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Ostropomycetidae
The Ostropomycetidae are a subclass of mostly lichen-forming fungi in the class Lecanoromycetes. The subclass was circumscribed in 2004 by Catherine Reeb, François M. Lutzoni, and Claude Roux. It contains ten orders and 36 families. Arctomiaceae is the only family in the Ostropomycetidae that associates with cyanobacteria of the order Nostocales as its main photobiont partner. Taxa of uncertain classification The following taxa are of uncertain classification (incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...) in the Ostropomycetidae: Family incertae sedis: * Epigloeaceae Genera incertae sedis: *'' Amphorothecium'' – 1 sp. *'' Anzina'' – 1 sp. *'' Aspilidea'' – 1 sp. *'' Bachmanniomyces'' – 8 spp. *'' Dictyocatenulata'' – 1 sp. *'' Malvinia'' ...
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Baeomycetales
The Baeomycetales are an order of mostly lichen-forming fungi in the subclass Ostropomycetidae, in the class Lecanoromycetes. It contains 8 families, 33 genera and about 170 species. As a result of molecular phylogenetics research published in the late 2010s, several orders were folded into the Baeomycetales, resulting in a substantial increase in the number of taxa. Taxonomy The family Baeomycetaceae was originally proposed by Barthélemy Charles Joseph Dumortier in 1829 (under the spelling ''Baeomyceae''); he included two genera, ''Baeomyces'' and ''Calicium''. Baeomycetaceae was initially classified in the Lecanorales, and Baeomycetaceae and Cladoniaceae were thought to be closely related, sharing a phylogenetic origin in Lecideaceae. It was transferred to the order Helotiales based on the structure of its ascus, which is similar to those in genus '' Leotia''. However, the Helotiales consists of mostly non-lichenised fungi. The first DNA studies conducted with ''Baeomyces ...
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Sarrameanales
Sarrameanaceae is a family of lichen-forming fungi in the monotypic order Sarrameanales. It contains two genera, '' Loxospora'', and '' Sarrameana'', the type genus. The family was circumscribed by Josef Hafellner in 1984. The order Sarrameanales was proposed by Brendan Hodkinson and James Lendemer in 2011, as they had noted that previously published large-scale molecular phylogenetic studies had shown that the group of species contained in the family Sarrameanaceae were distinct and separate from the clade containing all of the other orders of the Ostropomycetidae. However, the name Sarrameanales was not validly published according to the rules of botanical nomenclature, because it was not accompanied by a suitable description. Despite this, the order continues to be used in lichenological literature. Sarrameanales is in the Ostropomycetidae; within this subclass, Sarrameanales and the order Schaereriales form a clade which has a sister relationship with a clade containing t ...
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Gyalectales
''Gyalectales'' is an order of lichen-forming fungi in the class Lecanoromycetes. It contains 5 families, 15 genera and about 550 species. Taxonomy The Gyalectales were introduced in a 1974 publication by Aino Henssen and Martin Jahns, but not formally published until 1986 by David Hawksworth and Ove Eriksson. Phylogeny An early (2002) phylogenetics study showed that the order Ostropales, as was then circumscribed, was paraphyletic, and proposed that the Ostropales sensu lato included the Gyalectales and Trapeliaceae. Although they had traditionally been considered to be only distantly related, molecular studies suggested a much closer phylogenetic relationship. As a result, of the molecular data, Kauff and Lutzoni subsumed the Gyalectales into the Ostropales, as the latter name was published earlier (1932 vs. 1986). In 2018, Kraichak and colleagues used a recently developed "temporal phylogenetic" approach to identify temporal bands for specific taxonomic ranks. Base ...
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Pertusariales
The Pertusariales are an order of fungi in the class Lecanoromycetes. It contains the following families: Agyriaceae The Agyriaceae are a family of lichenized fungi in the order Pertusariales. It contains two genera: ''Agyrium ''Agyrium'' is a genus of saprophytic fungi in the family Agyriaceae. It probably evolved from a lichen ancestor, as it is clos ..., Coccotremataceae, Icmadophilaceae, Megasporaceae, Microcaliciaceae, Ochrolechiaceae, Pertusariaceae, Varicellariaceae, and Variolariaceae. Many of these fungi form lichens. Gallery Image:Pertusaria_paratuberculifera_(EU).jpg, '' Pertusaria paratuberculifera'' (2 verrucae) Image:Pertusaria_paratuberculifera_(EU1).jpg, '' Pertusaria paratuberculifera'' (8 spores per ascus) References Lichen orders Lecanoromycetes orders Taxa named by David Leslie Hawksworth Taxa named by Maurice Choisy Taxa described in 1986 {{Pertusariales-stub ...
