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Saltasauroidea
Saltasauroidea is a superfamily of titanosaurs named by França and colleagues in 2016 based on their phylogenetic results, for a clade uniting Aeolosaurini and Saltasauridae, as well as the intermediate genera ''Baurutitan'', ''Diamantinasaurus'' and ''Isisaurus''. The group was not defined or discussed in the text, but was supported by Carballido and colleagues in 2022 as a useful designation for subdividing titanosaurs. As there was no discussion about the intentions for the clade, Carballido gave it the definition of all taxa closer to ''Saltasaurus'' than ''Patagotitan'', encompassing half of Eutitanosauria as the sister taxon to the inversely defined Colossosauria. Carballido ''et al.'' placed Nemegtosauridae Nemegtosauridae is a Scientific classification, family of titanosaurian sauropod dinosaurs based on their diplodocidae, diplodocid-like skulls. Only three species are known: ''Nemegtosaurus'', ''Quaesitosaurus'' and possibly ''Tapuiasaurus'', each ... and Saltasauridae ...
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Eutitanosauria
Eutitanosauria is a clade of titanosaurs, encompassing the more derived members of the group and characterized by the absence of the hyposphene-hypantrum articulation and possibly the presence of osteoderms. The group was first named by Sanz and colleagues in 1999, who used it to unite the group of '' Argyrosaurus'', ''Lirainosaurus'', ''Saltasaurus'' and the Peiropolis titanosaur. However, this definition was not used as it made the group equivalent to Saltasauridae, so Saldago redefined it in 2003 to be all titanosaurs closer to ''Saltasaurus'' than ''Epachthosaurus''. This definition created Eutitanosauria as the sister group to Epachthosaurinae (''Epachthosaurus'' but not ''Saltasaurus''), but was problematic due to the variable nature of ''Epachthosaurus''. Eutitanosauria was often broadly similar to Lithostrotia, and has often been unused or unlabelled on phylogenies. Sometimes ''Epachthosaurus'' would be more primitive than ''Malawisaurus'', making Eutitanosauria more encom ...
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Nemegtosaurus
''Nemegtosaurus'' (meaning 'Reptile from the Nemegt') was a sauropod dinosaur from the Late Cretaceous Period of what is now Mongolia. It was named after the Nemegt Basin in the Gobi Desert, where the remains — a single skull — were found. The skull resembles diplodocoids in being long and low, with pencil-shaped teeth. However, recent work has shown that ''Nemegtosaurus'' is in fact a titanosaur, closely related to animals such as ''Saltasaurus'', ''Alamosaurus'' and ''Rapetosaurus''. Discovery and taxonomy The skull of ''Nemegtosaurus'' comes from the same beds as the titanosaur ''Opisthocoelicaudia'', which is known from a skeleton lacking the neck and skull. Originally, the referral of ''Nemegtosaurus'' to Diplodocoidea and ''Opisthocoelicaudia'' to Camarasauridae argued that the two represented different species. Both of these genera represent advanced titanosaurians, however, raising the possibility that the two are in fact the same animal. Relocation of the ''Nemegtosa ...
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Colossosauria
Colossosauria is a clade of titanosaur sauropods from the latest Early Cretaceous through the Late Cretaceous of South America. The group was originally named by Bernardo González-Riga ''et al.'' in 2019 in paleontology, 2019 and defined as the "most inclusive clade containing ''Mendozasaurus neguyelap'' but not ''Saltasaurus loricatus'' or ''Epachthosaurus sciuttoi''". The clade contains different taxa depending on the phylogenetic analysis used, in the defining paper the only subgroups were Rinconsauria and Lognkosauria, but alternate phylogenies published previously had also included various similar titanosaurs such as ''Aeolosaurus'', ''Bonitasaura'', ''Drusilasaura'', ''Overosaurus'' and ''Quetecsaurus''. The phylogenetic analysis of González-Riga ''et al.'' (2019) placed Colossosauria as sister taxa to ''Epachthosaurus'', ''Pitekunsaurus'' and a larger clade including Saltasauridae. Due to the labile position of ''Epachthosaurus'' in titanosaur phylogeny, Carballido and c ...
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Muyelensaurus
''Muyelensaurus'' (meaning "Muyelen lizard", after an indigenous name for the Colorado River in Argentina) is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous of Argentina. It was more slender than other titanosaurs. Fossils have been recovered from the Plottier Formation in the Neuquén province of Patagonia. The type species is ''M. pecheni''.''Muyelensaurus''
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The name ''Muyelensaurus'' first appeared in a 2007 paper by



Aeolosaurini
Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found ''Aeolosaurus'', ''Gondwanatitan'', '' Maxakalisaurus'', '' Panamericansaurus'' and '' Rinconsaurus'' to be aeolosaurids. Aeolosaurini is characterized by several synapomorphies of the caudal vertebrae, such as angled centra, elongate prezygapophyses, and neural arches shifted anteriorly relative to the centra. In life, their tails may have been strongly curved downward as a result of these traits, which may have increased the force exerted by the caudofemoralis longus muscle in retracting the hindlimb. Some aeolosaurins, such as ''Shingopana'' and ''Overosaurus'', were relatively small compared to other titanosaurs, whereas others, such as ''Aeolosaurus maximus'', were large. Phylogeny Aeolosaurini was defined by Franco-Rosas, Salgado, Rosas and Carvalho (2004) as the ste ...
