Mendozasaurus Neguyelap
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''Mendozasaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
titanosaurian Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
. It was a member of
Titanosauria Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
, which were massive sauropods that were common on the southern landmasses during the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
. It is represented by several partial skeletons from a single locality within the
Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...
(lower Upper Cretaceous)
Sierra Barrosa Formation The Sierra Barrosa Formation is a geologic formation of the Neuquén Basin in the northern Patagonian provinces of Mendoza and Neuquén. The formation dates to the Late Cretaceous, middle to late Coniacian, and belongs to the Río Neuquén Subg ...
in the south of Mendoza Province, northern
Neuquén Basin Neuquén Basin ( es, Cuenca Neuquina) is a sedimentary basin covering most of Neuquén Province in Argentina. The basin originated in the Jurassic and developed through alternating continental and marine conditions well into the Tertiary. The bas ...
, Argentina. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Mendozasaurus neguyelap'', was described by Argentine paleontologist Bernardo Javier González Riga in 2003. ''Mendozasaurus'' is the first dinosaur named from
Mendoza Province Mendoza, officially Province of Mendoza, is a province of Argentina, in the western central part of the country in the Cuyo region. It borders San Juan to the north, La Pampa and Neuquén to the south, San Luis to the east, and the republic o ...
,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, for which it was named.


Description

This species belonged to the discovered
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Lognkosauria Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and u ...
, a transitional group of titanosaurs which included the gigantic '' Futalognkosaurus'' and ''
Puertasaurus ''Puertasaurus'' is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Ar ...
''. Like both of these animals, ''Mendozasaurus'' had a long neck with very wide cervical neural spines. Holtz estimated its length at . In 2010
Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...
estimated ''Mendozasaurus'' at 20 meters (65.6 feet) in length and 16 tonnes (17.6 short tons) in weight.


Classification

A phylogenetic analysis conducted by González Riga and colleagues in 2018 recovered ''Mendozasaurus'' as the most basal member of Lognkosauria, including ''Futalognkosaurus'' and the gigantic titanosaurs ''
Argentinosaurus ''Argentinosaurus'' is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, ''Argentinosaurus'' is one of the largest known land animals o ...
'', ''
Notocolossus ''Notocolossus'' is a genus of titanosaurian sauropod dinosaur from late Cretaceous strata of Mendoza Province, Argentina. Discovery and naming A fossil of a large sauropod was discovered by the Argentine paleontologist Dr. Bernardo Javier ...
'', ''
Patagotitan ''Patagotitan'' is a genus of titanosaurian sauropod dinosaur from the Cerro Barcino Formation in Chubut Province, Patagonia, Argentina. The genus contains a single species known from multiple individuals: ''Patagotitan mayorum'', first announce ...
'' and ''Puertasaurus''. The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
shows the position of ''Mendozasaurus'' in
Lognkosauria Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkosaurians can be distinguished from other titanosaurs by the wide and u ...
according to González Riga and colleagues, 2018.


References


Further reading

* González Riga, B.J. (2003). "A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina". ''Amehginiana'' 40: 155–172. * González Riga, B.J. (2005). "Nuevos restos fósiles de ''Mendozasaurus neguyelap'' (Sauropoda: Titanosauridae) del Cretácico Tardío de Mendoza, Argentina". ''Ameghiniana'' 42 (3): 535–538. * González Riga, B.J. and Astini, R. (2007).
Fossil preservation of large titanosaur sauropods in overbank fluvial facies: a case study in the Cretaceous of Argentina
. ''Journal of South American Earth Sciences'' 23: 290–303.


External links


Sauropod dinosaurs
Complementary information on South American titanosaurs Lithostrotians Coniacian life Late Cretaceous dinosaurs of South America Cretaceous Argentina Fossils of Argentina Neuquén Basin Fossil taxa described in 2003 {{Sauropodomorph-stub