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Tapejaroidea
Tapejaroidea (or tapejaroids) is a group of pterosaurs belonging to the clade Ornithocheiroidea. Tapejaroids lived from the Early to Late Cretaceous periods, with one possible member, ''Tendaguripterus'', extending the fossil range to the Late Jurassic period. Tapejaroidea contains two groups, the Dsungaripteridae and the Azhdarchoidea, which in turn includes the azhdarchids, the group that contains some of the largest flying animals. The group was named by Brazilian paleontologist Alexander Wilhelm Armin Kellner in 1996. Classification Tapejaroidea was named by paleontologist Alexander Kellner from Brazil in 1996, and in 2003 it was given a phylogenetic definition by Kellner himself as the most recent common ancestor of ''Dsungaripterus'', '' Tapejara'' and ''Quetzalcoatlus'', and all their descendants. Tapejaroidea, in Kellner's 2003 study, was recovered as the sister taxon of the Pteranodontoidea, both within the group Ornithocheiroidea, and consisting of the groups Dsungarip ...
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Neoazhdarchia
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionar ...
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Azhdarchoidea
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early Cretaceous, Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and ''Tapejara (pterosaur), Tapejara'', and all its descendants.Unwin, D. M., (20 ...
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Ornithocheiroidea
Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fossil remains found all over the world except Antarctica. Ornithocheiroids were the most advanced group of pterosaurs, as the group includes the clade Azhdarchoidea, of which its members lived until the Maastrichtian stage of the Late Cretaceous, around 66 million years ago. Notable pterosaurs from this group include the pteranodontians ''Pteranodon'' and ''Nyctosaurus'', the ornithocheirid ''Ornithocheirus'', the anhanguerid ''Tropeognathus'', as well as the azhdarchids ''Hatzegopteryx'' and ''Quetzalcoatlus''. Classification The name Ornithocheiroidea was originally defined as an apomorphy-based taxon by Christopher Bennett in 1994. It was given a relationship-based definition in 2003 by Alexander Kellner, who defined it as the least in ...
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Pteranodontoidea
Pteranodontoidea (or pteranodontoids, from Greek meaning "toothless wings") is an extinct clade of ornithocheiroid pterosaurs from the Early to Late Cretaceous (early Valanginian to late Maastrichtian stages) of Asia, Africa, Europe, North America and South America. It was named by Alexander Wilhelm Armin Kellner in 1996. In 2003, Kellner defined the clade as a node-based taxon consisting of the last common ancestor of ''Anhanguera'', ''Pteranodon'' and all its descendants. The clade Ornithocheiroidea is sometimes considered to be the senior synonym of Pteranodontoidea, however it depends on its definition.Unwin, D. M., (2003): On the phylogeny and evolutionary history of pterosaurs. pp. 139-190. — ''in'' Buffetaut, E. & Mazin, J.-M., (eds.): ''Evolution and Palaeobiology of Pterosaurs''. Geological Society of London, Special Publications 217, London, 1-347 Brian Andres (2008, 2010, 2014) in his analyses, converts Ornithocheiroidea using the definition of Kellner (2003) to a ...
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Dsungaripterus Weii
''Dsungaripterus'' is a genus of dsungaripterid pterosaur with an average wingspan of . ''Dsungaripterus'' lived during the Early Cretaceous in what is now China, and its first fossil was found in the Junggar Basin. Description ''Dsungaripterus weii'' had a wing span of . Like most dsungaripteroids it had a rather robust skeleton with thick walls and stouty bodily proportions, suggesting a mostly terrestrial lifestyle. The flight style of these animals is unclear, but it was probably punctuated by abrupt landings and extensive flapping. The skull of ''Dsungaripterus'', long, bore a low bone crest that ran down from the base of the skull to halfway to the beak. ''Dsungaripteruss head and neck were together almost long. Its most notable feature are its long, narrow, upcurved jaws with a pointed tip. It had no teeth in the front part of its jaws, which were probably used to remove prey from cracks in rocks and/or the sandy, muddy inland environments it inhabited. It had knobbly ...
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Dsungaripteridae
Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings. Classification In 1964 Young created a family to place the recently found Chinese genus ''Dsungaripterus''. Later on, also ''Noripterus'' (then now with the name "Phobetor" which was already occupied, therefore the quotation marks) were assigned to the family. In 2003, Alexander Kellner gave the exact definition as a clade:Kellner, A.W.A., 2003. Pterosaur phylogeny and comments on the evolutionary history of the AN group. In: Buffetaut, E., Mazin, J.M. (Eds.), ''Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publication'' 217, 105–137. the group was composed out of the latest common ancestor of ''Dsungaripterus'', ''Noripterus'' and “Phobetor”, and all its descendants. As synapomorphies he gave the next six characteristics: a relatively small eye-socket, which is pla ...