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Lecanoromycetes
Lecanoromycetes is the largest class of lichenized fungi. It belongs to the subphylum Pezizomycotina in the phylum Ascomycota. The asci (spore-bearing cells) of the Lecanoromycetes most often release spores by rostrate dehiscence. Genera of uncertain placement The are several genera in the Lecanoromycetes that have not been placed into any order or family. These are: *''Argopsis'' – 1 sp. *''Ascographa ''Ascographa'' is a genus of fungi in the Helotiales order. The relationship of this taxon to other taxa within the order is unknown (''incertae sedis''), and it has not yet been placed with certainty into any family. This is a monotypic In b ...'' - 1 sp. *'' Bartlettiella'' – 1 sp. *'' Bouvetiella'' – 1 sp. *'' Buelliastrum'' – 1 sp. *'' Haploloma'' – 1 sp. *'' Hosseusia'' – 3 spp. *'' Korfiomyces'' – 1 sp. *'' Maronella'' – 1 sp. *'' Notolecidea'' – 1 sp. *'' Petractis'' – 3 spp. *'' Piccolia'' – 10 spp. *'' Ravenelula'' – 2 spp. *'' R ...
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Anzina
''Anzina'' is a genus of lichenized fungi of uncertain familial and ordinal classification in the subclass Ostropomycetidae. It is a monotypic genus, containing the single species ''Anzina carneonivea''. This lichen was first described by Italian botanist Martino Anzi in 1868, as ''Gyalolechia carneonivea''. German lichenologist Christoph Scheidegger circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ... the genus ''Anzina'' in 1982. References Lecanoromycetes Lichen genera Monotypic Lecanoromycetes genera Taxa described in 1982 {{Lecanoromycetes-stub ...
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Agyriales
The Agyriales are an order of fungi in the class Lecanoromycetes. It contains the following families: Agyriaceae, Anamylopsoraceae and Schaereriaceae. Many of these fungi form lichens. In 2001, rRNA gene sequences of ''Anamylopsora pulcherrima'' ( Anamylopsoraceae), ''Placopsis gelida'', ''Trapelia involuta'' and ''Trapelia placodioides'' (Agyriaceae) showed that the Agyriineae family was outside the Lecanorales order (where it had been placed) and the study suggested a sister group relationship to Ostropales/Pertusariales orders. They were once thought to be a synonym of Pertusariales The Pertusariales are an order of fungi in the class Lecanoromycetes. It contains the following families: Agyriaceae The Agyriaceae are a family of lichenized fungi in the order Pertusariales. It contains two genera: ''Agyrium ''Agyri ..., until 2020 who re-established the order. References {{Taxonbar, from=Q2712199 Lichen orders Lecanoromycetes orders Taxa described in 193 ...
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Schaereriales
''Schaereria'' is a genus of lichen-forming fungi. It is the sole genus in the family Schaereriaceae, which itself is the only family in the Schaereriales, an order in the subclass Ostropomycetidae of the class Lecanoromycetes. Taxonomy The genus name of ''Schaereria'' is in honour of Ludwig Emanuel Schaerer (1785–1853), who was a Swiss pastor and lichenologist. Genus ''Schaereria'' was circumscribed by German lichenologist Gustav Wilhelm Körber in 1855, with '' Schaereria lugubris'' assigned as the type species. The genus was accepted a few years later by Theodor Magnus Fries. It subsequently fell into disuse as William Nylander placed it in synonymy with ''Lecidea''. Josef Poelt and Antonín Vězda resurrected the genus in 1977, and included '' S. cinereorufa''. ''Schaereria'' is one of several dozen genera whose species were previously included in the large genus ''Lecidea''. However, ''Lecidea'' has a different ascus structure than ''Schaereria''. The family Schae ...
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Cyanobacteria
Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, blue-green algae, although they are not usually scientifically classified as algae. They appear to have originated in a freshwater or terrestrial environment. Sericytochromatia, the proposed name of the paraphyletic and most basal group, is the ancestor of both the non-photosynthetic group Melainabacteria and the photosynthetic cyanobacteria, also called Oxyphotobacteria. Cyanobacteria use photosynthetic pigments, such as carotenoids, phycobilins, and various forms of chlorophyll, which absorb energy from light. Unlike heterotrophic prokaryotes, cyanobacteria have internal membranes. These are flattened sacs called thylakoids where photosynthesis is performed. Phototrophic eukaryotes such as green plants perform photosynthesis in plast ...
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Nostocales
The Nostocales are an order of cyanobacteria containing most of its species. It includes filamentous forms, both simple or branched, and both those occurring as single strands or multiple strands within a sheath. Some members show a decrease in width from the base, and some have heterocysts Heterocysts or heterocytes are specialized nitrogen-fixing cells formed during nitrogen starvation by some filamentous cyanobacteria, such as ''Nostoc punctiforme'', ''Cylindrospermum stagnale'', and ''Anabaena sphaerica''. They fix nitrogen from .... References Bacteria orders {{cyanobacteria-stub ...
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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Photobiont
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (