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This article classifies the subgroups of the order Coleoptera (
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s) down to the level of
families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...
, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020, with common names from bugguide.net. *Order
Coleoptera Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
**Suborder †
Protocoleoptera The Protocoleoptera are a Paraphyly, paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern bee ...
***Superfamily † Tshekardocoleoidea Rohdendorf, 1944 ****Family † Tshekardocoleidae Rohdendorf, 1944 ****Family † Labradorocoleidae Ponomarenko, 1969 ****Family † Oborocoleidae Kukalová, 1969 ***Superfamily † Permocupedoidea Martynov, 1933 ****Family † Permocupedidae Martynov, 1933 ****Family † Taldycupedidae Rohdendorf, 1961 ***Superfamily † Permosynoidea Tillyard, 1924 ****Family † Ademosynidae Ponomarenko, 1968 ****Family † Permosynidae Tillyard, 1924 **Suborder
Archostemata The Archostemata are the smallest suborder of beetles, consisting of 50 living species in five families and over 200 described fossil species. They are an ancient lineage with a number of primitive characteristics. Antenna (biology), Antennae may ...
***Superfamily Cupedoidea Laporte, 1836 ****Family Crowsoniellidae Iablokoff-Khnzorian, 1983 ****Family Cupedidae Laporte, 1836 ****Family Micromalthidae Barber, 1913 ****Family
Ommatidae The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. Howev ...
Sharp and Muir, 1912 ****Family Jurodidae Ponomarenko, 1985 ****Family † Triadocupedidae Ponomarenko, 1966 ****Family † Magnocoleidae Hong, 1998 ****Family † Obrieniidae Zherikhin and Gratshev, 1994 ***Superfamily † Asiocoleoidea Rohdendorf, 1961 ****Family † Asiocoleidae Rohdendorf, 1961 ****Family † Tricoleidae Ponomarenko, 1969 ***Superfamily † Rhombocoleoidea Rohdendorf, 1961 ****Family † Rhombocoleidae Rohdendorf, 1961 ***Superfamily † Schizocoleoidea Rohdendorf, 1961 (formerly Schizophoroidea Ponomarenko, 1968) ****Family † Phoroschizidae Bouchard and Bousquet, 2020 (formerly Schizophoridae Ponomarenko, 1968) ****Family † Catiniidae Ponomarenko, 1968 ****Family † Schizocoleidae Rohdendorf, 1961 **Suborder
Myxophaga Myxophaga is the second-smallest suborder of the Coleoptera after Archostemata, consisting of roughly 65 species of small to minute beetles in four families. The members of this suborder are aquatic and semiaquatic, and feed on algae. Descript ...
***Superfamily Lepiceroidea Hinton, 1936 (1882) ****Family Lepiceridae Hinton, 1936 (1882) ***Superfamily Sphaeriusoidea Erichson, 1845 ****Family Torridincolidae Steffan, 1964 ****Family
Hydroscaphidae The Hydroscaphidae are a small family of water beetles known commonly as skiff beetles. As of 2010, there are 23 species in the family. Several are recently described. These beetles are small, most under in length. They are tan to brown in col ...
LeConte, 1874 ****Family Sphaeriusidae Erichson, 1845 **Suborder
Adephaga The Adephaga (from Greek ἀδηφάγος, ''adephagos'', "gluttonous") are a suborder of beetles, and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collecti ...
****Family † Tritarsusidae Hong, 2002 (formerly Tritarsidae Hong, 2002) ****Family
Gyrinidae The whirligig beetles are water beetles, comprising the family Gyrinidae, that usually swim on the surface of the water if undisturbed, though they swim underwater when threatened. They get their common name from their habit of swimming rapidly ...
Latreille, 1810 (whirligig beetles) ****Family Trachypachidae Thomson, 1857 (false ground beetles) ****Family
Rhysodidae Rhysodinae is a subfamily (sometimes called wrinkled bark beetles) in the family Carabidae. There are 19 genera and at least 380 described species in Rhysodinae. The group of genera making up Rhysodinae had been treated as the family Rhysodidae i ...
