Tapejaroidea (or tapejaroids) is a group of pterosaurs belonging to the clade

Ornithocheiroidea Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fo ...

. Tapejaroids lived from the

Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

periods, with one possible member, ''

Tendaguripterus ''Tendaguripterus'' was a genus of pterodactyloid pterosaur from the Kimmeridgian to Tithonian-age Upper Jurassic Middle Saurian Beds (Tendaguru Formation) of Tendaguru, Lindi Region, Tanzania. Discovery and naming During the German paleontolog ...

'', extending the fossil range to the

Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...

period. Tapejaroidea contains two groups, the

Dsungaripteridae Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings. Classification In 1964 Young created a family to place the recently ...

and the

Azhdarchoidea Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with ...

, which in turn includes the

azhdarchids Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cret ...

, the group that contains some of the largest flying animals. The group was named by Brazilian paleontologist Alexander Wilhelm Armin Kellner in 1996.

Classification

Tapejaroidea was named by paleontologist

Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...

from Brazil in 1996, and in 2003 it was given a

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...

definition by Kellner himself as the most recent common ancestor of ''

Dsungaripterus ''Dsungaripterus'' is a genus of dsungaripterid pterosaur with an average wingspan of . ''Dsungaripterus'' lived during the Early Cretaceous in what is now China, and its first fossil was found in the Junggar Basin. Description ''Dsungaripte ...

'', '' Tapejara'' and ''

Quetzalcoatlus ''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, ...

'', and all their descendants. Tapejaroidea, in Kellner's 2003 study, was recovered as the sister taxon of the

Pteranodontoidea Pteranodontoidea (or pteranodontoids, from Greek meaning "toothless wings") is an extinct clade of ornithocheiroid pterosaurs from the Early to Late Cretaceous (early Valanginian to late Maastrichtian stages) of Asia, Africa, Europe, North Americ ...

, both within the group

, and consisting of the groups

and

. However, in a phylogenetic analysis made by Jaime Headden and Hebert Bruno Nascimento Campos in 2014, Tapejaroidea was recovered within the Azhdarchoidea, as a clade comprising the families

Tapejaridae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primi ...

and Thalassodromidae. The

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

of their analysis is shown below: More recently, the original definition of Tapejaroidea has been used in a number of phylogenetic analyses conducted in 2019 and 2020, meaning that Tapejaroidea and Pteranodontoidea were once again recovered as the sister taxa and within the larger Ornithocheiroidea.Borja Holgado, Rodrigo V. Pêgas, José Ignacio Canudo, Josep Fortuny, Taissa Rodrigues, Julio Company & Alexander W.A. Kellner, 2019, "On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria", ''Scientific Reports'' 9: 4940 The

below represents the phylogenetic analysis conducted by Kellner and colleagues in 2019, where they recovered Tapejaroidea as the more inclusive group containing both the Dsungaripteridae and the Azhdarchoidea. In 2021, Pêgas ''et al.'' named and officially registered two new clades: Azhdarchomorpha, the most inclusive clade containing ''Azhdarcho'' but not ''Tapejara'' or ''Thalassodromeus'', and Alanqidae, containing ''Alanqa'' but not ''Chaoyangopterus'' or ''Azhdarcho''. Their phylogeny is shown below:

References

{{Taxonbar, from=Q2237060 Ornithocheiroids Maastrichtian extinctions