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Epachthosaurus
''Epachthosaurus'' (meaning "heavy lizard") was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America. Discovery and naming The type species, ''E. sciuttoi'', was described by Powell in 1990. The bones assigned to it by Powell in 1990 were, originally, assigned to '' Antarctosaurus sp.'', and then to ''Argyrosaurus superbus?'', before being named as a new taxon. The holotype specimen is MACN-CH 1317, which consists of an incomplete posterior dorsal vertebra. Another specimen, the paratype MACN-CH 18689, consists of a cast of six articulated caudal vertebrae, the partial sacrum, and a fragmentary pubic peduncle from the right ilium. A nearly complete specimen referred to ''Epachthosaurus'', UNPSJB-PV 920, was recovered during field research conducted as part of the project ''Los vertebrados de la Formación Bajo Barreal, Provincia de Chubut, ...
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Opisthocoelicaudia
''Opisthocoelicaudia'' is a genus of sauropod dinosaur of the late Cretaceous, Late Cretaceous Period discovered in the Gobi Desert of Mongolia. The type species is ''Opisthocoelicaudia skarzynskii''. A well-preserved skeleton lacking only the head and neck was unearthed in 1965 by Polish and Mongolian scientists, making ''Opisthocoelicaudia'' one of the best known sauropods from the Late Cretaceous. Tooth marks on this skeleton indicate that large carnivorous dinosaurs had fed on the wikt:carcass, carcass and possibly had carried away the now-missing parts. To date, only two additional, much less complete specimens are known, including part of a pectoral girdle, shoulder and a fragmentary tail. A relatively small sauropod, ''Opisthocoelicaudia'' measured about in length. Like other sauropods, it would have been characterised by a small head sitting on a very long neck and a barrel shaped trunk carried by four column-like legs. The name ''Opisthocoelicaudia'' means "posterior ca ...
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Opisthocoelicaudiinae
Opisthocoelicaudiinae is a subfamily of titanosaurian dinosaurs from the Late Cretaceous. It was named by John McIntosh in 1990. Opisthocoelicaudiines are known from Mongolia, Argentina, and the United States. Two genera were assigned to Opisthocoelicaudiinae by Gonzalez ''et al.'' (2009): ''Alamosaurus'' and ''Opisthocoelicaudia'' (the type genus), a conclusion also found by Díez Díaz ''et al.'' (2018). The hands of opisthocoelicaudiines lacked wrist bones and phalanges. It was suggested by Averianov and Lopatin in 2022 that ''Opisthocoelicaudia'' was not in fact closely related to ''Saltasaurus'', and instead to ''Nemegtosaurus'' and '' Quaesitosaurus'', which are both also Laurasian, as well as isolated teeth from the Turonian of Uzbekistan and the Santonian of Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the ...
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Saltasaurinae
Saltasaurinae is a subfamily of titanosaurian sauropods known from the late Cretaceous period of South America, India and Madagascar. Description Saltasaurines are relatively small sauropods with the general body shape of a small head, long neck, four limbs, and a long tail. They range from the small '' Ibirania'' at around , to the larger ''Neuquensaurus'' at . A currently unnamed fragmentary sauropod from Madagascar may turn out to be a saltasaurine longer than ''Neuquensaurus''. The weight of saltasaurines is very light compared to that of some of the largest dinosaurs. Thomas R. Holtz Jr. found the genera range from around , with ''Saltasaurus'' and an unnamed genus on both extremes, respectively.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> Saltasaurinae is the only known group of sauropods found with armour from almost every species. The most probable reason for the bony stu ...
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Neuquensaurus
''Neuquensaurus'' (meaning "Neuquén lizard") is a genus of saltasaurid sauropod dinosaur that lived in the Late Cretaceous, about 80 million years ago in Argentina and Uruguay in South America. Its fossils were recovered from outcrops of the Anacleto Formation around Cinco Saltos, near the Neuquén river from which its name is derived. History In 1893, Richard Lydekker named ''Titanosaurus australis'', based on a series of caudal vertebrae and limb elements. The remains had been found by Santiago Roth and F. Romero in the Neuquén Province of Argentina at the Neuquén River, and were by Lydekker assigned to a single individual. Six caudal vertebrae, with the inventory numbers MLP Ly 1-6-V-28-1, were the holotype of the species. They had probably been found in a layer of the Anacleto Formation. Some elements that had been referred to ''Titanosaurus australis'' were reassigned to '' Laplatasaurus araukanicus'' by Friedrich von Huene in 1929. The same year, von Huene named a ''Ti ...
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Lognkosauria
Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and unusually thick cervical rib loops on their neck vertebrae, their extremely robust neck neural spines, the relatively narrow neural canal, and their huge vaulted neural arches. They also had very wide dorsal vertebrae with wing-like side processes, and extremely wide rib cages. Their dorsal side processes are also fairly in-line with the level of the neural canal, instead of being attached further up the neural arch as in lithostrotians. Skull material from ''Malawisaurus'', the sister taxon to Lognkosauria, indicates that lognkosaurians at least began with the big-nosed, rounded head shape of earlier titanosaurs and more basal macronarians. Classification Lognkosauria was defined as the clade encompassing the most recent common ancestor o ...
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Diamantinasauria
Diamantinasauria is an extinct clade of non- lithostrotian titanosaurian sauropod dinosaurs, known from the early Late Cretaceous (Cenomanian-Turonian) of South America and Australia. It was named by Poropat and colleagues in 2021, and contains four genera: ''Australotitan'', ''Savannasaurus'' and ''Diamantinasaurus'' from the Winton Formation of Queensland, as well as ''Sarmientosaurus'' from the Bajo Barreal Formation of Patagonia. The existence of the clade indicates connectivity between Australia and South America via Antarctica during the Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ... period. Though Diamantinasauria has been recovered consistently as a clade in the phylogenetic analyses of Poropat and colleagues, its placement within Titanosauria has fluctuate ...
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