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Coleoptera
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species. Found in almost every habitat except the sea and the polar regions, they interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops. Beetles typically have a particularly hard e ...
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Coccinellidae
Coccinellidae () is a widespread family of small beetles ranging in size from . They are commonly known as ladybugs in North America and ladybirds in Great Britain. Some entomologists prefer the names ladybird beetles or lady beetles as they are not true bugs. Many of the species have conspicuous aposematic colours and patterns, such as red with black spots, that warn potential predators that they are distasteful. The majority of the more than 6,000 described species are generally considered beneficial insects, because many prey on herbivorous hemipterans such as aphids or scale insects, which are agricultural pests. Many coccinellids lay their eggs directly in aphid and scale insect colonies, ensuring their larvae have an immediate food source. However, some species such as the herbivorous Mexican bean beetle are agricultural pests. Etymology The name ''coccinellids'', created by Pierre André Latreille, is derived from the Latin word ''coccineus'' meaning "scarlet". ...
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List Of Subgroups Of The Order Coleoptera
This article classifies the subgroups of the order Coleoptera (beetles) down to the level of families, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020, with common names from bugguide.net. *Order Coleoptera **Suborder † Protocoleoptera ***Superfamily †Tshekardocoleoidea Rohdendorf, 1944 ****Family †Tshekardocoleidae Rohdendorf, 1944 ****Family † Labradorocoleidae Ponomarenko, 1969 ****Family † Oborocoleidae Kukalová, 1969 ***Superfamily † Permocupedoidea Martynov, 1933 ****Family †Permocupedidae Martynov, 1933 ****Family †Taldycupedidae Rohdendorf, 1961 ***Superfamily † Permosynoidea Tillyard, 1924 ****Family †Ademosynidae Ponomarenko, 1968 ****Family †Permosynidae Tillyard, 1924 **Suborder Archostemata ***Superfamily Cupedoidea Laporte, 1836 ****Family Crowsoniellidae Iablokoff-Khnzorian, 1983 ****Family Cupedidae Laporte, 1836 ****Family Micromalthidae Barber, 1913 ...
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Myxophaga
Myxophaga is the second-smallest suborder of the Coleoptera after Archostemata, consisting of roughly 65 species of small to minute beetles in four families. The members of this suborder are aquatic and semiaquatic, and feed on algae. Description Myxophaga have several diagnostic features: the antennae are more or less distinctly clubbed with usually fewer than nine segments, mesocoxal cavities are open laterally and bordered by a mesepimeron and metanepisternum, the hind wings are rolled apically in the resting positions. Internally, they are characterised by the presence of six malpighian tubules and the testes are tube-like and coiled. Beetles of this suborder are adapted to feed on algae. Their mouthparts are characteristic in lacking galeae and having a mobile tooth on their left mandible. Taxonomy There are four extant families in the suborder Myxophaga divided between two superfamilies, containing about 65 described species, and at least one extinct family. Superfami ...
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Colorado Potato Beetle
The Colorado potato beetle (''Leptinotarsa decemlineata''), also known as the Colorado beetle, the ten-striped spearman, the ten-lined potato beetle, or the potato bug, is a major pest of potato crops. It is about long, with a bright yellow/orange body and five bold brown stripes along the length of each of its elytra. Native to the Rocky Mountains, it spread rapidly in potato crops across America and then Europe from 1859 onwards. Taxonomy The Colorado potato beetle was first observed in 1811 by Thomas Nuttall and was formally described in 1824 by American entomologist Thomas Say.see pp. 453–454: "Doryphora, Illig.: D. 10-lineata"./ref> The beetles were collected in the Rocky Mountains, where they were feeding on the buffalo bur, '' Solanum rostratum''. The genus '' Leptinotarsa'' is assigned to the chrysolmelid beetle tribe Chrysomelini (in subfamily Chrysomelinae). Description Adult beetles typically are in length and in width. They weigh 50-170 mg. The beetles ...
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Insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch f ...
