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Colossosauria
Colossosauria is a clade of titanosaur sauropods from the latest Early Cretaceous through the Late Cretaceous of South America. The group was originally named by Bernardo González-Riga ''et al.'' in 2019 in paleontology, 2019 and defined as the "most inclusive clade containing ''Mendozasaurus neguyelap'' but not ''Saltasaurus loricatus'' or ''Epachthosaurus sciuttoi''". The clade contains different taxa depending on the phylogenetic analysis used, in the defining paper the only subgroups were Rinconsauria and Lognkosauria, but alternate phylogenies published previously had also included various similar titanosaurs such as ''Aeolosaurus'', ''Bonitasaura'', ''Drusilasaura'', ''Overosaurus'' and ''Quetecsaurus''. The phylogenetic analysis of González-Riga ''et al.'' (2019) placed Colossosauria as sister taxa to ''Epachthosaurus'', ''Pitekunsaurus'' and a larger clade including Saltasauridae. Due to the labile position of ''Epachthosaurus'' in titanosaur phylogeny, Carballido and c ...
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Rinconsauria
Rinconsauria is an extinct clade of giant titanosaurian sauropods known from the late Cretaceous period of Argentina. Systematics Rinconsauria was coined by Calvo et al. (2007) to include their new titanosaur ''Muyelensaurus'' and the previously described ''Rinconsaurus''. Santucci and Arruda-Campos (2011) recovered Rinconsauria as part of Aeolosaurini, as did Franca et al. (2016) and Silva et al. (2019). However, cladistic analyses by Gonzalez-Riga et al. (2019) and Mannion et al. (2019) found ''Aeolosaurus'' to be in a phylogenetically disparate position than Rinconsauria, with Rinconsauria as sister to Lognkosauria Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and u ... in the clade Colossosauria.Mannion, P.D.; Upchurch, P.; Jin, X.; Zheng, W. (2019). New information on the Cretaceo ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Bonitasaura
''Bonitasaura'' is a titanosaurian dinosaur hailing from uppermost layers of the Late Cretaceous (Santonian) Bajo de la Carpa Formation, Neuquén Group of the eastern Neuquén Basin, located in Río Negro Province, Northwestern Patagonia, Argentina. The remains, consisting of a partial sub-adult skeleton jumbled in a small area of fluvial sandstone, including lower jaw with teeth, partial vertebrae series and limb bones, were described by Sebastian Apesteguía in 2004. The genus name ''Bonitasaura'' refers to the fossil quarry's name, "La Bonita", while the name of the type species, ''B. salgadoi'', pays homage to Leonardo Salgado, a renowned Argentine paleontologist.Gallina, P. A. (2011)Notes on the axial skeleton of the titanosaur Bonitasaura salgadoi (Dinosauria-Sauropoda).''Anais da Academia Brasileira de Ciências'', ''83''(1), 235-246. Description ''Bonitasaura'' measured in length, and had a skull similar to another group of sauropods, the diplodocids. The lower jaw ha ...
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Chucarosaurus
''Chucarosaurus'' (meaning "indomitable reptile") is an extinct genus of titanosaurian dinosaur from the Late Cretaceous (middle Cenomanian–lower Turonian) Huincul Formation of Argentina. The genus contains a single species, ''C. diripienda'', known from various limb bones. Discovery and naming The ''Chucarosaurus'' holotype specimen, MPCA PV 820, was discovered in sediments of the Huincul Formation, dated to the middle Cenomanian–lower Turonian ages of the late Cretaceous period, at Pueblo Blanco Natural Reserve in Río Negro Province, Argentina. The specimen consists of a complete left humerus, partial left radius, complete left metacarpal II, left ischium, partial left femur and fibula, partial right tibia, and partial indeterminate metapodial, all of which belong to one individual. An additional specimen (MPCA PV 821), consisting of a left femur and tibia, was also referred as a paratype. In 2023, Agnolin ''et al''. described ''Chucarosaurus diripienda'', a new ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Aeolosaurus
''Aeolosaurus'' (; "Aeolus' lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. ''Aeolosaurus'' is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of ''Aeolosaurus rionegrinus'' consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species ''A. maximus'' was transferred over to the new genus '' Arrudatitan'' in 2021. Etymology This dinosaur is named after the Greek mythological figure Aeolus, Keeper of the Winds in Homer's Odyssey, because of the frequent wind ...
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Mendozasaurus Neguyelap
''Mendozasaurus'' is a genus of titanosaurian sauropod dinosaur. It was a member of Titanosauria, which were massive sauropods that were common on the southern landmasses during the Cretaceous. It is represented by several partial skeletons from a single locality within the Coniacian (lower Upper Cretaceous) Sierra Barrosa Formation in the south of Mendoza Province, northern Neuquén Basin, Argentina. The type species, ''Mendozasaurus neguyelap'', was described by Argentine paleontologist Bernardo Javier González Riga in 2003. ''Mendozasaurus'' is the first dinosaur named from Mendoza Province, Argentina, for which it was named. Description This species belonged to the discovered clade Lognkosauria, a transitional group of titanosaurs which included the gigantic '' Futalognkosaurus'' and ''Puertasaurus''. Like both of these animals, ''Mendozasaurus'' had a long neck with very wide cervical neural spines. Holtz estimated its length at . In 2010 Gregory S. Paul estimated ''Mend ...
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Saltasaurus Loricatus
''Saltasaurus'' (which means "lizard from Salta") is a genus of saltasaurid dinosaur of the Late Cretaceous period of Argentina. Small among sauropods, though still heavy by the standards of modern creatures, ''Saltasaurus'' was characterized by a short neck and stubby limbs. It was the first genus of sauropod known to possess armour of bony plates embedded in its skin. Such small bony plates, called osteoderms, have since been found on other titanosaurians. Discovery The fossils of ''Saltasaurus'' were excavated by José Bonaparte, Martín Vince and Juan C. Leal between 1975 and 1977 at the Estancia "El Brete". The find was in 1977 reported in the scientific literature. ''Saltasaurus'' was named and described by Bonaparte and Jaime E. Powell in 1980. The type species is ''Saltasaurus loricatus''. Its generic name is derived from Salta Province, the region of north-west Argentina where the first fossils were recovered. The specific name means "protected by small armoured pla ...
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Epachthosaurus Sciuttoi
''Epachthosaurus'' (meaning "heavy lizard") was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America. Discovery and naming The type species, ''E. sciuttoi'', was described by Powell in 1990. The bones assigned to it by Powell in 1990 were, originally, assigned to '' Antarctosaurus sp.'', and then to '' Argyrosaurus superbus?'', before being named as a new taxon. The holotype specimen is MACN-CH 1317, which consists of an incomplete posterior dorsal vertebra. Another specimen, the paratype MACN-CH 18689, consists of a cast of six articulated caudal vertebrae, the partial sacrum, and a fragmentary pubic peduncle from the right ilium. A nearly complete specimen referred to ''Epachthosaurus'', UNPSJB-PV 920, was recovered during field research conducted as part of the project ''Los vertebrados de la Formación Bajo Barreal, Provincia de Chubut ...
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Pitekunsaurus
''Pitekunsaurus'' is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Anacleto Formation of Neuquén, Argentina. It was described by L. Filippi and A. Garrido in 2008. The type species is ''P. macayai''. The generic name is derived from Mapudungun ''pitekun'', meaning "to discover", the epitheton honours the discoverer, oil company explorer Luis Macaya, who found the fossil in April 2004. Classification ''Pitekunsaurus'' appears to be a genus of aeolosaurid, most closely related to the genera ''Gondwanatitan'' and ''Aeolosaurus''. Its braincase closely resembles that of the aeolosaurid ''Muyelensaurus'' as well as the titanosaurs ''incertae sedis'' '' Vahiny'' and ''Jainosaurus''. It cannot be compared to the more closely related ''Aeolosaurus'' and ''Gondwanatitan ''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in t ...
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Drusilasaura
''Drusilasaura'' is an extinct genus of possible lognkosaurian titanosaur sauropod dinosaur which lived during the late Cretaceous (Cenomanian-Turonian stage) of Santa Cruz Province of southern Patagonia, Argentina. ''Drusilasaura'' is known from the holotype MPM-PV 2097/1 to 2097/19, a partial skeleton including four dorsal vertebrae, a sacral vertebra, six caudal vertebrae, a left scapula, dorsal rib fragments and other fragments. It was found by palaeontologist Marcelo Tejedor searching fossil mammals, in layers of the Upper Member of the Bajo Barreal Formation, on the María Aike Ranch owned by the Ortiz de Zárate family. A team from the '' Laboratorio de Paleontología de Vertebrados'' of the ''Universidad Nacional de la Patagonia San Juan Bosco'' subsequently collected the remains.Navarrete, C.; Casal, G. & Martínez, R., (2008), "Nuevos materiales de Titanosauridae (Lognkosauria?) de la Formación Bajo Barreal, Cretácico Superior, Santa Cruz, Argentina". In: ''CON ...
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Overosaurus
''Overosaurus'' (meaning "Overo lizard", after the Cerro Overo locality) is an extinct genus of sauropod dinosaurs, containing only a single species, ''Overosaurus paradasorum''. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia (in southern Argentina). ''Overosaurus paradasorum'' was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately . It was a ground-dwelling herbivore. Discovery and naming The only known specimen of ''Overosaurus paradasorum'' was recovered at the Cerro Overo locality in Patagonia, Argentina. The specimen was collected in 2002 by researchers from the Museo Carmen Funes, the Museo Argentino Urquiza, and the Natural History Museum of Los Angeles County, in terrestrial sediments deposited during the Campanian stage of the Late Cretaceous period, approximately 84 to 78 million years ago. This speci ...
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