Keresdrakon

''Keresdrakon'' is an extinct genus of tapejaromorph pterosaur from the Goio-Erê Formation of Brazil, which dates back to the Late Cretaceous period (Turonian-Campanian stages), 94 to 71 million years ago. ''Keresdrakon'' contains a single species, ''Keresdrakon vilsoni''.

Discovery

In 1971, Alexandre Dobruski and his son João Gustavo Dobruski, discovered a fossil site near Cruzeiro do Oeste in Paraná. Only in 2011, paleontologists Paulo César Manzig and Luiz C. Weinschütz visited the location. A bone bed proved to be present with hundreds of specimens of a pterosaur that in 2014 was named ''Caiuajara''. Among them were some bones belonging to a second pterosaur species. These remains, both of ''Caiuajara'' and the new taxon, were prepared by volunteer Vilson Greinert.

In 2019, the type species ''Keresdrakon vilsoni'' was named and described by Alexander Wilhelm Armin Kellner, Luiz Carlos Weinschütz, Borja Holgado, Renan Alfredo Machado Bantim and Juliana Manso Sayão. The generic name combines the Keres, evil spirits from Greek mythology, with the Greek ''drakon'', "dragon". The specific name honors Greinert. As the name appeared in an electronic publication, ''Life Science Identifiers'' were needed for its validity. These were E26A65E0-2859-4CD3-B773-B99D568D366C for the genus and 8358C917-6C12-4390-AAB8-37D82723BBCD for the species.

The holotype, CP.V 2069, was found in bonebed C, a sandstone layer. The strata in which the fossils have been discovered were initially thought to belong to the Rio Paraná Formation but later were assigned to the Goio-Erê Formation. The age of this formation is disputed; it could be as old as the Aptian but also as young as the Campanian. The holotype consists of a partial skeleton with skull. It contains the snout, a quadrate bone, the lower jaws, two neck vertebrae, four back vertebrae, the breastbone, belly ribs, the right shoulderblade, two humeri, a radius, two thighbones, a shinebone, the left ilium, the right pubic bone and the left ischium. The skeleton is not articulated. The bones have largely been three-dimensionally preserved, with little compression, and represent a subadult individual. Additional material from the same bonebed was assigned to the taxon as paratypes and referred material.

Phylogeny

The phylogenetic analysis in the description recovered ''Keresdrakon'' as a tapejaromorph azhdarchoid, as the sister taxon to the rest of the clade.

A 2022 study by Rodrigo Pêgas and colleagues recovered ''Keresdrakon'' as the sister taxon to ''Alanqa'' in a basal position among azhdarchoids.

Ecology

The fossils of ''Keresdrakon'' were found in direct association with the tapejarid ''Caiuajara'' and the noasaurid theropod ''Vespersaurus'', confirming that these species were sympatric. Among pterosaurs, this is the first direct evidence of sympatry, i.e. direct association instead of being found in the same stratigraphic unit. Of the three taxa in the bone beds, ''Caiuajara'' is the most common, and ''Keresdrakon'' is the least common. The lizard ''Gueragama'' was also likely part of this paleocommunity. Sedimentological data indicates that the environment of the Goio-Erê Formation was a desert, and the azhdarchid-like rostrum of ''Keresdrakon'' implies it may have been an opportunistic predator in this environment.

Gallery

File:Keresdrakon skull parts.jpg, Skull fragments
File:Keresdrakon forelimbs.jpg, Forelimb fossils
File:Keresdrakon cervicals.jpg, Cervical vertebrae
File:Keresdrakon hip.jpg, Hip fragments
File:Keresdrakon sternum.jpg, Sternum

See also

* List of pterosaur genera
* Timeline of pterosaur research
* Pterosaur size

References

{{Taxonbar, from=Q66661621

Late Cretaceous pterosaurs of South America
Fossil taxa described in 2019
Taxa named by Alexander Kellner
Tapejaromorphs