genus of carnivorous megaraptoran theropod dinosaur from the Sierra Barrosa Formation, part of the Neuquén Group of Patagonia, in Argentina, South America. It is known from a single specimen that consists of ...
barrosaensis'', and on its implications for inferring the phylogenetic placement of
megaraptora Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proporti ...
ns within Theropoda, is published by Rolando,
Novas A nova (plural novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", which is Latin for "new") that slowly fades over weeks or months. Causes of the dramat ...
& Agnolín (2019). * New fossil material of a megaraptorid belonging or related to the species ''
Australovenator ''Australovenator'' (meaning "southern hunter") is a genus of megaraptoran theropod dinosaur from Cenomanian (Late Cretaceous)-age Winton Formation (dated to 95 million years ago) of Australia. It is known from partial cranial and postcranial rem ...
wintonensis'' is described from the Lower Cretaceous (
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
) Eric the Red West site (
Eumeralla Formation The Eumeralla Formation is a geological formation in Victoria, Australia whose strata date back to the Early Cretaceous. It is Aptian to Albian in age. Dinosaur remains are among the fossils that have been recovered from the formation, particularl ...
;
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
) by Poropat ''et al.'' (2019). * A study comparing different methods of assessing morphological diversity of
coelurosauria Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrann ...
n
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
s is published online by Schaeffer ''et al.'' (2019). * A study on the anatomy of the skull of '' Bicentenaria argentina'' is published online by Aranciaga‐Rolando, Cerroni & Novas (2019). * New postcranial bones of ''
Kileskus ''Kileskus'' (meaning ''lizard'' in the Khakas language) is a genus of tyrannosauroid dinosaur known from partial remains found in Middle Jurassic (Bathonian stage) Itat Formation of Sharypovsky District, Krasnoyarsk Krai (Russia). Fossils rec ...
aristotocus'', providing new information on the anatomy of this species, are described from the Middle Jurassic (
Itat Formation The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from t ...
)
Itat Formation The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from t ...
(
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
) by Averianov ''et al.'' (2019). * A study on the agility and turning capability of tyrannosaurids and other large theropods is published by Snively ''et al.'' (2019), who argue that tyrannosaurids could turn with greater agility, thus pivoting more quickly, than other large theropods, which enhanced their ability to pursue and subdue prey. * A study on the taxonomic identity of the tyrannosaurid specimen CMN 11315 from the lower
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
Tolman Member of the
Horseshoe Canyon Formation The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller. The Horseshoe Canyon Formation is part of the ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) is published online by Mallon ''et al.'' (2019). * A study on the taxonomic identity of the juvenile tyrannosaurid specimen
TMP TMP can refer to any of the following: Chemistry * 2,2,6,6-Tetramethylpiperidine, an organic chemistry reagent * Thymidine monophosphate, a nucleotide * Trimethoprim, an antibiotic * Trimethyl phosphate, a solvent * Trimethylolpropane, a precurso ...
1994.143.1, formerly assigned to the genus ''
Daspletosaurus ''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three species ...
'', is published by Voris ''et al.'', who reinterpret this specimen as belonging to the species ''
Gorgosaurus ''Gorgosaurus'' ( ; ) is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period (Campanian), between about 76.6 and 75.1 million years ago. Fossil remains have been found in the Can ...
libratus'', and describe a new
postorbital The ''postorbital'' is one of the bones in vertebrate skulls which forms a portion of the dermal skull roof and, sometimes, a ring about the orbit. Generally, it is located behind the postfrontal and posteriorly to the orbital fenestra. In some ve ...
from the
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) belonging to a small juvenile ''
Daspletosaurus ''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three species ...
''. * A study on the tooth replacement patterns in tyrannosaurid theropods, as indicated by data from a juvenile specimen of ''
Tarbosaurus ''Tarbosaurus'' ( ; meaning "alarming lizard") is a genus of tyrannosaurid dinosaur that flourished in Asia about 70 million years ago, at the end of the Late Cretaceous Period, considered to contain a single known species, ''Tarbosaurus bataa ...
bataar'', is published by Hanai & Tsuihiji (2019). * A study on teeth of ''Tarbosaurus bataar'' and its potential prey species from the
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilian ...
(
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
), aiming to infer the diet of this dinosaur and seasonal climatic variations in the area of Mongolia in the early
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
on the basis of stable isotope data from
tooth enamel Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The ...
, is published online by Owocki ''et al.'' (2019). * A study on the complexity and modularity of the skull of ''
Tyrannosaurus ''Tyrannosaurus'' is a genus of large theropoda, theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosa ...
rex'' is published by Werneburg ''et al.'' (2019). * Traces preserved on a tail vertebra of a
hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
dinosaur from the
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Hell Creek Formation The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of ...
(
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) are described by Peterson & Daus (2019), who interpret their finding as feeding traces produced by a late-stage juvenile ''Tyrannosaurus rex''. * A large specimen of ''Tyrannosaurus rex'' ( RSM P2523.8) with an estimated body mass exceeding other known ''T. rex'' specimens and representatives of all other gigantic terrestrial theropods is described by Persons,
Currie Currie ( gd, Currach, IPA: kʰuːᵲəx is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edi ...
& Erickson (2019). * A study testing the biomechanical performance of the skull of ''Tyrannosaurus rex'' is published online by Cost ''et al.'' (2019). * A study on tooth replacement in a well-preserved skull of ''Tyrannosaurus rex'' from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
Hell Creek Formation The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of ...
(
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) is published online by Sattler & Schwarz (2019). * A study aiming to determine the processes contributing to the preservation of soft tissue structures and proteins of ''Tyrannosaurus rex'' is published by Boatman ''et al.'' (2019). * Teeth on a (probably non-tyrannosaurid) tyrannosauroid and a dromaeosaurid are described from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
Mount Laurel Formation The Mount Laurel Formation is a Mesozoic geologic formation located in New Jersey and Delaware. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation.Weishampel, et al. (2004). "Dinosaur d ...
(
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) by Brownstein (2019). * Description of an
ornithomimid Ornithomimidae (meaning "bird-mimics") is a family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period of Laura ...
specimen UALVP 16182, putatively assigned to the genus ''
Dromiceiomimus ''Dromiceiomimus'' is a genus of ornithomimid theropod from the Late Cretaceous (early Maastrichtian) of Alberta, Canada. The type species, ''D. brevitertius'', is considered a synonym of ''Ornithomimus edmontonicus'' by some authors, while other ...
'', and a study on the validity of this genus is published by Macdonald &
Currie Currie ( gd, Currach, IPA: kʰuːᵲəx is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edi ...
(2019). * A study on the
morphometrics Morphometrics (from Greek μορϕή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
of teeth of ''
Richardoestesia ''Richardoestesia'' is a morphogenus of theropod dinosaur teeth, originally described from the Late Cretaceous of what is now North America. It currently contains two species, ''R. gilmorei'' and ''R. isosceles''. It has been used as a morphotax ...
asiatica'' from the Upper Cretaceous Khodzhakul, Bissekty and Aitym formations of
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
is published by Averianov & Sues (2019). * A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of a
metatarsal bone The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the ...
of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''
Xixianykus ''Xixianykus'' is a genus of alvarezsaurid theropod dinosaur from the late Cretaceous period of China.Xu, X., Wang, D.Y., Sullivan, C., Hone, D.W.E., Han, F.L., Yan, R.H. and Du, F.M. (2010). "A basal parvicursorine (Theropoda: Alvarezsauridae) ...
zhangi'' is published by Qin, Zhao & Xu (2019). * A study on the anatomy of the skull of ''
Beipiaosaurus ''Beipiaosaurus'' is a genus of therizinosauroid theropod dinosaurs that lived in Asia during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the discovery of ''Yutyrannus ...
inexpectus'' is published by Liao & Xu (2019). * A study on form, function and evolution of skulls of members of
Oviraptorosauria Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or wit ...
is published online by Ma ''et al.'' (2019). * A study on the wing performance of ''
Caudipteryx ''Caudipteryx'' (which means "tail feather") is a genus of peacock-sized theropod dinosaurs that lived in the Barremian age of the early Cretaceous (about 124.6 million years ago). They were feathered and extremely birdlike in their overall appea ...
'' is published by Talori ''et al.'' (2019). * A study on the
aerodynamic Aerodynamics, from grc, ἀήρ ''aero'' (air) + grc, δυναμική (dynamics), is the study of the motion of air, particularly when affected by a solid object, such as an airplane wing. It involves topics covered in the field of fluid dyn ...
capacity of feathered forelimbs of ''Caudipteryx'' is published by Talori & Zhao (2019). * Description of an avimimid bonebed assemblage from the
Iren Dabasu Formation The Iren Dabasu Formation (also known as Erlian Formation) is a Late Cretaceous geologic formation in the Iren Nor region of Inner Mongolia. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the fo ...
of northern China, providing new information on the growth of avimimids, is published by Funston ''et al.'' (2019). * Description of new
caenagnathid Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids ...
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
s from the
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) and a study on their
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
is published online by Funston ''et al.'' (2019). * A reconstruction of the architecture of the
oviraptorid Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one ...
egg clutch, based on data from five clutches from the Upper Cretaceous Nanxiong Group (
Jiangxi Jiangxi (; ; formerly romanized as Kiangsi or Chianghsi) is a landlocked province in the east of the People's Republic of China. Its major cities include Nanchang and Jiujiang. Spanning from the banks of the Yangtze river in the north int ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) is presented by Yang ''et al.'' (2019), who re-evaluate the hypothesis of thermoregulatory contact incubation of eggs as an explanation for the discoveries of associations of adult oviraptorosaurs with egg clutches. * A study on the reproductive biology of oviraptorids, based on data from a partial clutch of eggs from the Upper Cretaceous Nanxiong Group, is published online by Yang ''et al.'' (2019). * A study on the function of the enlarged "sickle claw" on the second toe of
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
theropods is published by Bishop (2019). * An ungual Phalanx bone, phalanx of a dromaeosaurid theropod is described from the Blagoveshchensk area (
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
) by Bolotskii, Bolotskii & Sorokin (2019). * A study on the anatomy, taphonomy, environmental setting and phylogenetic position of ''Halszkaraptor escuilliei'' is published by Brownstein (2019); the study is subsequently criticized by Cau (2020). * A study on a fossil lizard found in the abdomen of a specimen of ''Microraptor zhaoianus'' from the Lower Cretaceous Jiufotang Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
), evaluating its implications for the knowledge of dromaeosaurid digestion, is published by O’Connor ''et al.'' (2019). * Description of the anatomy of the skull of ''Saurornitholestes langstoni'', based on data from an almost complete skeleton from the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
), is published online by
Currie Currie ( gd, Currach, IPA: kʰuːᵲəx is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edi ...
& Evans (2019). * Histology, Histological analysis of the forelimb bones of ''Daliansaurus liaoningensis'' is presented by Shen ''et al.'' (2019). * Evidence indicating that the pennaceous feathers of ''Anchiornis'' were composed of both feather Beta-keratin, β-keratins and Alpha-keratin, α-keratins is presented by Pan ''et al.'' (2019); the study is subsequently criticized by Saitta & Vinther (2019). * Isolated theropod teeth, interpreted as most likely representing at least two species, are described from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
Valtos Sandstone Formation, Valtos Sandstone and Lealt Shale Formations of Skye (Scotland) by Young ''et al.'' (2019).


