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Eutitanosauria
Eutitanosauria is a clade of titanosaurs, encompassing the more derived members of the group and characterized by the absence of the hyposphene-hypantrum articulation and possibly the presence of osteoderms. The group was first named by Sanz and colleagues in 1999, who used it to unite the group of '' Argyrosaurus'', ''Lirainosaurus'', ''Saltasaurus'' and the Peiropolis titanosaur. However, this definition was not used as it made the group equivalent to Saltasauridae, so Saldago redefined it in 2003 to be all titanosaurs closer to ''Saltasaurus'' than ''Epachthosaurus''. This definition created Eutitanosauria as the sister group to Epachthosaurinae (''Epachthosaurus'' but not ''Saltasaurus''), but was problematic due to the variable nature of ''Epachthosaurus''. Eutitanosauria was often broadly similar to Lithostrotia, and has often been unused or unlabelled on phylogenies. Sometimes ''Epachthosaurus'' would be more primitive than ''Malawisaurus'', making Eutitanosauria more encom ...
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Titanosauria
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ran ...
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Titanosaurs
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ran ...
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Saltasauroidea
Saltasauroidea is a superfamily of titanosaurs named by França and colleagues in 2016 based on their phylogenetic results, for a clade uniting Aeolosaurini and Saltasauridae, as well as the intermediate genera ''Baurutitan'', ''Diamantinasaurus'' and ''Isisaurus''. The group was not defined or discussed in the text, but was supported by Carballido and colleagues in 2022 as a useful designation for subdividing titanosaurs. As there was no discussion about the intentions for the clade, Carballido gave it the definition of all taxa closer to ''Saltasaurus'' than ''Patagotitan'', encompassing half of Eutitanosauria as the sister taxon to the inversely defined Colossosauria. Carballido ''et al.'' placed Nemegtosauridae Nemegtosauridae is a Scientific classification, family of titanosaurian sauropod dinosaurs based on their diplodocidae, diplodocid-like skulls. Only three species are known: ''Nemegtosaurus'', ''Quaesitosaurus'' and possibly ''Tapuiasaurus'', each ... and Saltasauridae ...
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Patagotitan
''Patagotitan'' is a genus of titanosaurian sauropod dinosaur from the Cerro Barcino Formation in Chubut Province, Patagonia, Argentina. The genus contains a single species known from multiple individuals: ''Patagotitan mayorum'', first announced in 2014 and then validly named in 2017 by José Carballido, Diego Pol, and colleagues. Contemporary studies estimated the length of the type specimen, a young adult, at with a weight estimated at , later revised to . Discovery Remains of ''Patagotitan mayorum'', a part of a lower thighbone, were initially discovered in 2008 by a farm laborer, Aurelio Hernández, in the desert near La Flecha, Argentina, about west of Trelew. Excavation was done by palaeontologists from the Museum of Paleontology Egidio Feruglio. The lead scientists on the excavation were Jose Luis Carballido and Diego Pol, with partial funding from The Jurassic Foundation. Between January 2013 and February 2015, seven paleontological field expeditions were carri ...
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Saltasauroidea
Saltasauroidea is a superfamily of titanosaurs named by França and colleagues in 2016 based on their phylogenetic results, for a clade uniting Aeolosaurini and Saltasauridae, as well as the intermediate genera ''Baurutitan'', ''Diamantinasaurus'' and ''Isisaurus''. The group was not defined or discussed in the text, but was supported by Carballido and colleagues in 2022 as a useful designation for subdividing titanosaurs. As there was no discussion about the intentions for the clade, Carballido gave it the definition of all taxa closer to ''Saltasaurus'' than ''Patagotitan'', encompassing half of Eutitanosauria as the sister taxon to the inversely defined Colossosauria. Carballido ''et al.'' placed Nemegtosauridae Nemegtosauridae is a Scientific classification, family of titanosaurian sauropod dinosaurs based on their diplodocidae, diplodocid-like skulls. Only three species are known: ''Nemegtosaurus'', ''Quaesitosaurus'' and possibly ''Tapuiasaurus'', each ... and Saltasauridae ...
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Colossosauria
Colossosauria is a clade of titanosaur sauropods from the latest Early Cretaceous through the Late Cretaceous of South America. The group was originally named by Bernardo González-Riga ''et al.'' in 2019 in paleontology, 2019 and defined as the "most inclusive clade containing ''Mendozasaurus neguyelap'' but not ''Saltasaurus loricatus'' or ''Epachthosaurus sciuttoi''". The clade contains different taxa depending on the phylogenetic analysis used, in the defining paper the only subgroups were Rinconsauria and Lognkosauria, but alternate phylogenies published previously had also included various similar titanosaurs such as ''Aeolosaurus'', ''Bonitasaura'', ''Drusilasaura'', ''Overosaurus'' and ''Quetecsaurus''. The phylogenetic analysis of González-Riga ''et al.'' (2019) placed Colossosauria as sister taxa to ''Epachthosaurus'', ''Pitekunsaurus'' and a larger clade including Saltasauridae. Due to the labile position of ''Epachthosaurus'' in titanosaur phylogeny, Carballido and c ...
