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Neoazhdarchia
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionar ...
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Thalassodromidae
Thalassodrominae or Thalassodromidae (meaning "sea runners", due to previous misconceptions of skimming behavior; they are now thought to be terrestrial predators) is a group of azhdarchoid pterosaurs from the Cretaceous period. Its traditional members come from Brazil, however, other possible members also come from other places, including the United States, Morocco, and Argentina. Thalassodrominae is considered either to be a subfamily within the pterosaur family Tapejaridae, or as a distinct family, Thalassodromidae, within the clade Neoazhdarchia, closely related to dsungaripterids or azhdarchids. Classification Thalassodrominae traditionally includes only two genera, ''Thalassodromeus'' and ''Tupuxuara'', and was defined to include them and any other descendants of their most recent common ancestor. The classification of thalassodromines is controversial. Some studies, including one by Lü and colleagues in 2008, have found that the thalassodromines are more closely related t ...
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Tapejarid
Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primitive genera were found in China, indicating that the family has an Asian origin. Description Tapejarids were small to medium-sized pterosaurs with several unique, shared characteristics, mainly relating to the skull. Most tapejarids possessed a bony crest arising from the snout (formed mostly by the premaxillary bones of the upper jaw tip). In some species, this bony crest is known to have supported an even larger crest of softer, fibrous tissue that extends back along the skull. Tapejarids are also characterized by their large nasoantorbital fenestra, the main opening in the skull in front of the eyes, which spans at least half the length of the entire skull in this family. Their eye sockets were small and pear-shaped. Studies of tapejarid ...
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Eoazhdarcho
''Eoazhdarcho'' is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is ''Eoazhdarcho liaoxiensis''. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. Etymology The genus name, ''Eoazhdarcho'', combines a Greek ''eos'', "dawn" with the name of the genus ''Azhdarcho'', with the implication it was an early related form of the latter. The specific name, ''liaoxiensis'', refers to the ancient region Liaoxi. Description ''Eoazhdarcho'' is based on holotype GMN-03-11-02, a partial skeleton and lower jaw, and is distinguished from other pterosaurs by the proportions of its bones. The metacarpals are very elongated but the cervical vertebrae and hind limbs are not. It was relatively small by azhdarchoid standards, with a wingspan of about 1.6 meters (5.2 feet). Classification The describers first assigned ''Eoazhdarcho'' t ...
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Quetzalcoatlus Northropi
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas at ...
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Arambourgiania
''Arambourgiania'' is an extinct genus of azhdarchid pterosaur from the Late Cretaceous period (Maastrichtian stage) of Jordan, and possibly the United States.Harrell, T. Lynn Jr.; Gibson, Michael A.; Langston, Wann Jr. (2016). "A cervical vertebra of ''Arambourgiania philadelphiae'' (Pterosauria, Azhdarchidae) from the Late Campanian micaceous facies of the Coon Creek Formation in McNairy County, Tennessee, USA" ''Bull. Alabama Mus. Nat. Hist.'' 33:94–103 ''Arambourgiania'' was among the largest members of its family, the Azhdarchidae, and it is also one of the largest flying animals ever known. The incomplete left ulna of the "Sidi Chennane azhdarchid" from Morocco may have also belonged to ''Arambourgiania''. History of discovery In the early 1940s, a railway worker during repairs on the Amman-Damascus railroad near Russeifa found a two foot long fossil bone. In 1943 this was acquired by the director of a nearby phosphate mine, Amin Kawar, who brought it to the attention ...
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Quetzalcoatlus
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas ...
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Pterodactyloidea
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like ''Nyctosaurus'' and ''Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. "Pteroda ...
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Pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Tapejaroidea
Tapejaroidea (or tapejaroids) is a group of pterosaurs belonging to the clade Ornithocheiroidea. Tapejaroids lived from the Early to Late Cretaceous periods, with one possible member, ''Tendaguripterus'', extending the fossil range to the Late Jurassic period. Tapejaroidea contains two groups, the Dsungaripteridae and the Azhdarchoidea, which in turn includes the azhdarchids, the group that contains some of the largest flying animals. The group was named by Brazilian paleontologist Alexander Wilhelm Armin Kellner in 1996. Classification Tapejaroidea was named by paleontologist Alexander Kellner from Brazil in 1996, and in 2003 it was given a phylogenetic definition by Kellner himself as the most recent common ancestor of ''Dsungaripterus'', '' Tapejara'' and ''Quetzalcoatlus'', and all their descendants. Tapejaroidea, in Kellner's 2003 study, was recovered as the sister taxon of the Pteranodontoidea, both within the group Ornithocheiroidea, and consisting of the groups Dsungarip ...
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Tapejaromorpha
Tapejaromorpha is a group of pterosaurs within the clade Azhdarchoidea. The fossil remains of tapejaromorphs dated back to the Cretaceous period. The Tapejaromorpha was defined in 2014 by Andres and colleagues. They made Tapejaromorpha the most inclusive clade containing ''Tapejara'' but not ''Quetzalcoatlus''. Martill and Naish had previously proposed this same definition for Tapejaridae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primiti ..., but did not formalize it. Andres and colleagues instead defined Tapejaridae more narrowly, as the clade ''Tapejara'' + ''Sinopterus''. References {{pterosaur-stub ...
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Cretornis
''Cretornis'' is a pterosaur genus from the late Cretaceous period (Turonian stage) of what is now the Jizera Formation in the Czech Republic, dating to about 92 million years ago. It only contains a single species, ''Cretornis hlavaci''. Discovery and naming The fossils were discovered in 1880 by workers at a quarry in Zářecká Lhota near the town of Choceň, who were getting gravel to repair a local road. A certain Mrs. Tomková, a croupier from Choceň, then alerted František Hlaváč, a Choceň pharmacist and fossil collector, to the find. He recognised exceptionality of that one, secured the rest of the fossils and then sent them to naturalist Professor Antonín Frič in Prague. In 1881, Antonín Frič identified it as a pterosaur and named it as the type species ''Cretornis Hlaváči''. The generic name is derived from Latin ''creta'', "chalk", in reference to the Cretaceous, and Greek ὄρνις, ''ornis'', "bird", as Frič originally thought that the fossil bones ...
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Microtuban
''Microtuban'' is an extinct genus of azhdarchoid pterosaur known from the Late Cretaceous of northern Lebanon. Discovery ''Microtuban'' is known only from a partially preserved skeleton lacking the skull, holotype SMNK PAL 6595. It was acquired from a local fossil dealer by the ''Staatliches Museum für Naturkunde Karlsruhe'' and was, as can be deduced from the qualities of the stone matrix, collected in the locality of Hjoûla in a marine layer of the Sannine Formation, dating to the early Cenomanian stage of the Late Cretaceous, about 99.6–96 million years ago. The fossil dealer however, indicated it came from a quarry at nearby Hâqel. Regardless, it is only the second pterosaur fossil found in Lebanon and a rare example of a pterosaur from the African continental plate, to which the area in Cretaceous times belonged. In this period the site was located hundreds of miles from the mainland coastline. The specimen consists of the last cervical and first dorsal vertebrae, a ...
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