= Sauropodomorphs

= * A study aiming to explain high diversity of early evolutionary branches of Sauropodomorpha, sauropodomorph dinosaurs is published online by Müller & Garcia (2019). * A study on the anatomy and phylogenetic relationships of ''Pampadromaeus barberenai'' is published by Langer ''et al.'' (2019). * A Dinosauriformes, dinosauriform femur, possibly of a juvenile specimen of the species ''Pampadromaeus barberenai'', is described from the Late Triassic of southern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
by Müller ''et al.'' (2019). * A study on the anatomy of the braincase of ''Saturnalia tupiniquim'' is published by Bronzati, Langer & Rauhut (2019). * Description of all available skull bones of ''Saturnalia tupiniquim'' except the braincase, evaluating the implications of this taxon for the knowledge of the early evolution of the sauropodomorph feeding behaviour, is published by Bronzati, Müller & Langer (2019). * A study on the phylogenetic relationships of ''Unaysaurus tolentinoi'' is published online by McPhee ''et al.'' (2019). * A study on the anatomy of the skull of ''Macrocollum itaquii'' and on the phylogenetic relationships of this species is published online by Müller (2019). * A study on the bony labyrinth scale and geometry through
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
in ''
Massospondylus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by S ...
carinatus'', evaluating whether the putative gait change from quadrupedal juvenile to bipedal adult is reflected in labyrinth morphology, will be published by Neenan ''et al.'' (2019). * Description of the anatomy of the postcranial skeleton of the neotype specimen of ''Massospondylus carinatus'' is published by Barrett ''et al.'' (2019). * Redescription of the anatomy of the skull of ''Jingshanosaurus xinwaensis'' is published online by Zhang ''et al.'' (2019), who consider ''Chuxiongosaurus lufengensis'' to be a Synonym (taxonomy), junior synonym of ''J. xinwaensis''. * A study on the anatomy of the
axial skeleton The axial skeleton is the part of the skeleton that consists of the bones of the head and trunk (anatomy), trunk of a vertebrate. In the human skeleton, it consists of 80 bones and is composed of six parts; the human skull, skull (22 bones), als ...
of ''Xingxiulong chengi'' is published online by Wang ''et al.'' (2019). * A study on changes of body mass and center of mass of ''
Mussaurus ''Mussaurus'' (meaning " mouse lizard") is a genus of herbivorous sauropodomorph dinosaur that lived in southern Argentina during the Early Jurassic, with a maximum age of 192.78 ± 0.14 Ma. It receives its name from the small size of the sk ...
patagonicus'' during its
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
, and on their potential relationship with the locomotor stance of this dinosaur, is published by Otero ''et al.'' (2019). * A study on the leverage of forelimb muscles in the transition from the narrow‐gauge stance of Basal (phylogenetics), basal sauropods to a wide‐gauge stance in titanosaurs is published by Klinkhamer ''et al.'' (2019). * A study on the hind foot posture and Biomechanics, biomechanical capabilities of ''Rhoetosaurus brownei'' is published by Jannel ''et al.'' (2019). * A study on the age of the fossils of ''Rhoetosaurus brownei'' is published by Todd ''et al.'' (2019). * An isolated tooth-crown of a member of Eusauropoda, possibly a member of Mamenchisauridae or Euhelopodidae, is described from the
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
Qigu Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Maisch & Matzke (2019), representing the first record of a eusauropod from this formation reported so far. * A Cervical vertebrae, cervical vertebra of a member of the genus ''Omeisaurus'' is described from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
Lower Member of the Shaximiao Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Tan ''et al.'' (2019), providing new information on the skeletal
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
of this genus, and representing the easternmost occurrence of ''Omeisaurus'' reported so far. * Possible Mamenchisauridae, mamenchisaurid teeth are described from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
Itat Formation The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from t ...
(
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
) by Averianov ''et al.'' (2019). * A study on the age of the fossils of members of the genus ''Mamenchisaurus'' from the Suining Formation in the Sichuan Basin (China) is published by Wang ''et al.'' (2019). * A study on the anatomy and affinities of ''Lapparentosaurus madagascariensis'' is published by Raveloson, Clark & Rasoamiaramana (2019). * Partial vertebra of a sauropod dinosaur belonging to the group Turiasauria is described from the
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
Wealden Supergroup (
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
) by Mannion (2019). * Description and a study on the affinities of sauropod teeth from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
(Bathonian) Sakahara Formation (Madagascar) is published online by Bindellini & Dal Sasso (2019), who report evidence of presence of Titanosauriformes in the Bathonian. * Description of isolated sauropod vertebrae from the Oxford Clay Formation (
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
), indicative of a higher sauropod biodiversity in this formation than previously recognised, is published by Holwerda, Evans & Liston (2019). * Revision of the taxonomic diversity of sauropod dinosaurs from a historic Carnegie Museum of Natural History, Carnegie Museum locality (Red Fork of the Powder River Quarry B) in northern Wyoming (Morrison Formation) is published by Tschopp ''et al.'' (2019). * A study on pneumatic structures in the vertebrae of ''Pilmatueia faundezi'' is published online by Windholz, Rodolfo Coria, Coria & Zurriaguz (2019). * A study on the anatomy of the appendicular skeleton of ''Europasaurus holgeri'' and on the phylogenetic relationships of this species is published online by Carballido ''et al.'' (2019). * Redescription of Brachiosauridae, brachiosaurid fossil material from the Upper Jurassic Morrison Formation (Colorado,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), including a mostly complete skull discovered in 1883, is published online by D'Emic & Carrano (2019). * A study on the phylogenetic relationships of ''Galvesaurus herreroi'' is published by Pérez-Pueyo ''et al.'' (2019). * A study on the phylogenetic relationships of the Late Jurassic sauropod dinosaurs from the Tendaguru Formation of
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
(''Australodocus bohetii'', ''Janenschia robusta'' and ''Tendaguria tanzaniensis'') is published by Mannion ''et al.'' (2019). * The first confirmed fossil of a sauropod dinosaur from Ethiopia (an isolated tooth) is reported from the Upper Jurassic Mugher Mudstone by Goodwin ''et al.'' (2019). * A study on the affinities of the sauropod dinosaur known from an isolated Metacarpal bones, metacarpal from the Upper Jurassic (Oxfordian (stage), Oxfordian) Jagua Formation (Cuba) is published online by Apesteguía, Izquierdo & Iturralde-Vinent (2019). * A study on isolated sauropod teeth from the Early Cretaceous Teete locality (Batylykh Formation) (Yakutia,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
), representing the northernmost sauropod record in the Northern Hemisphere reported so far, is published online by Averianov ''et al.'' (2019). * Redescription of ''Jiangshanosaurus lixianensis'', a study on the anatomy of ''Dongyangosaurus sinensis'' and a study on the phylogenetic relationships of these species is published by Mannion ''et al.'' (2019). * New fossil material of Titanosauriformes, titanosauriform sauropods is described from the Upper Cretaceous Daijiaping Formation (Hunan,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Han ''et al.'' (2019). * A study on the long bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
in early juvenile titanosaur sauropods, evaluating its implications for the knowledge of early stages of development of these dinosaurs, is published online by González ''et al.'' (2019). * A study on the neurology and phylogenetic affinities of a titanosaurian braincase from the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
locality of Fox-Amphoux-Métisson (southeastern
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
) is published by Knoll ''et al.'' (2019). * Tail vertebrae of lithostrotian titanosaurs are described from the Lower Cretaceous
Ilek Formation The Ilek Formation is a Lower Cretaceous geologic formation in Western Siberia. Many different fossils have been recovered from the formation. It overlies the Late Jurassic Tyazhin Formation and underlies the Albian Kiya Formation. The formation ...
(Krasnoyarsk Krai,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
) by Averianov, Ivantsov & Skutschas (2019). * A study on the anatomy of the appendicular skeleton of South American titanosaur sauropods and on its implications for the knowledge of the phylogenetic relationships of these sauropods is published by Bernardo Javier González Riga, González Riga ''et al.'' (2019), who name a new clade Colossosauria. * Description of titanosaur sauropod vertebrae from the Upper Cretaceous Lameta Formation (
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
) is published by Jeffrey A. Wilson, Wilson ''et al.'' (2019). * Description of the anatomy of the braincase of ''Malawisaurus dixeyi'' is published by Andrzejewski ''et al.'' (2019), who present digital reconstructions of the
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
and inner ear of this species based on CT scanning. * A study on the anatomy and phylogenetic relationships of ''Uberabatitan ribeiroi'' is published by Silva ''et al.'' (2019). * A study on vertebral pneumaticity in ''Uberabatitan ribeiroi'', indicating that
diagenesis Diagenesis () is the process that describes physical and chemical changes in sediments first caused by water-rock interactions, microbial activity, and compaction after their deposition. Increased pressure and temperature only start to play a ...
can obliterate traces of bone pneumaticity, is published online by Aureliano ''et al.'' (2019). * Fossils of a titanosaur sauropod related to ''Rapetosaurus'' and the indeterminate Italian titanosaur specimen Museo Civico di Storia Naturale di Milano, MSNM V7157 are described from the Algora vertebrate fossil site located in the Cenomanian strata of the Arenas de Utrillas Formation (
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) by Mocho ''et al.'' (2019). * Description of five articulated sauropod dorsal vertebrae from the Upper Cretaceous
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilian ...
, possibly belonging to the species ''Nemegtosaurus mongoliensis'', is published by Averianov & Lopatin (2019), who also study the anatomy of sauropod Femur, femora from the Nemegt Formation, and argue that ''N. mongoliensis'' is likely to be distinct from ''Opisthocoelicaudia skarzynskii''.


Ornithischians

* A study on the evolution of the Femur, femoral fourth trochanter in ornithischian dinosaurs is published online by Persons &
Currie Currie ( gd, Currach, IPA: kʰuːᵲəx is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edi ...
(2019). * A study on the taphonomy and
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of the ornithischian (ankylosaurian and ornithopod) fossils from the La Cantalera-1 site (
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
Blesa Formation,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) is published by Perales-Gogenola ''et al.'' (2019).


