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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Titanosauridae
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a '' nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ...
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Epachthosaurus
''Epachthosaurus'' (meaning "heavy lizard") was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America. Discovery and naming The type species, ''E. sciuttoi'', was described by Powell in 1990. The bones assigned to it by Powell in 1990 were, originally, assigned to '' Antarctosaurus sp.'', and then to ''Argyrosaurus superbus?'', before being named as a new taxon. The holotype specimen is MACN-CH 1317, which consists of an incomplete posterior dorsal vertebra. Another specimen, the paratype MACN-CH 18689, consists of a cast of six articulated caudal vertebrae, the partial sacrum, and a fragmentary pubic peduncle from the right ilium. A nearly complete specimen referred to ''Epachthosaurus'', UNPSJB-PV 920, was recovered during field research conducted as part of the project ''Los vertebrados de la Formación Bajo Barreal, Provincia de Chubut, ...
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Sarmientosaurus
''Sarmientosaurus'' is a genus of titanosaurian sauropod dinosaur belonging to the Titanosauria. It lived in what is now South America, specifically Argentina, during the Upper Cretaceous Period about 95 million years ago. The type species is ''Sarmientosaurus musacchioi''. Discovery In 1997, paleontologist Rubén D.F. Martínez, at the ''Estancia Laguna Palacios'' of the Goicoechea family in Chubut province, discovered a sauropod skull. This proved to be connected to the first few cervical vertebrae. In 2016, the type species ''Sarmientosaurus musacchioi'' was named and described by Rubén Darío Francisco Martínez, Matthew Carl Lamanna, Fernando Emilio Novas, Ryan C. Ridgely, Gabriel Andrés Casal, Javier E. Martínez, Javier R. Vita and Lawrence M. Witmer. The generic name refers to the town of Sarmiento. The specific name honours the late Eduardo Musacchio, an educator at the ''Universidad Nacional de la Patagonia San Juan Bosco''. The Life Science Identifiers are 5 ...
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Volgatitan
''Volgatitan'' (meaning "Volga giant") is a genus of titanosaurian sauropod dinosaur from the Early Cretaceous of the Ulyanovsk Oblast, Russia. The type species, type and only species is ''Volgatitan simbirskiensis'', known from seven from a single individual. It is the oldest known titanosaur from the northern hemisphere, and is considered important for being related to the Lognkosauria, a group known only from South America later in the Late Cretaceous. It was first species description, described in November 2018 by Russian palaeontologists Alexander Averianov and Vladimir Efimov. Classification Amerianov and Efimov recovered ''Volgatitan'' as a lithostrotian titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th .... They found Lithostrotia to be divided into two main lineag ...
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Eutitanosauria
Eutitanosauria is a clade of titanosaurs, encompassing the more derived members of the group and characterized by the absence of the hyposphene-hypantrum articulation and possibly the presence of osteoderms. The group was first named by Sanz and colleagues in 1999, who used it to unite the group of '' Argyrosaurus'', ''Lirainosaurus'', ''Saltasaurus'' and the Peiropolis titanosaur. However, this definition was not used as it made the group equivalent to Saltasauridae, so Saldago redefined it in 2003 to be all titanosaurs closer to ''Saltasaurus'' than ''Epachthosaurus''. This definition created Eutitanosauria as the sister group to Epachthosaurinae (''Epachthosaurus'' but not ''Saltasaurus''), but was problematic due to the variable nature of ''Epachthosaurus''. Eutitanosauria was often broadly similar to Lithostrotia, and has often been unused or unlabelled on phylogenies. Sometimes ''Epachthosaurus'' would be more primitive than ''Malawisaurus'', making Eutitanosauria more encom ...
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Tapuiasaurus
''Tapuiasaurus'' (meaning " Tapuia lizard") is a genus of titanosaur which lived during the Lower Cretaceous period (Aptian age) in what is now Minas Gerais, Brazil. Its fossils, including a partial skeleton with a nearly complete skull, have been recovered from the Quiricó Formation of the São Francisco Basin in Minas Gerais, eastern Brazil. This genus was named by Hussam Zaher, Diego Pol, Alberto B. Carvalho, Paulo M. Nascimento, Claudio Riccomini, Peter Larson, Rubén Juárez Valieri, Ricardo Pires Domingues, Nelson Jorge da Silva Jr. and Diógenes de Almeida Campos in 2011, and the type species is ''Tapuiasaurus macedoi''. Classification ''Tapuiasaurus'' was originally assigned to Nemegtosauridae by its original describers, but two subsequent cladistic analyses have recovered it as only distantly related to ''Nemegtosaurus'', with Wilson ''et al.'' (2016) recovering the genus outside the Lithostrotia, and Carballido ''et al.'' (2017) recovering it as closely relate ...
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Lirainosaurinae
Lirainosaurinae is a subfamily of lithostrotian titanosaur sauropods from the Late Cretaceous of France, Spain, and Romania. Systematics Lirainosaurinae was defined by Díez Díaz ''et al.'' (2018) as comprising "''Lirainosaurus'', ''Ampelosaurus'', their common ancestor, and all of its descendants", and includes the two definitional taxa as well as ''Atsinganosaurus'', '' Lohuecotitan'', and ''Paludititan''. The five genera are known from Europe and all within the timespan of the late Campanian (''Lirainosaurus'') to early Maastrichtian (''Ampelosaurus'' and ''Atsinganosaurus''). Lirainosaurinae was recovered by Diez ''et al.'' (2018) as phylogenetically intermediate between the clades traditionally considered Saltasauridae and a clade containing taxa normally found in Aeolosaurini and Lognkosauria Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. Description Lognkos ...
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Malawisaurus
''Malawisaurus'' (meaning "Malawi lizard") is an extinct genus of titanosaurian sauropod dinosaur. It is known from the Dinosaur Beds of northern Malawi, which probably date to the Aptian stage of the Early Cretaceous. The type species is ''M. dixeyi'' and the specific name honours Frederick Augustus Dixey. Discovery and naming ''Malawisaurus dixeyi'' was originally described in 1928 by Sidney H. Haughton as a species of ''Gigantosaurus'' (an invalid name for the diplodocid currently known as ''Tornieria''). Haughton considered it closely related to the species ''G. robustus'' (later the type species of ''Janenschia''). The holotype was discovered in the "Dinosaur Beds" of Malawi (then known as the Nyasaland Protectorate), which are usually considered to be of Barremian-Aptian age based on K–Ar dating, though they have also been suggested to be Late Cretaceous in age based on the vertebrate assemblage, and possibly also the Lupata Group. In 1993 it was placed in the newly na ...
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Mnyamawamtuka
''Mnyamawamtuka'' (pronounced "Mm-nya-ma-wah-mm-too-ka" ; meaning "beast of the Mtuka river drainage" in Kiswahili) is a genus of lithostrotian titanosaur sauropod dinosaur from the Cretaceous Galula Formation in Tanzania. The type and only species is ''M. moyowamkia''. Discovery and naming In 2004, a sauropod skeleton was found at the Mtuka River, twenty kilometres from Lake Rukwa. It was excavated between 2005 and 2008. In 2019, the type species ''Mnyamawamtuka moyowamkia'' was named and described by Eric Gorscak and Patrick M. O’Connor. The generic name is a contraction of the Kiswahili ''Mnyama wa Mtuka'', the "Beast of the Mtuka". The describers made explicit that they considered "beast" to be an apt name for a member of the Titanosauria. The specific name is a contraction of ''moyo wa mkia'', meaning the "heart of the tail" in Kiswahili, a reference to the heart-shaped cross-section of the rear facet of the middle tail vertebrae. The holotype, RRBP 05834, was found ...
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Rapetosaurus
''Rapetosaurus'' ( ) is a genus of titanosaurian sauropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period. Only one species, ''Rapetosaurus krausei'', has been identified. Like other sauropods, ''Rapetosaurus'' was a quadrupedal herbivore; it is calculated to have reached lengths of 15 metres (49 ft). Description ''Rapetosaurus'' was a fairly typical sauropod, with a short and slender tail, a very long neck and a huge, elephant-like body. Its head resembles the head of a diplodocid, with a long, narrow snout and nostrils on the top of its skull. It was a herbivore and its small, pencil-like teeth were good for ripping the leaves off trees but not for chewing. It was fairly modest in size, for a titanosaur. The juvenile specimen measured from head to tail, and "probably weighed about as much as an elephant". An adult would have been about twice as long ( in length) which is still less than half the length of its ...
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Sonidosaurus
''Sonidosaurus'' (meaning "Sonid lizard", after Sonid, the large geographical area that includes the type locality ) is a genus of sauropod dinosaur from the Late Cretaceous. It was a titanosaur which lived in what is now Inner Mongolia. The type species, ''Sonidosaurus saihangaobiensis'', was described by Xu, Zhang, Tan, Zhao, and Tan in 2006. It was a small titanosaur, about 9 meters (30 ft) long. It was first discovered in the Iren Dabasu Formation in 2001 in a quarry which would later yield the remains of ''Gigantoraptor'' starting in 2005; it was only slightly larger than the contemporaneous giant caenagnathid ''Gigantoraptor ''Gigantoraptor'' () is a genus of large oviraptorosaur dinosaur that lived in Asia during the Late Cretaceous period. It is known from the Iren Dabasu Formation of Inner Mongolia, where the first remains were found in 2005. ''Gigantoraptor'' wa ...''. References External links ''Sonidosaurus'' at Dinodata.de (in German) Late Creta ...
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