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Menucocelsior
''Menucocelsior'' (meaning "the taller one from Salitral Ojo del Agua") is a genus of medium-sized titanosaurian sauropod dinosaur from the Late Cretaceous Allen Formation of Argentina. The type and only species is ''Menucocelsior arriagadai''. Discovery The Salitral Ojo del Agua area of Río Negro, Argentina was home to an entire sauropod fauna during the late Cretaceous period. Fossil material belonging to saltasaurines, aeolosaurines, and other titanosaurs have been discovered. In 2022, Rolando ''et al.'' designated one specimen as the type species for a new titanosaurian taxon, ''Menucocelsior arriagadai.'' The generic name combines the Mapudungun ''menuco'', meaning "watering hole", a translation of Ojo del Agua, with the Latin ''celsior'', meaning "higher" or "major." The specific name honours "Beto" Arriagada, the owner of the ranch where the discoveries were made. Description and classification The holotype, MPCN-PV-798, consists of seventeen caudal vertebrae and seve ...
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Futalognkosaurus
''Futalognkosaurus'' ( ; meaning "giant chief lizard") is a genus of titanosaurian dinosaur. The herbivorous ''Futalognkosaurus'' lived approximately 87 million years ago in the Portezuelo Formation, in what is now Argentina, of the Coniacian stage of the late Cretaceous Period. The fish and fossilized leaf debris on the site, together with other dinosaur remains, suggest a warm tropical climate in Patagonia during this period. Discovery Fossils of ''Futalognkosaurus'' were found in the Neuquén province of Argentina in 2000, and were scientifically described in 2007, alongside sauropods, ''Megaraptor'', ''Unenlagia'', iguanodonts, peirosaurids, and pterosaurs. The holotype was found near the margins of a river. The holotype was hypothesised to have been washed into the river after its death. Due to its giant size the carcass likely acted as a barrier, altering thr colurse of the river around it, lasting long enough for fish and bivalves to live in ir, and eventually creating an ...
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Rinconsaurus
''Rinconsaurus'' is a genus of titanosaur sauropod dinosaur from the Late Cretaceous in what is now Argentina. The type species, ''Rinconsaurus caudamirus'', was described by Calvo and Riga in 2003, and is based on three partial skeletons. Description Like all sauropods, ''Rinconsaurus'' was a large long-necked quadrupedal animal, with a long, whip-like tail and four pillar-like legs. ''Rinconsaurus'' was an unusually slender sauropod. Although fossil discoveries are incomplete, and no complete necks or heads have been found, fully grown ''Rinconsaurus'' are estimated to have been 11 meters (36 ft) long and approximately 2.5 meters (8 ft) high at the shoulder. The body mass of ''Rinconsaurus'' has been estimated to be between 3.21 and 5.39 tonnes. Discovery and species Fossils of ''Rinconsaurus'' were discovered in 1997 by Gabriel Benítezin strata belonging to the Bajo de la Carpa Formation near Rincón de los Sauces, in the Neuquen province of Argentina. ...
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Rinconsauria
Rinconsauria is an extinct clade of giant titanosaurian sauropods known from the late Cretaceous period of Argentina. Systematics Rinconsauria was coined by Calvo et al. (2007) to include their new titanosaur ''Muyelensaurus'' and the previously described ''Rinconsaurus''. Santucci and Arruda-Campos (2011) recovered Rinconsauria as part of Aeolosaurini, as did Franca et al. (2016) and Silva et al. (2019). However, cladistic analyses by Gonzalez-Riga et al. (2019) and Mannion et al. (2019) found ''Aeolosaurus'' to be in a phylogenetically disparate position than Rinconsauria, with Rinconsauria as sister to Lognkosauria Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and u ... in the clade Colossosauria.Mannion, P.D.; Upchurch, P.; Jin, X.; Zheng, W. (2019). New information on the Cretaceo ...
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Diamantinasauria
Diamantinasauria is an extinct clade of non- lithostrotian titanosaurian sauropod dinosaurs, known from the early Late Cretaceous (Cenomanian-Turonian) of South America and Australia. It was named by Poropat and colleagues in 2021, and contains four genera: ''Australotitan'', ''Savannasaurus'' and ''Diamantinasaurus'' from the Winton Formation of Queensland, as well as ''Sarmientosaurus'' from the Bajo Barreal Formation of Patagonia. The existence of the clade indicates connectivity between Australia and South America via Antarctica during the Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ... period. Though Diamantinasauria has been recovered consistently as a clade in the phylogenetic analyses of Poropat and colleagues, its placement within Titanosauria has fluctuate ...
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Lognkosauria
Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and unusually thick cervical rib loops on their neck vertebrae, their extremely robust neck neural spines, the relatively narrow neural canal, and their huge vaulted neural arches. They also had very wide dorsal vertebrae with wing-like side processes, and extremely wide rib cages. Their dorsal side processes are also fairly in-line with the level of the neural canal, instead of being attached further up the neural arch as in lithostrotians. Skull material from ''Malawisaurus'', the sister taxon to Lognkosauria, indicates that lognkosaurians at least began with the big-nosed, rounded head shape of earlier titanosaurs and more basal macronarians. Classification Lognkosauria was defined as the clade encompassing the most recent common ancestor o ...
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