= Thyreophorans

= * Studies on the anatomy of the skull and postcranial skeleton of ''Scelidosaurus harrisonii'' are published online by
Norman Norman or Normans may refer to: Ethnic and cultural identity * The Normans, a people partly descended from Norse Vikings who settled in the territory of Normandy in France in the 10th and 11th centuries ** People or things connected with the Norm ...
(2019). * A study on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''Bienosaurus lufengensis'', and on the taxonomic validity and phylogenetic relationships of this dinosaur, is published by Raven ''et al.'' (2019). * A study on the Morphology (biology), morphological diversity of Stegosauria, stegosaurs through the evolutionary history of the group is published by Romano (2019). * Two new stegosaurian specimens from the northernmost outcrops of the Morrison Formation in
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
, one of which is the northernmost occurrence of a dinosaur from the Morrison Formation reported so far, are described by Woodruff, Trexler & Susannah Maidment, Maidment (2019). * Description of a new specimen of ''Miragaia longicollum'' and a study on the taxonomic validity and phylogenetic relationships of this species is published by Costa & Octávio Mateus, Mateus (2019), who transfer the species ''Alcovasaurus longispinus'' to the genus ''Miragaia''. * Plates of an armored dinosaur from the Lower Jurassic (Sinemurian-Pliensbachian) Lower Kota Formation (
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
) are redescribed by Peter Galton, Galton (2019), who considers these fossils to be more similar to plates of ankylosaurians than Basal (phylogenetics), basal thyreophorans, and interprets them as the earliest ankylosaurian fossils reported so far. * Description of an assemblage of 12 partial, articulated or associated ankylosaurian skeletons and thousands of isolated bones and teeth from the Cretaceous (
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The ...
) Iharkút vertebrate locality (Hungary) will be published by Ősi ''et al.'' (2019). * A study on the evolution of morphological traits associated with tail weaponry in ankylosaurs and glyptodonts, aiming to quantitatively test the hypothesis that tail weaponry of these groups is an example of convergent evolution, is published online by Victoria Arbour, Arbour & Zanno (2019). * Murray, Riguetti & Rozadilla (2019) describe ankylosaur
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...
s from the Upper Cretaceous (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
-
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
) Allen Formation (Argentina). * Description of three new skull specimens of ''Talarurus plicatospineus'' from the Upper Cretaceous (Cenomanian–
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The ...
) Bayan Shireh Formation (
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
), and a study on the phylogenetic relationships of this species, is published online by Park ''et al.'' (2019). * Evidence for sexual dimorphism in ''Tianzhenosaurus youngi'' is presented by Ma (2019). * A study on the brain morphology and topography of cranial nerves of ''Bissektipelta archibaldi'' is published by Alifanov & Saveliev (2019). * A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''Antarctopelta oliveroi'' and on its implication for the knowledge of paleobiology of this species is published by Cerda ''et al.'' (2019).


= Cerapods

= * A study on the age of the Kulinda locality (south-eastern Siberia,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
) which yielded fossils of ''Kulindadromeus zabaikalicus'' is published by Cincotta ''et al.'' (2019). * First Photogrammetry, photogrammetric models of the type locality burrow of ''Oryctodromeus cubicularis'' are presented by Wilson & Varricchio (2019). * A study on the taphonomy of fossils of ''Oryctodromeus cubicularis'' is published by Krumenacker ''et al.'' (2019), who also report discovery of new burrows of this dinosaur. * New fossil material of ornithopod dinosaurs is described from the Cretaceous Flat Rocks locality (Wonthaggi Formation,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
) by Herne ''et al.'' (2019), who also revise ''Qantassaurus intrepidus'' and study the phylogenetic relationships of the Victoria (Australia), Victorian ornithopods. * Two minuscule ornithopod Femur, femora, likely belonging to individuals around the point of hatching, are described from the Cenomanian
Griman Creek Formation The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian of the Early-Late Cretaceous.Bell et al., 2019 It is most notable being a major s ...
(Australia) by Kitchener ''et al.'' (2019). * Description of new fossil material of large ornithopod dinosaurs from the Lower Cretaceous localities in El Castellar (Maestrazgo Basin, Teruel,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
), and a study on the implications of these fossils for the knowledge of ornithopod diversity in the Lower Cretaceous of the Iberian Peninsula, is published by Verdú ''et al.'' (2019). * Description of the anatomy of the skeleton of ''Talenkauen santacrucensis'' is published by Rozadilla, Agnolín &
Novas A nova (plural novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", which is Latin for "new") that slowly fades over weeks or months. Causes of the dramat ...
(2019). * A study on the anatomy of the skeleton of ''Macrogryphosaurus gondwanicus'' is published online by Rozadilla, Cruzado-Caballero & Jorge O. Calvo, Calvo (2019). * Skeletal pathologies affecting a subadult specimen of ''
Tenontosaurus ''Tenontosaurus'' ( ; ) is a genus of medium- to large-sized ornithopod dinosaur. It was a relatively medium sized ornithopod, reaching in length and in body mass. It had an unusually long, broad tail, which like its back was stiffened with a n ...
tilletti'' from the Antlers Formation of southeastern Oklahoma are described by Hunt ''et al.'' (2019). * A study on the anatomy and phylogenetic relationships of the ornithopod dinosaurs from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
of Crimea, including ''Riabininohadros weberae'', is published by Averianov & Lopatin (2019). * Redescription of the fossil material of ''Orthomerus dolloi'' and a study on the phylogenetic affinities of this taxon is published online by Madzia, Jagt & Mulder (2019). * A study on patterns and processes of morphological evolution of Hadrosauroidea, hadrosauroid dinosaurs is published by Stubbs ''et al.'' (2019). * A study on the nature of the Fluvial processes, fluvial systems of Laramidia during the Late Cretaceous, as indicated by data from vertebrate and invertebrate fossils from the Kaiparowits Formation of southern Utah, and on the behavior of
hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
dinosaurs over these landscapes, will be published by Crystal ''et al.'' (2019). * Evidence of three‐dimensional preservation of eumelanin‐bearing bodies, dermal cells and blood vessel fragments in a hadrosaur specimen Peabody Museum of Natural History, YPMPU 016969 is presented by Fabbri ''et al.'' (2019). * A study on the osteology and phylogenetic relationships of ''"Tanius laiyangensis"'' is published online by Zhang ''et al.'' (2019). * A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of tibiae of ''Maiasaura peeblesorum'', focusing on the composition, frequency and cortical extent of localized vascular changes, is published by Woodward (2019). * A study on hadrosaurine skulls from the