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Dsungaripterus
''Dsungaripterus'' is a genus of dsungaripterid pterosaur with an average wingspan of . ''Dsungaripterus'' lived during the Early Cretaceous in what is now China, and its first fossil was found in the Junggar Basin. Description ''Dsungaripterus weii'' had a wing span of . Like most dsungaripteroids it had a rather robust skeleton with thick walls and stouty bodily proportions, suggesting a mostly terrestrial lifestyle. The flight style of these animals is unclear, but it was probably punctuated by abrupt landings and extensive flapping. The skull of ''Dsungaripterus'', long, bore a low bone crest that ran down from the base of the skull to halfway to the beak. ''Dsungaripteruss head and neck were together almost long. Its most notable feature are its long, narrow, upcurved jaws with a pointed tip. It had no teeth in the front part of its jaws, which were probably used to remove prey from cracks in rocks and/or the sandy, muddy inland environments it inhabited. It had knobbly ...
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Noripterus Complicidens
''Noripterus'' (meaning "lake wing" from Mongolian ''nuur'', "lake" and Greek ''pteron'', "wing") is a genus of dsungaripterid pterodactyloid pterosaur from Lower Cretaceous-age Lianmuqin Formation in the Junggar Basin of Xinjiang, China. It was first named by Yang Zhongjian (also known as C.C. Young in older sources) in 1973. Additional fossil remains have been recovered from Tsagaantsav Svita, Mongolia. Description The first, holotype specimen of ''Noripterus'' ( IVPP V.4062, type locality IVPP 64045) preserved the front part of the skull and lower jaws, vertebrae, and partial limbs and pelvis. ''Noripterus'' was quite similar to the contemporaneous ''Dsungaripterus'', though it was estimated to be a third shorter. It has long narrow neck vertebrae and, like ''Dsungaripterus'', a crest and no teeth in the front of the lower jaw. The teeth that are present are well-developed and spaced fairly far apart. The sharp snout is straight and not pointed upwards as with ''Dsungaripteru ...
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Keresdrakon
''Keresdrakon'' is an extinct genus of tapejaromorph pterosaur from the Goio-Erê Formation of Brazil, which dates back to the Late Cretaceous period (Turonian-Campanian stages), 94 to 71 million years ago. ''Keresdrakon'' contains a single species, ''Keresdrakon vilsoni''. Discovery In 1971, Alexandre Dobruski and his son João Gustavo Dobruski, discovered a fossil site near Cruzeiro do Oeste in Paraná. Only in 2011, paleontologists Paulo César Manzig and Luiz C. Weinschütz visited the location. A bone bed proved to be present with hundreds of specimens of a pterosaur that in 2014 was named ''Caiuajara''. Among them were some bones belonging to a second pterosaur species. These remains, both of ''Caiuajara'' and the new taxon, were prepared by volunteer Vilson Greinert. In 2019, the type species ''Keresdrakon vilsoni'' was named and described by Alexander Wilhelm Armin Kellner, Luiz Carlos Weinschütz, Borja Holgado, Renan Alfredo Machado Bantim and Juliana Manso Sayão. ...
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Banguela Oberlii
''Banguela'' is a genus of azhdarchoid pterosaur from the Early Cretaceous period (Albian stage) of what is now Brazil. Only one species is known, ''Banguela oberlii''. Discovery and naming The Swiss collector Urs Oberli acquired a pterosaur jaw fragment from the Chapada do Araripe. In 2005, this was described by André Jacques Veldmeijer and colleagues, and referred to ''Thalassodromeus sethi''.Veldmeijer, A.J., Signore, M. & Meijer, H.J.M., 2005, "Description of two pterosaur (Pterodactyloidea) mandibles from the lower Cretaceous Santana Formation, Brazil", ''DEINSEA'' 11: 67–86 In 2014 it was named and described by Jaime Headden and Hebert Bruno Nascimento Campos as a separate genus ''Banguela'', with the type species ''Banguela oberlii''. The generic name is a Brazilian Portuguese word for "toothless one", especially used as an affectionate term for elderly women. The specific name honors Oberli. The holotype, NMSG SAO 251093, was probably found in the Romualdo Format ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Chaoyangopterus Zhangi
''Chaoyangopterus'' is a genus of chaoyangopterid pterosaur known from a partial skeleton found in Liaoning, China. ''Chaoyangopterus'' was found in rocks dating back to the Aptian-age Lower Cretaceous Jiufotang Formation of Dapingfang, Chaoyang. Etymology ''Chaoyangopterus'' was named and described in 2003 by Wang Xiaolin and Zhou Zhonghe. The type species is ''Chaoyangopterus zhangi''. The genus name is derived from Chaoyang and a Latinized Greek ''pteron'', "wing". The specific name honors journalist Zhang Wanlian for his efforts in protecting fossil sites. Description ''Chaoyangopterus'' is based on holotype IVPP V13397, which includes the front of the skull, the lower jaws, the neck vertebrae, the shoulder and pelvic girdles, and the limbs. The skull is about long and toothless, and its wingspan is estimated to have been around . Wang and Zhou concluded that it compared most closely to ''Nyctosaurus'' and classified it as a nyctosaurid, although they found that i ...
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