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Protocoleoptera
The Protocoleoptera are a paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern beetle groups during the following Triassic. Protocoleopterans typically possess prognathous (horizontal) heads, distinctive elytra with regular window punctures, culticles with tubercles or scales, as well as a primitive pattern of ventral sclerites, similar to the modern archostematan families Ommatidae and Cupedidae. They are thought to have been xylophagous and wood boring. Kirejtshuk ''et al.'' (2014) argue that the name "Protocoleoptera" should not be used for the group, as Protocoleoptera was originally proposed for the family Protocoleidae , now considered a member of the extinct order Protelytroptera (a stem-group of the modern Dermaptera, the earwigs). Because of this, Cai ''et al.'' (2022) proposed a replacement name, Alphacoleoptera ...
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Adephaga
The Adephaga (from Greek ἀδηφάγος, ''adephagos'', "gluttonous") are a suborder of beetles, and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collectively known as adephagans. The largest family is Carabidae (ground beetles) which comprises most of the suborder with over 40,000 species. Adephaga also includes a variety of aquatic beetles, such as predaceous diving beetles and whirligig beetles. Anatomy Adephagans have simple antennae with no pectination or clubs. The galeae of the maxillae usually consist of two segments. Adult adephagans have visible notopleural sutures. The first visible abdominal sternum is completely separated by the hind coxae, which is one of the most easily recognizable traits of adephagans. Five segments are on each foot. Wings The transverse fold of the hind wing is near the wing tip. The median nervure ends at this fold, where it is joined by a ...
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Archostemata
The Archostemata are the smallest suborder of beetles, consisting 45 living species in five families. They are an ancient lineage with a number of primitive characteristics. Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are paedogenic, '' Micromalthus debilis''. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic. The term "Archostemata" is used more broadly by some authors to include both modern archostematans as well as stem-group beetles like " protocoleopterans", which some modern archostematans closely resemble to due to their plesiomorphic morphology. Genetic research suggests that modern archostematans are a monophyletic group. Some genetic studies have recovered archostematans as the sister group of Myxophaga. Taxonomy There are five extant families. * Family Crowsoniellidae Iablokoff-Khnzorian, 1983 * Family Cupedidae Laporte, 1838 ...
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Rove Beetles
The rove beetles are a family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With roughly 63,000 species in thousands of genera, the group is currently recognized as the largest extant family of organisms. It is an ancient group, with fossilized rove beetles known from the Triassic, 200 million years ago, and possibly even earlier if the genus ''Leehermania'' proves to be a member of this family. They are an ecologically and morphologically diverse group of beetles, and commonly encountered in terrestrial ecosystems. One well-known species is the devil's coach-horse beetle. For some other species, see list of British rove beetles. Anatomy As might be expected for such a large family, considerable variation exists among the species. Sizes range from <1 to , with most in the 2–8 mm range, and the form is generally elongated, with some rove beetles being ovoid ...
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Dynastinae
Dynastinae or rhinoceros beetles are a subfamily of the scarab beetle family (Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include Hercules beetles, unicorn beetles or horn beetles. Over 1500 species and 225 genera of rhinoceros beetles are known. Many rhinoceros beetles are well known for their unique shapes and large sizes. Some famous species are, for example, the Atlas beetle (''Chalcosoma atlas''), common rhinoceros beetle (''Xylotrupes ulysses''), elephant beetle (''Megasoma elephas''), European rhinoceros beetle (''Oryctes nasicornis''), Hercules beetle (''Dynastes hercules''), Japanese rhinoceros beetle or ''kabutomushi'' (''Allomyrina dichotoma''), ox beetle (''Strategus aloeus'') and the Eastern Hercules beetle (''Dynastes tityus''). Description and ecology The Dynastinae are among the largest of beetles, reaching more than in length, but are completely harmless to humans because they cannot bite or sting. So ...
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Polyphaga
Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the beetle species so far discovered. Key characteristics of Polyphaga are that the hind coxa (base of the leg) does not divide the first and second abdominal/ventral plates which are known as sternites. Also, the notopleural suture (found under the pronotal shield) is not present. Etymology The name of ''polyphaga'' is derived from two Greek words: , meaning 'many', and , meaning 'to eat', so the suborder is called the “eaters of many things”. Classification The five main infraorders are: * Bostrichiformia — including furniture beetles and skin beetles * Cucujiformia — includes lady beetles, longhorn beetles, weevils, checkered beetles and leaf beetles * Elateriformia — includes click beetles and fireflies * Scarabaeiform ...
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Permian
The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the Perm Governorate, region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the Sauropsida, sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditi ...
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