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
), aiming to assess the influence of
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
on skull morphology, and evaluating proposed synonymy between ''Gryposaurus incurvimanus'' and ''G. notabilis'', is published online by Lowi-Merri & Evans (2019). * Three juvenile specimens of ''Prosaurolophus maximus'', providing new information on the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
of this taxon, are described from the Bearpaw Formation (
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) by Drysdale ''et al.'' (2019). * A study on the structure and contents of a large piece of amber attached to a jaw of a specimen of ''Prosaurolophus maximus'' from the Cretaceous
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
), evaluating the implications of this finding for the knowledge of the habitat and taphonomy of the dinosaur, is published by McKellar ''et al.'' (2019). * A study on the impact of bone tissue structure, early Diagenesis, diagenetic regimes and other Taphonomy, taphonomic variables on the preservation potential of soft tissues in vertebrate fossils, as indicated by data from fossils of ''Edmontosaurus annectens'' from the Standing Rock Hadrosaur Site (
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
Hell Creek Formation The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of ...
, South Dakota), is published by Ullmann, Pandya & Nellermoe (2019), who report the first recovery of osteocytes and vessels from a fossil vertebral centrum and ossified tendons. * A femur of an early juvenile hadrosaurid, probably belonging to the species ''Edmontosaurus annectens'', is described from the Hell Creek Formation (
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
, United States) by Farke & Yip (2019), providing new information on Ontogeny, ontogenetic changes in the skeleton of this dinosaur. * Skull remains of nestling-sized hadrosaurids, probably belonging to the species ''Edmontosaurus annectens'', are described from the Hell Creek Formation (Montana, United States) by Wosik, Goodwin & Evans (2019). * A study of three-dimensionally preserved squamous skin of a member of the genus ''Edmontosaurus'' from the Upper Cretaceous (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
) Wapiti Formation (
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) is published by Barbi ''et al.'' (2019). * The first definitive Lambeosaurinae, lambeosaurine fossil (an isolated skull bone) is described from the Liscomb Bonebed of the Prince Creek Formation (
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) by Takasaki ''et al.'' (2019). * Fossils of a lambeosaurine related to the Eurasian Tsintaosaurini are described from the lower
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
of the Els Nerets locality (eastern Tremp Syncline, northeastern
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) by Conti ''et al.'' (2019). * A study on the microwear of hadrosaur teeth from the La Parrita locality (
Cerro del Pueblo Formation The Cerro del Pueblo Formation is a geological Formation (geology), formation in Coahuila, Mexico whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel et al., ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
) and on its implications for the knowledge of jaw mechanics and feeding ecology of these hadrosaurs is published by Rivera-Sylva ''et al.'' (2019). * A study on bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of ''Psittacosaurus lujiatunensis'' through its
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
is published by Zhao ''et al.'' (2019). * A study on the morphological changes in the braincase of ''Psittacosaurus lujiatunensis'' through its ontogeny, based on data from three specimens from the Lower Cretaceous Yixian Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
), is published by Bullar ''et al.'' (2019). * A three-dimensional virtual
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
of a member of the genus ''Auroraceratops'' is reconstructed on the basis of a well-preserved skull by Zhang ''et al.'' (2019). * Studies on the preservation of fossils of ''Auroraceratops rugosus'', on their stratigraphic provenance, and on the anatomy and phylogenetic relationships of this species are published by Suarez ''et al.'' (2019), Suarez ''et al.'' (2019), Morschhauser ''et al.'' (2019), Li ''et al.'' (2019), Morschhauser ''et al.'' (2019) and Morschhauser ''et al.'' (2019). * A study on the nature of the observed variation in morphology and size of skulls of ''Bagaceratops rozhdestvenskyi'' is published online by Czepiński (2019), who considers the species ''Gobiceratops minutus'', ''Lamaceratops tereschenkoi'', ''Platyceratops tatarinovi'' and ''Magnirostris dodsoni'' to be Synonym (taxonomy), junior synonyms of ''B. rozhdestvenskyi''. * The first postcranial skeleton of ''Bagaceratops'' reported so far is described from the Upper Cretaceous Barun Goyot Formation (
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
) by Kim, Yun & Lee (2019). * A study on the anatomy of the appendicular skeleton of ''
Protoceratops ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
andrewsi'' and on its implications for the knowledge of the locomotor abilities of this species throughout its ontogeny is published by Słowiak, Tereshchenko & Fostowicz-Frelik (2019). * A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
and skeletal growth of ''Avaceratops'' and ''Yehuecauhceratops'' is published online by Hedrick ''et al.'' (2019). * New information on the anatomy of the skeleton of ''Pachyrhinosaurus perotorum'' is presented by Tykoski, Anthony Fiorillo, Fiorillo & Chiba (2019), who also provide a new diagnosis of this species. * A study on the morphological variation of the skulls of specimens of ''Styracosaurus albertensis'' is published online by Holmes ''et al.'' (2019). * A study on the affinities of two Chasmosaurinae, chasmosaurine skulls from the
Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...
(
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
), previously referred to the species ''Chasmosaurus belli'', is published by Campbell ''et al.'' (2019), who transfer the species ''Vagaceratops irvinensis'' to the genus ''Chasmosaurus'', and consider the two studied skulls to be fossils of members of the genus ''Chasmosaurus'' of uncertain specific assignment, with morphology intermediate between ''C. belli'' and ''C. irvinensis''. * A study on the taphonomy of hadrosaurid and ceratopsid fossils from the Scabby Butte locality (St. Mary River Formation;
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
) is published online by Campbell, Ryan & Anderson (2019). *A study on the taxonomic status of ''Teyuwasu barberenai'', in which it was proposed as a second specimen of the Herrerasauridae, herrerasaurid ''Staurikosaurus pricei'' rather than a separate genus and species, is published by Garcia, Müller & Dias-da-Silva (2019). *A study on a putative Sauropodomorpha, sauropodomorph ilium, from Carnian rocks of the Candelária Sequence/Santa Maria Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
), aims to discuss muscle attachment scars, in the context of
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
and phylogeny of Basal (phylogenetics), basal Dinosauriformes, dinosauriforms, focusing in Saturnaliinae, saturnaliine sauropodomorphs is published by Garcia ''et al.'' (2019).


New taxa


Birds


Research

* A study on early bird evolution, aiming to determine their divergence times and evolutionary rates, is published by Zhang & Wang (2019). * A study on the impact of varying oxygen concentrations, global temperatures and air densities on the flight performance of extinct birds and on major diversification events which took place during the evolution of birds is published by Serrano ''et al.'' (2019). * A study aiming to determine whether there is a relationship between the volume of Lacuna (histology), lacunae of osteocytes derived from static osteogenesis and biological parameters such as genome size, body mass, growth rate, metabolic rate or red blood cell size in extant birds is published by Grunmeier & D'Emic (2019), who evaluate the implications of their finding for inferring Physiology, physiological paraments in extinct birds, and potentially other vertebrates, on the basis of osteocyte lacunar volumes. * A study on the expression patterns of the anterior genes ''GLI3, Gli3'' and ''ALX4, Alx4'' in limb buds of emu, chicken and zebra finch embryos, and on their implications for the knowledge of evolution of the Origin of birds#Digit homology, avian digital pattern, is published by Kawahata ''et al.'' (2019). * A study on the diversity of regulatory gene expression profiles of amniote Digit (anatomy), digits, evaluating its implications for the knowledge of the origin of the avian digital pattern, is published by Stewart ''et al.'' (2019). * A study on the total mass of the dentition of Mesozoic birds, and on the impact of the reduction and loss of teeth on total body mass of Mesozoic birds, is published by Zhou, Sullivan & Zhang (2019). * A review of the available evidence of the diet of Mesozoic birds, especially those known from the Lower Cretaceous Jehol Lagerstätte (China), is published by O’Connor (2019). * A study on the early evolution of the digestive system of birds, as indicated by data from paravians from the Jurassic Yanliao Biota and the Cretaceous Jehol Biota (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
), is published online by O'Connor & Zhou (2019). * A study on the early evolution of the diel activity patterns in
diapsid Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago ...
lineages, focusing on the common ancestor branch of living birds, is published by Yu & Wang (2019). * A study on the diversity of melanosome
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
in Iridescence, iridescent feathers of extant birds, and on its implications for inferring iridescence in fossil feathers in general and in
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
birds cf. ''Primotrogon'' and ''Scaniacypselus'' in particular, is published by Nordén ''et al.'' (2019). * A study on the morphology of melanosomes from feathers of extant birds that express non-iridescent structural colour, and on its implications for the possibility of detection of these melanosomes in the fossil record in general and in Crown group#Stem groups, stem group Coraciidae, roller ''Eocoracias'' in particular, is published by Babarović ''et al.'' (2019). * Amino acids are detected in two specimens of fossil feathers from the Cretaceous amber from Myanmar and
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
Baltic amber by McCoy ''et al.'' (2019). * Description of new amber specimens preserving feathers from the Cretaceous of Myanmar, evaluating the implications of these feathers for the knowledge of the development of the rachis-dominated feathers of Mesozoic birds, is published by Carroll, Luis M. Chiappe, Chiappe & Bottjer (2019). * A study on ''Praeornis sharovi'' from the Late Jurassic of Kazakhstan is published by Agnolin, Rozadilla & Carvalho (2019), who interpret the fossil as a tail feather of a Basal (phylogenetics), basal bird. * A geochemical halo of the calamus of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
feather of ''Archaeopteryx lithographica'', detected using Laser-induced fluorescence, Laser-Stimulated Fluorescence, is reported by Kaye ''et al.'' (2019), who also assess the implications of their findings for the identification of this feather; the study is subsequently criticized by Carney, Tischlinger & Shawkey (2020). * A study on the postcranial skeleton of the Specimens of Archaeopteryx#The Berlin specimen, Berlin specimen of ''Archaeopteryx lithographica'', reporting pneumatic structures visible under ultraviolet light and confirming that numerous postcranial bones of ''Archaeopteryx'' were reduced in mass via hollow interiors, is published by Schwarz ''et al.'' (2019). * A comparative study of all named taxa referred to Confuciusornithiformes, taxonomic revision of the group and a study on the phylogenetic relationships of members of the group is published by Wang, O'Connor & Zhou (2019). * Evidence of beak preservation in a referred specimen of ''Confuciusornis sanctus'' (originally the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
of ''Jinzhouornis zhangjiyingia'') is presented by Falk ''et al.'' (2019). * A study on bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of ''Confuciusornis sanctus'', and on its implications for the knowledge of the life history of this species, is published online by Anusuya Chinsamy-Turan, Chinsamy ''et al.'' (2019). * Fully fledged feathering is reported in a hatchling Enantiornithes, enantiornithine specimen from the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
Las Hoyas locality in
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
(first described by Knoll ''et al.'', 2018) by Kate ''et al.'' (2019). * A remarkably well-preserved foot of an enantiornithine bird, accompanied by part of the wing plumage, is described from the Cretaceous amber from Myanmar by Xing ''et al.'' (2019). * A foot of a bird (likely a member of Enantiornithes; made the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
of the species ''Fortipesavis prehendens'' in a later publication), revealing a morphology previously unrecognized in Mesozoic birds, and a range of feathers representing multiple morphotypes are reported for the Cretaceous amber from Myanmar by Xing ''et al.'' (2019). * O'Connor ''et al.'' (2019) describe the integument preserved in four juvenile enantiornithine specimens from the Early Cretaceous Jehol Biota, interpreted by the authors as mid to late immature feathers. * Description of two new specimens of ''Protopteryx fengningensis'' from the Lower Cretaceous Huajiying Formation (China), preserving most of the skeleton and plumage, and providing new information on the anatomy and flight performance of members of this species, is published online by Luis M. Chiappe, Chiappe ''et al.'' (2019). * A study on the bone microstructure of ''Yanornis'', and on its implications for the knowledge of the growth strategy of this bird, is published online by Wang ''et al.'' (2019). * A study on the evolution and function of avian predentary found in Mesozoic Ornithuromorpha, ornithuromorphs, based on data from a specimen of ''Yanornis martini'', is published by Bailleul ''et al.'' (2019). * A study comparing the hindlimb
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
of Hesperornithiformes, hesperornithiforms and modern foot-propelled diving birds is published by Bell, Wu & Luis M. Chiappe, Chiappe (2019). * A study on the paleobiogeography of hesperornithiforms, evaluating its implications for the knowledge of the paleoecology of the Late Cretaceous Western Interior Seaway, is published by Wilson (2019). * A large bird femur referred to the species ''Gargantuavis philoinos'', providing new information on the anatomy of this species, is described from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
of southern
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
by Buffetaut & Angst (2019), who name a new family Gargantuaviidae. * Description of a well-preserved pelvis of ''Gargantuavis'' from the Maastrichtian Sânpetru Formation (
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
), preserving characteristics previously unknown in ''Gargantuavis'' and constituting the first record of this genus outside the area of the Late Cretaceous Ibero-Armorican Island, is published online by Gerald Mayr, Mayr ''et al.'' (2019), who evaluate the implications of this finding for the knowledge of the phylogenetic relationships of ''Gargantuavis''; the study is subsequently criticized by Éric Buffetaut, Buffetaut & Angst (2020). * A study on the evolution of body size of Palaeognathae, palaeognath birds is published by Crouch &
Clarke Clarke is a surname which means "clerk". The surname is of English and Irish origin and comes from the Latin . Variants include Clerk and Clark. Clarke is also uncommonly chosen as a given name. Irish surname origin Clarke is a popular surname i ...
(2019). * A study on the hindlimb anatomy and phylogenetic relationships of ''Palaeotis weigelti'' is published by Gerald Mayr, Mayr (2019). * A study on wing anatomy, body mass, wing surface area, wing span and probable flight parameters of ''Calciavis grandei'' is published online by Torres, Mark Norell, Norell & Clarke (2019). * A study on changes of ostrich eggshell bead diameter throughout the Holocene, testing the proposed relationship between changes of ostrich eggshell bead diameter and the spread of herding in Africa, is published by Miller & Sawchuk (2019). * A study on the taxonomic identification of Rhea (bird), rhea bones from four archaeological sites in the Mendoza Province (Argentina), based on genetic data, is published by Abbona ''et al.'' (2019). * A study aiming to evaluate whether Invasive species in New Zealand, introduced deers and hares fill the same ecological niches as extinct moa birds in New Zealand, as indicated by data from pollen extracted from moa coprolites and mammal feces, is published by Wood & Wilmshurst (2019). * A study on the anatomy of the Bone#Cancellous bone, cancellous bone in the femur, tibiotarsus and fibula of three moa species is published by Bishop, Scofield & Hocknull (2019). * A study on population densities and on the relationship between body mass and population densities in moa birds is published online by Latham ''et al.'' (2019). * A femur of a very large specimen of ''Struthio dmanisensis, "Struthio" dmanisensis'' is described from the Early Pleistocene of the Crimean Peninsula by Zelenkov ''et al.'' (2019), who transfer this species to the genus ''Pachystruthio'' and estimate body mass of this species. * A study on the microstructure of the bones of ''Vegavis iaai'' is published by Garcia Marsà, Agnolín &
Novas A nova (plural novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", which is Latin for "new") that slowly fades over weeks or months. Causes of the dramat ...
(2019). * Eggshells and a small ovoid-shaped egg of Neognathae, neognathous birds, probably members of the family Presbyornithidae, as well as a carpometacarpus of a presbyornithid are described from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of the Glib Zegdou Formation (Algeria) by Garcia ''et al.'' (2019). * A study on the putative Cariamiformes, cariamiform femur from the
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
Sandwich Bluff Member of the
Lopez de Bertodano Formation The Lopez de Bertodano Formation is a geological formation in the James Ross archipelago of the Antarctic Peninsula. The strata date from the end of the Late Cretaceous (upper-lower Maastrichtian stage) to the Danian stage of the lower Paleocene ...
(Vega Island,
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
) reported by Case ''et al.'' (2006) is published by West ''et al.'' (2019), who reinterpret this specimen as a fossil of an unnamed large-bodied member of the genus ''Vegavis''. * A study on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen and other fossils attributed to the species ''Cayaoa bruneti'' is published by De Mendoza & Tambussi (2019), who present a revised diagnosis of this species. * A study on the phylogenetic relationships of ''Cayaoa bruneti'' is published by De Mendoza (2019). * A study on the Cenozoic fossil record of Anatidae, anatids from Eurasia is published by Zelenkov (2019). * A study on the morphology of the postcranial skeleton of the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
-
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Galliformes, galliform ''Palaeortyx'', and on the phylogenetic relationships of this taxon, is published by Zelenkov (2019). * A study on the phylogenetic relationships of extant and fossil members of Strisores is published by Chen ''et al.'' (2019). * Description of new fossil material of ''Pellornis mikkelseni'', providing new information on the anatomy of this species, and a study on the phylogenetic relationships of this species is published by Musser, Ksepka & Field (2019). * A study on the phylogenetic relationships of the adzebills, as indicated by data from near-complete mitochondrial genome sequences, is published by Boast ''et al.'' (2019). * A study on the phylogenetic relationships of the adzebills, as indicated by morphological and molecular data, is published by Musser & Joel Cracraft, Cracraft (2019). * A study on two Humerus, humeri of Rail (bird), rails belonging to the genus ''Dryolimnas'' from the Pleistocene of the Picard Island (Seychelles) is published by Hume & Martill (2019), who interpret these humeri as bones of a flightless rail, and consider them to be evidence of repeated evolution flightlessness in members of the genus ''Dryolimnas'' inhabiting the Aldabra Atoll – before the atoll was completely submerged in the late Pleistocene, as well as after it emerged from the ocean again. * A study on the phylogenetic relationships and evolutionary history of living and extinct flightless lineages of the white-throated rail from the Aldabra Group is published by van de Crommenacker ''et al.'' (2019). * A revision of extinct endemic rails of the Mascarene Islands and a study on their ecology and extinction chronologies is published by Hume (2019). * A study on the taxonomic status of the Canary Islands oystercatcher (''Haematopus meadewaldoi'') is published online by Senfeld ''et al.'' (2019). * A study on the fossil material attributed to the species ''Becassius charadriioides'' is published online by De Pietri, Gerald Mayr, Mayr & Scofield (2019), who assign this species to the family Glareolidae. * A nearly complete tarsometatarsus of the least seedsnipe (''Thinocorus rumicivorus'') is described from the Ensenadan of Argentina by Picasso, De Mendoza & Gelfo (2019). * A study aiming to determine the drivers of the extinction of the great auk, based on data from mitochondrial genome sequences from across its geographic range, is published by Thomas ''et al.'' (2019). * Pedal Phalanx bone, phalanx of a penguin affected by osteomyelitis is described from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of West Antarctica by Jadwiszczak & Rothschild (2019). * A set of skeletal elements of a penguin attributable to the species ''Delphinornis larseni'', providing new information on the anatomy of this species, is described from the Eocene Submeseta Formation (Seymour Island, Antarctica) by Jadwiszczak & Mörs (2019). * The first skull reliably assigned to ''Anthropornis grandis'' is described from the Eocene (
Bartonian The Bartonian is, in the ICS's geologic time scale, a stage or age in the middle Eocene Epoch or Series. The Bartonian Age spans the time between . It is preceded by the Lutetian and is followed by the Priabonian Age. Stratigraphic definitio ...
) Submeseta Formation (Seymour Island, Antarctica) by Acosta Hospitaleche ''et al.'' (2019). * A study on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''Tereingaornis moisleyi'', evaluating the taxonomic validity of this species, is published online by Thomas ''et al.'' (2019). * A fossil humerus of the Magellanic penguin or a relative of this species is described from
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
by Acosta Hospitaleche ''et al.'' (2019), representing the first fossil of a penguin from Uruguay reported so far. * A study on changes in the population size of the Adélie penguin colonies and relative krill abundance in the Prydz Bay (
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
) during the 2nd millennium, as indicated by data from ornithogenic sediment cores from the Vestfold Hills, will be published by Gao ''et al.'' (2019). * A vertebra of a stork similar to the maguari stork is described from the late Pleistocene of the Santa Vitória Formation (Rio Grande do Sul,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Lopes, Pereira & Ferigolo (2019), who evaluate the implications of this finding for reconstructions of local paleoenvironment. * Restudy of a putative bill of an ibis-like bird from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
La Meseta Formation (
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
) described by Jadwiszczak, Gaździcki & Tatur (2008) is published by Agnolin, Bogan & Rozadilla (2019), who consider this specimen to be more likely to be a dorsal spine of a Chimaera, chimaeroid Chondrichthyes, cartilaginous fish. * A study on the body mass evolution in the clade Telluraves, incorporating data from 76 extinct species, is published by Crouch & Mason-Gamer (2019). * A study on the demographic history of the Andean condors in southern South America and on the causes of their survival of late Quaternary megafauna extinctions is published by Perrig ''et al.'' (2019). * Hindlimb bones of an extinct eagle of uncertain phylogenetic placement are described from the late Quaternary of Hispaniola by David Steadman, Steadman, Milan & Mychajliw (2019). * A study on the origin and evolution of the Haast's eagle and the Eyles's harrier, as indicated by complete mitochondrial genome data, is published by Knapp ''et al.'' (2019). * Evidence from Neanderthal-associated sites in Europe indicating that Neanderthals practiced catching the golden eagles is presented by Finlayson ''et al.'' (2019). * Evidence of Châtelperronian Neanderthals using Pes (anatomy), pedal Phalanx bone, phalanges of imperial eagles for symbolic practices is reported from the Cova Foradà site (
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) by Rodríguez-Hidalgo ''et al.'' (2019). * A study on the impact of the climate changes of the last 35,000 years on the long-eared owls and burrowing owls, as indicated by data from fossils from the La Brea Tar Pits, is published by Madan, Donald Prothero, Prothero & Syverson (2019). * A study on the geographical origin and evolutionary history of Coraciiformes, based on data from extant taxa and from fossils, is published by McCullough ''et al.'' (2019). * New skull remains of ''Phorusrhacos longissimus'' are described from the Cerro de los fósiles site in the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Santa Cruz Formation (Argentina) by Degrange ''et al.'' (2019). * A study on the phylogenetic relationships of the Bahaman caracara, based on data from a nearly complete mitochondrial genome recovered from a bone of a member of this species, is published by Oswald ''et al.'' (2019). * A study on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''Calcardea junnei'' is published by Gerald Mayr, Mayr, Philip D. Gingerich, Gingerich & Smith (2019), who reject the interpretation of this species as a heron, and claim that this bird resembled parrot-like taxon ''Vastanavis'' from the early
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. * A study on the identity of a parakeet specimen held at National Museums Scotland, interpreted as most likely originating from Mauritius by Cheke & Jansen (2016), is published by Jones ''et al.'' (2019), who consider this parakeet to be the only known skin specimen of extinct Echo parakeet, Réunion parakeet. * Complete genomic sequence of a specimen of the Carolina parakeet is generated by Gelabert ''et al.'' (2019), who evaluate the implications of their findings for the knowledge of the phylogenetic relationships of this species, its demographic history and adaptation to a toxic diet. * A study on the phylogenetic relationships,
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
and diversification rates of passerine birds throughout their evolutionary history, aiming to evaluate the impact of major events in Earth history on the evolution of passerines, is published by Oliveros ''et al.'' (2019). * Dussex ''et al.'' (2019) sequence whole genomes of the huia and the South Island kōkako, and evaluate whether the loss of genomic diversity played a role in their extinction. * A review of Cretaceous and
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
bird fossils from the James Ross Basin (
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
) is published by Acosta Hospitaleche ''et al.'' (2019). * A study on drivers of bird distribution shifts throughout the Cenozoic is published by Saupe ''et al.'' (2019). * A review of the bird fossil assemblage from the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
locality of Menat, Puy-de-Dôme, Menat (Puy-de-Dôme,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
), including a new fossil specimen with exceptional soft tissue preservation, is published by Gerald Mayr, Mayr, Hervet & Buffetaut (2019). * New bird fossils, including the oldest European record of the Gastornithidae which is temporally well-constrained, are described from the Paleocene localities from the North Sea Basin in Belgium (Maret) and France (Templeuve and Rivecourt-Petit Pâtis) by Mayr & Smith (2019). * A revision of bird fossils from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
(Ypresian) fossil sites of the North American Okanagan Highlands, mainly in British Columbia (
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
), is published by Mayr ''et al.'' (2019), who report, among other findings, a skeleton of a possible member of the family Songziidae, and fossil wings which might constitute the earliest known record of Gaviiformes. * An assemblage of 54 bird bones from early Eocene marine sediments of the Ampe quarry near Egem in Belgium is described by Mayr & Smith (2019). * New Eocene bird fossils, including remains of members of Crown group#Pan-group, Pan-Charadriiformes, a member of Pan-Mirandornithes and a member or a relative of the family Quercymegapodiidae, are described from the Bumban Member of the Naranbulag Formation (
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
) by Hood ''et al.'' (2019). * Flamingo-like and Anatidae, anatid-like fossil bird footprints are described from the Vinchina Formation (Argentina) by Farina ''et al.'' (2019), who name new ichnotaxa ''Phoenicopterichnum lucioi'' and ''P. vinchinaensis''. * A study on the date of extinction of the Tristan moorhen, the Inaccessible Island finch and the Tristan albatross on the main island of the Tristan da Cunha archipelago, aiming to place these extinctions in the context of the changing island ecosystems of the nineteenth and early twentieth centuries, is published by Bond, Carlson & Burgio (2019). * Description of a fossil bird assemblage from the early Pliocene of the Na Burguesa-1 site (Mallorca,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) is published by Torres-Roig ''et al.'' (2019). * A study on the impact of Plio-Pleistocene environmental changes on the bird fauna of New Zealand is published by Rawlence ''et al.'' (2019). * Description of Late Pleistocene and Holocene bird remains from Jerimalai (cave), Jerimalai and Matja Kuru 1 sites in East Timor will be published by Meijer, Louys & O'Connor (2019). * Description of bird remains from the Castelcivita Caves, Grotta di Castelcivita site (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
) and a study on their implications for the knowledge of local environment and human-bird interactions in the Paleolithic is published by Fiore ''et al.'' (2019). * Description of bird remains from the Qesem cave (Israel) dated to between 420 and 200 Year#SI prefix multipliers, ka, and a study on their implications for the knowledge of interactions of birds and humans occupying the site, is published by Blasco ''et al.'' (2019). * A study on the phylogenetic relationships of the dodo and the great auk, as indicated by data from proteins extracted from bone material, is published by Horn ''et al.'' (2019). * A study on bone surface modifications of Pleistocene bird fossils from Mata Menge site (Flores, Indonesia) is published by Meijer ''et al.'' (2019), who report no unambiguous evidence for exploitation of birds from Mata Menge by early hominins. * A study on the impact of human colonization of New Zealand on the diversity dynamics of New Zealand bird fauna is published by Valente, Etienne & Garcia-R (2019).


New taxa


Pterosaurs


Research

* A study on the evolution of vertebral Skeletal pneumaticity, pneumaticity in pterosaurs is published by Buchmann & Rodrigues (2019). * A study on the development of pterosaur embryos is published by Unwin & Deeming (2019). * A study on three pterosaur coprolites from the
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
of Poland, probably produced by Ctenochasmatidae, ctenochasmatids, and on the probable diet of their producers is published by Qvarnström ''et al.'' (2019). * Pterosaur#Pycnofibers, Pycnofibers showing diagnostic features of feathers are reported in two specimens of Anurognathidae, anurognathid pterosaurs (probably belonging either to the genus ''Jeholopterus'' or ''Dendrorhynchoides'') from the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
of
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
by Yang ''et al.'' (2019); the study is subsequently criticized by Unwin & Martill (2020). * Redescription of the holotype specimen of ''Mythunga camara'' is published by Pentland & Poropat (2019). * Partial left Metacarpal bones, metacarpal of a large Ornithocheiridae, ornithocheirid pterosaur is described from the Lower Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
) Wessex Formation (
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
) by Martill & Coram (2019). * New specimen of ''Sinopterus atavismus'' (a nearly complete skeleton) is described from the Lower Cretaceous (
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
) Jiufotang Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Zhang ''et al.'' (2019). * A study on intervertebral foramina in ''Vectidraco'', ''Anhanguera (pterosaur), Anhanguera'' and ''Coloborhynchus'', and on their implications for inferring palaeoecology and locomotion of these pterosaurs, is published by Martin‐Silverstone, Sykes & Darren Naish, Naish (2019). * The first pterosaur postcranial bone (a left ulna) from the
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
Lohan Cura Formation (Argentina) is described by Bellardini & Codorniú (2019). * A study on the melanin content of the soft tissue headcrest of a specimen of ''Tupandactylus imperator'' is published by Pinheiro ''et al.'' (2019). * Alexander Kellner, Kellner ''et al.'' (2019) describe fossils of Pterodactyloidea, pterodactyloid pterosaurs from the Upper Cretaceous Santa Marta Formation and
Snow Hill Island Formation The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
(
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
).


New taxa


Other archosaurs


Research

* Fossils of an immature Lagerpetidae, lagerpetid, representing a new morphotype different from other known lagerpetids, and co-occurring with dinosaurs and Silesauridae, silesaurids, are described from the Candelária Sequence of the
Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. Th ...
Santa Maria Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Garcia ''et al.'' (2019). * A study on the
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
and gross
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
of a growth series of Femur, femora of ''Dromomeron romeri'' is published by Griffin ''et al.'' (2019). * A study reevaluating the taxonomic validity of ''Lagosuchus talampayensis'' is published by Agnolin & Ezcurra (2019). * A study on the microstructure of the long bones (femur and tibiae) of ''Lewisuchus admixtus'' is published by Garcia Marsà, Agnolín &
Novas A nova (plural novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", which is Latin for "new") that slowly fades over weeks or months. Causes of the dramat ...
(2019). * New specimen of ''Lewisuchus admixtus'' is described from the Chañares Formation (Argentina) by Ezcurra ''et al.'' (2019). * A study on the anatomy of the skeleton of ''Asilisaurus kongwe'' is published online by Nesbitt, Langer & Ezcurra (2019). * A study on the anatomy of the braincase of ''Silesaurus opolensis'' is published by Piechowski, Niedźwiedzki & Tałanda (2019). * Coprolites containing beetle remains, most likely produced by ''Silesaurus opolensis'', are described from the Upper Triassic Krasiejów locality (Poland) by Qvarnström ''et al.'' (2019). * A study on bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of ''Sacisaurus agudoensis'' is published by Veiga ''et al.'' (2019). * A study on the phylogenetic relationships of ''Pisanosaurus mertii'' is published by Baron (2019), who interprets this taxon as a likely Silesauridae, silesaurid.


New taxa


References

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