Laporte, 1840 ****Family
Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal ...
Latreille, 1802 (ground beetles) ****Family
Haliplidae The Haliplidae are a family (biology), family of water beetles that swim using an alternating motion of the legs. They are therefore clumsy in water (compared e.g. with the Dytiscidae or Hydrophilidae), and prefer to get around by crawling. The f ...
Aubé, 1836 (crawling water beetles) ****Family † Triaplidae Ponomarenko, 1977 ****Family † Colymbotethidae Ponomarenko, 1994 ****Family † Parahygrobiidae Ponomarenko, 1977 ****Family †
Coptoclavidae Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Adephaga, Dytiscoidea, which contains living ...
Ponomarenko, 1961 ****Family † Liadytidae Ponomarenko, 1977 ****Family Meruidae Spangler and Steiner, 2005 ****Family Noteridae Thomson, 1860 (burrowing water beetles) ****Family Amphizoidae LeConte, 1853 ****Family Aspidytidae Ribera, Beutel, Balke and Vogler, 2002 ****Family Hygrobiidae Régimbart, 1879 (1837) ****Family
Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive", are the predaceous diving beetles, a family of water beetles. They occur in virtually any freshwater habitat around the world, but a few species l ...
Leach, 1815 (predaceous diving beetles) **Suborder
Polyphaga Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the b ...
***Infraorder
Staphyliniformia Staphyliniformia is a large infraorder of beetles. It contains over 70,000 described species from all regions of the world. Most species occur in moist habitats - various kinds of rotting plant debris, fungi, dung, carrion, many live in fresh wat ...
****Superfamily
Histeroidea Histeroidea is a Taxonomic rank, superfamily of beetles in the infraorder Staphyliniformia. Characteristics Characteristic to Histeroidea are an accessory posterior ridge (locking device) behind the hind margin and presence of Comstock-Needham s ...
Gyllenhal, 1808 *****Family
Histeridae Histeridae is a family (biology), family of beetles commonly known as clown beetles or hister beetles. There are more than 410 genera and 4,800 described species in Histeridae worldwide, with more than 500 species in North America. They can be id ...
Gyllenhal, 1808 *****Family
Sphaeritidae ''Sphaerites'' is a genus of beetles, the only genus in the family Sphaeritidae, sometimes called the false clown beetles. There are five known species, which are widespread in temperate areas of the Northern Hemisphere, found in forested or upla ...
Shuckard, 1839 *****Family
Synteliidae ''Syntelia'' is a genus of beetles. It is the only genus in the family Synteliidae. There are seven known species, which are native to high-elevation regions in southern North America from central Mexico to Guatemala, and in eastern Asia, from I ...
Lewis, 1882 ****Superfamily
Hydrophiloidea Hydrophiloidea, known as water scavenger beetles, is a Taxonomic rank, superfamily of beetles. Until recently it included only a single family, the Hydrophilidae (water scavenger beetles), but several of the subfamilies have been removed and rais ...
Latreille, 1802 *****Family Epimetopidae Zaitzev, 1908 *****Family
Georissidae ''Georissus'', also called minute mud-loving beetles, is the only genus in the beetle family Georissidae (or Georyssidae). They are tiny insects living in wet soil, often near water. They are found on every continent except Antarctica. Character ...
Laporte, 1840 *****Family Helophoridae Leach, 1815 *****Family
Hydrochidae ''Hydrochus'' is the only living genus of beetle in the family Hydrochidae, which belongs to the superfamily Hydrophiloidea, and was formerly treated as a subfamily of Hydrophilidae. ''Hydrochus'' includes about 180 species, which are found worldw ...
Thomson, 1859 *****Family
Hydrophilidae Hydrophilidae, also known colloquially as water scavenger beetles, is a family of beetles. Aquatic hydrophilids are notable for their long maxillary palps, which are longer than their antennae. Several of the former subfamilies of Hydrophilidae ...
Latreille, 1802 (water scavenger beetles) *****Family
Spercheidae ''Spercheus'' is a genus of aquatic beetles which are placed in a family of their own, Spercheidae within the Hydrophiloidea. About 20 species are known from around the world except the Nearctic with the majority being from the Oriental and Afro ...
Erichson, 1837 ****Superfamily
Staphylinoidea Staphylinoidea is a superfamily of beetles. It is a very large and diverse group with worldwide distribution. Description Adult staphylinoids are generally small beetles no more than a few millimetres long, though Staphylinidae can reach 50 ...
Latreille, 1802 *****Family
Hydraenidae Hydraenidae is a family of very small aquatic animal, aquatic beetles, sometimes called "Minute moss beetles", with a worldwide distribution. They are around 0.8 to 3.3 mm in length. The adults store air on the underside of the body as wel ...
Mulsant, 1844 *****Family
Ptiliidae Ptiliidae is a family of very tiny beetles (including the smallest of all beetles) with a cosmopolitan distribution. They are colloquially called featherwing beetles, because the hindwings are narrow and feathery. There are approximately 600 des ...
Erichson, 1845 *****Family Agyrtidae Thomson, 1859 *****Family
Leiodidae Leiodidae is a family of beetles with around 3800 described species found worldwide. Members of this family are commonly called round fungus beetles due to the globular shape of many species, although some are more elongated in shape. They are g ...
Fleming, 1821 *****Family
Silphidae Silphidae is a family of beetles that are known commonly as large carrion beetles, carrion beetles or burying beetles. There are two family (biology), subfamilies: Silphinae and Nicrophorinae. Members of Nicrophorinae are sometimes known as bury ...
Latreille, 1806 *****Family
Staphylinidae The rove beetles are a family (biology), family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With over 66,000 species in thousand ...
Latreille, 1802 (rove beetles) ***Infraorder
Scarabaeiformia Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Some of its constituent families are undergo ...
****Superfamily
Scarabaeoidea Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Some of its constituent families are undergo ...
Latreille, 1802 *****Family Pleocomidae LeConte, 1861 (rain beetles) *****Family
Bolboceratidae Bolboceratidae is a family of beetle. It was historically treated as a subfamily of the earth-boring dung beetles, or "dor beetles" (family Geotrupidae), but has been considered a separate family by many authors since 1995. Some recent classific ...
Mulsant, 1842 *****Family
Geotrupidae Geotrupidae (from Greek γῆ ''(gē)'', earth, and τρῡπητής ''(trȳpētēs)'', borer) is a family of beetles in the order Coleoptera. They are commonly called earth-boring dung beetles or dor beetles. Most excavate burrows in which t ...
Latreille, 1802 (earth-boring scarab beetles) *****Family
Belohinidae ''Belohina inexpectata'' is a species of polyphagan beetles and the sole member of family Belohinidae. It is endemic to southern Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island ...
Paulian, 1959 *****Family
Passalidae Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles", "betsy bugs", or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their siz ...
Leach, 1815 (bess beetles) *****Family
Trogidae Trogidae, sometimes called hide beetles, is a family of beetles with a distinctive warty or bumpy appearance. Found worldwide, the family includes about 300 species contained in four or five genera. Trogids range in length from 2 to 20 mm. Thei ...
MacLeay, 1819 (hide beetles) *****Family
Glaresidae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy areas wo ...
Prudhomme de Borre, 1886 *****Family
Diphyllostomatidae The false stag beetles (''Diphyllostoma'') are a group of three species of rare beetles known only from California. Almost nothing is known of their life history beyond that the adults are diurnal and females are flightless; larvae have not been ...
Holloway, 1972 *****Family
Lucanidae Stag beetles comprise the family Lucanidae. It has about 1,200 species of beetles in four subfamilies.Smith, A.B.T. (2006). A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a c ...
Latreille, 1804 (stag beetles) *****Family
Ochodaeidae Ochodaeidae, also known as the sand-loving scarab beetles, is a small family of scarabaeiform beetles occurring in many parts of the world. These beetles are small, ranging from . Their bodies are elongate and convex, with black and brown colo ...
Streubel, 1846 (sand-loving scarab beetles) *****Family
Hybosoridae Hybosoridae, sometimes known as the scavenger scarab beetles, is a family of Scarabaeiformia, scarabaeiform beetles. The >600 species in 78 extant genera occur widely in the tropics, but little is known of their biology. Hybosorids are small, 5 ...
Erichson, 1847 *****Family
Glaphyridae Glaphyridae is a family of beetles, commonly known as bumble bee scarab beetles. There are eight extant genera with about 80 species distributed worldwide and two extinct genera described from the Aptian The Aptian is an age (geology), age in ...
MacLeay, 1819 (bumble bee scarab beetles) *****Family
Scarabaeidae The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly tre ...
Latreille, 1802 (scarab beetles) *****Family † Coprinisphaeridae Genise, 2004 (
ichnotaxon An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxon'' comes from the Ancient Greek (''íchnos'') meaning "track" and English , itself derived from ...
) *****Family † Pallichnidae Genise, 2004 (ichnotaxon) ***Infraorder
Elateriformia Elateriformia is an infraorder of polyphagan beetles. The two largest families in this group are buprestids, of which there are around 15,000 described species, and click beetles, of which there are around 10,000 described species. The infraor ...
****Superfamily
Scirtoidea Scirtoidea is a superfamily of beetles. It is traditionally considered to consist of four families: Clambidae, Decliniidae, Eucinetidae and Scirtidae. However, genetic studies have suggested that Clambidae and Eucinetidae belong to a separate su ...
Fleming, 1821 *****Family
Decliniidae Decliniidae is a family of beetles belonging to Scirtoidea. It contains the single genus ''Declinia'' with two species, ''D. relicta'' and ''D. versicolor'', found in the Russian Far East The Russian Far East ( rus, Дальний Вос ...
Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994 *****Family
Eucinetidae Eucinetidae is a family (biology), family of beetles, notable for their large coxal plates that cover much of the first ventrite of the abdomen, sometimes called plate-thigh beetles. The family is small for beetles, with about 50 species in 11 ...
Lacordaire, 1857 *****Family
Clambidae Clambidae is a family of beetles. They are known commonly as the minute beetlesMajka, C. G., & Langor, D. (2009)Clambidae (Coleoptera) of Atlantic Canada.''Journal of the Acadian Entomological Society'' 5(7), 32-40. or the fringe-winged beetles.< ...
Fischer von Waldheim, 1821 *****Family
Scirtidae Scirtidae is a family of beetles (Coleoptera). These beetles are commonly referred to as marsh beetles, as the larvae are typically associated with stagnant water, but can be found in flowing water. Adults prefer decomposing plant material near ...
Fleming, 1821 *****Family † Elodophthalmidae Kirejtshuk and Azar, 2008 *****Family † Mesocinetidae Kirejtshuk and Ponomarenko, 2010 ****Superfamily
Dascilloidea Dascilloidea is a superfamily of polyphagan beetles, comprising two families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis fo ...
Guérin-Méneville, 1843 (1834) *****Family Dascillidae Guérin-Méneville, 1843 (1834) *****Family
Rhipiceridae Rhipiceridae is a family of beetles found worldwide. The larva of rhipicerids are parasitoids of cicada nymphs. Rhipiceridae and Dascillidae form the super family Dascilloidea, within the Elateriformia. Taxonomic History The taxonomic history ...
Latreille, 1834 ****Superfamily
Buprestoidea Buprestoidea is a superfamily of beetles. It contains two families: * Buprestidae Buprestidae is a family (biology), family of beetles known as jewel beetles or metallic wood-boring beetles because of their glossy Iridescence, iridescent co ...
Leach, 1815 *****Family
Schizopodidae Schizopodidae is a family of beetles, in the large suborder Polyphaga. It was a subfamily until 1991, when it was elevated to family status. The family of Schizopodidae is a part of the superfamily, Buprestoidea, which is a member of the suborde ...
LeConte, 1859 *****Family
Buprestidae Buprestidae is a family (biology), family of beetles known as jewel beetles or metallic wood-boring beetles because of their glossy Iridescence, iridescent colors. Larvae of this family are known as flatheaded borers. The family is among the lar ...
Leach, 1815 ****Superfamily
Byrrhoidea Byrrhoidea is a superfamily of beetles belonging to Elateriformia that includes several families which are either aquatic or associated with a semi-aquatic habitat. Other than the superfamily Hydrophiloidea, most of the remaining Polyphagan beetle ...
Latreille, 1804 *****Family Byrrhidae Latreille, 1804 *****Family Protelmidae Jeannel, 1950 *****Family
Elmidae Elmidae, commonly known as riffle beetles, is a family of beetles in the superfamily Byrrhoidea described by John Curtis in 1830. Both adults and larvae are usually aquatic, living under rocks in fast-flowing shallow areas of streams, such as ...
Curtis, 1830 *****Family
Dryopidae Dryopidae is a family (biology), family of beetles, commonly named long-toed water beetles, in the superfamily Byrrhoidea. It was described by Gustaf Johan Billberg in 1820. Description Long-toed water beetles are named for their extended claws. ...
Billberg, 1820 (1817) *****Family Lutrochidae Kasap and Crowson, 1975 *****Family
Limnichidae Limnichidae, commonly called minute marsh-loving beetles, is a family of beetles belonging to Byrrhoidea. There are at least 30 genera and 350 described species in Limnichidae. They are found worldwide, with the greatest diversity in tropical regi ...
Erichson, 1846 *****Family
Heteroceridae Heteroceridae, the variegated mud-loving beetles, are a widespread and relatively common family of beetles found on every continent except for Antarctica. Around two hundred and fifty species of heterocerids are known to occur worldwide. They a ...
MacLeay, 1825 *****Family
Psephenidae Water-penny beetles are a family (biology), family (the Psephenidae) of 273 species (in 35 genera) of aquatic beetles found on all continents except Antarctica, in both tropical and temperate areas. The young, which live in water, resemble tiny ...
Lacordaire, 1854 *****Family Cneoglossidae Champion, 1897 *****Family Ptilodactylidae Laporte, 1836 *****Family
Podabrocephalidae ''Podabrocephalus'' is an enigmatic monotypic genus in the family Ptilodactylidae. For decades it had historically been placed in its own family, Podabrocephalidae, or sometimes included in the highly diverse family Cerambycidae The longhorn b ...
Pic, 1930 *****Family
Chelonariidae Chelonariidae or turtle beetles is a family of beetles in the superfamily Byrrhoidea. It was described by Blanchard in 1845. There are 3 genera with around 300 described species. Little is known of their ecology, though it seems they are associa ...
Blanchard, 1845 *****Family Eulichadidae Crowson, 1973 *****Family Callirhipidae Emden, 1924 ****Superfamily
Elateroidea The Elateroidea are a large superfamily of beetles. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species. Description Elateroidea is a morphologically diverse group ...
Leach, 1815 *****Family
Artematopodidae Artematopodidae is a family of soft-bodied plant beetles in the superfamily Elateroidea. They are mostly found in understory forest foliage. The life history of the group is obscure, larvae of the genera '' Eurypogon'' and '' Macropogon'' likely ...
Lacordaire, 1857 *****Family
Brachypsectridae The Brachypsectridae are a family of beetles commonly known as the Texas beetles. There are only two extant genera, '' Brachypsectra'' and '' Asiopsectra. Brachypsectra'' has a cosmopolitan distribution, mostly in arid regions, while ''Asiopsectr ...
Horn, 1881 *****Family
Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are Cosmopolitan distribution, cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the Red coat (Bri ...
Imhoff, 1856 (1815) (soldier beetles) *****Family
Cerophytidae The Cerophytidae are a family of beetles belonging to Elateroidea. Larvae are associated with rotting wood, on which they are presumed to feed.Costa, Cleide, Vanin, Sergio A., Lawrence, John F. and Ide, Sergio. "4.4. Cerophytidae Latreille, 1834 ...
Latreille, 1834 *****Family
Elateridae Elateridae or click beetles (or "typical click beetles" to distinguish them from the related families Cerophytidae and Eucnemidae, which are also capable of clicking) are a family of beetles. Other names include elaters, snapping beetles, sp ...
Leach, 1815 (click beetles; includes former Drilidae, Omalisidae, and Plastoceridae) *****Family
Eucnemidae Eucnemidae, or false click beetles, are a family of elateroid beetles based on the type genus '' Eucnemis''; they include about 1700 species, distributed worldwide. Description Closely related to the family Elateridae, specimens of Eucnemida ...
Eschscholtz, 1829 *****Family
Lampyridae The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...
Rafinesque, 1815 (fireflies) *****Family
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
Laporte, 1836 *****Family
Omethidae Omethidae is a family of Elateroidea sometimes known as the false soldier beetles. They are native to South, Southeast and Eastern Asia and the Americas. Their biology is obscure and their larvae are unknown. They appear to inhabit vegetation in ...
LeConte, 1861 (includes former Telegeusidae) *****Family
Phengodidae The beetle family Phengodidae is known also as glowworm beetles, whose larvae are known as glowworms. The females and larvae have bioluminescent organs. They occur throughout the New World from extreme southern Canada to Chile, numbering over ...
LeConte, 1861 *****Family
Rhagophthalmidae The Rhagophthalmidae are a family of beetles within the superfamily Elateroidea. Members of this beetle family have bioluminescent organs on the larvae, and sometimes adults, and are closely related to the Phengodidae ( American glowworm beetles ...
Olivier, 1907 *****Family Sinopyrophoridae Bi, 2018 *****Family Throscidae Laporte, 1840 nomen protectum *****Family † Berendtimiridae Winkler, 1987 *****Family † Praelateriidae Dolin, 1973 ****Superfamily
Rhinorhipoidea ''Rhinorhipus'' is a genus of beetles that contains a single species, ''Rhinorhipus tamborinensis'' from southern Queensland, Australia. It is the sole member of the family Rhinorhipidae and superfamily Rhinorhipoidea. It is an isolated lineage n ...
Lawrence, 1988 *****Family
Rhinorhipidae ''Rhinorhipus'' is a genus of beetles that contains a single species, ''Rhinorhipus tamborinensis'' from southern Queensland, Australia. It is the sole member of the family Rhinorhipidae and superfamily Rhinorhipoidea. It is an isolated lineage n ...
Lawrence, 1988 ***Infraorder
Bostrichiformia Bostrichiformia is an infraorder of polyphagan beetles. It contains two superfamilies, Derodontoidea and Bostrichoidea, which includes the Dermestidae, Ptinidae, Bostrichidae The Bostrichidae are a family of beetles with more than 700 descri ...
****Superfamily
Derodontoidea Derodontidae is a family of beetles, in its own superfamily, Derodontoidea, sometimes known as the tooth-necked fungus beetles. Beetles of this family are small, between 2 and 6 mm in length, typically with spiny margins on their pronotum ( ...
LeConte, 1861 *****Family
Derodontidae Derodontidae is a family of beetles, in its own superfamily, Derodontoidea, sometimes known as the tooth-necked fungus beetles. Beetles of this family are small, between 2 and 6 mm in length, typically with spiny margins on their pronotum ...
LeConte, 1861 *****Family Nosodendridae Erichson, 1846 *****Family
Jacobsoniidae Jacobsoniidae are a family of tiny beetles belonging to Staphylinoidea. The larvae and adults live under bark, in plant litter, fungi, bat guano and rotten wood. There are around 28 described species in three genera: Description Members of t ...
Heller, 1926 ****Superfamily
Bostrichoidea Bostrichoidea is a Taxonomic rank, superfamily of beetles. It is the type superfamily of the infraorder Bostrichiformia. Description The Bostrichoidea are united by the following features: modified cryptonephridism, the structure of the aedea ...
Latreille, 1802 *****Family
Dermestidae Dermestidae are a family of Coleoptera that are commonly referred to as skin beetles or carpet beetles. Other common names include larder beetles, hide or leather beetles, and khapra beetles. There are over 1,800 species described. Dermestids ha ...
Latreille, 1804 (carpet beetles) *****Family
Endecatomidae ''Endecatomus'' is a genus of beetles, the sole member of the family Endecatomidae. There are at least four described species in ''Endecatomus''. Endecatomidae was formerly treated as a subfamily of Ciidae or Bostrichidae. They are found in the H ...
LeConte, 1861 *****Family
Bostrichidae The Bostrichidae are a family of beetles with more than 700 described species. They are commonly called auger beetles, false powderpost beetles, or horned powderpost beetles. Anatomy The head of most auger beetles cannot be seen from above, as ...
Latreille, 1802 *****Family
Ptinidae Ptinidae is a family of beetles in the superfamily Bostrichoidea. There are at least 220 genera and 2,200 described species in Ptinidae worldwide. The family includes spider beetles and deathwatch beetle, as well as the Cigarette beetle, cigare ...
Latreille, 1802 ***Infraorder
Cucujiformia Cucujiformia is an infraorder of polyphagan beetles, representing most plant-eating beetles. The infraorder contains the seven superfamilies: * Chrysomeloidea (~7 families including longhorn beetles and leaf beetles) * Cleroidea (checkered be ...
****Superfamily Lymexyloidea Fleming, 1821 *****Family Lymexylidae Fleming, 1821 ****Superfamily
Cleroidea Cleroidea is a small Taxonomic rank, superfamily of beetles containing over 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elytra, and typically hairy or scaly. Description Cleroidea is ...
Latreille, 1802 *****Family
Phloiophilidae ''Phloiophilus edwardsii'' is the sole known species of the beetle family Phloiophilidae in the superfamily Cleroidea. It is native to Europe. The larvae are mycophagous, and have been observed feeding on Basidiomycota, basidiomycetes of the genu ...
Kiesenwetter, 1863 *****Family
Trogossitidae Trogossitidae, also known as bark-gnawing beetles, are a small family in the superfamily Cleroidea. Many taxa formerly within this family have been removed (as of 2019) to other families, such as Lophocateridae, Peltidae, Protopeltidae, Rentoniid ...
Latreille, 1802 *****Family Chaetosomatidae Crowson, 1952 *****Family Metaxinidae Kolibáč, 2004 *****Family
Thanerocleridae Thanerocleridae is a family of beetles belonging to the superfamily Cleroidea. It was formerly considered a subfamily of Cleridae, but was recently elevated to the rank of family. The family has 36 living species in 10 genera, which are found glo ...
Chapin, 1924 *****Family
Cleridae Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences. Cleridae have many Ecological nich ...
Latreille, 1802 (checkered beetles) *****Family
Acanthocnemidae ''Acanthocnemus nigricans'' is a species of cleroid beetle, the only species in the genus ''Acanthocnemus'' and the family Acanthocnemidae. Originally native to Australia, the species has spread in modern times to numerous parts of the world, in ...
Crowson, 1964 *****Family Phycosecidae Crowson, 1952 *****Family
Prionoceridae Prionoceridae is a small family of beetles, in the suborder Polyphaga. They form a group within the cleroid beetles and were formerly treated as a subfamily (Prionocerinae) within the family Melyridae. Very little is known of their life history ...
Lacordaire, 1857 *****Family
Mauroniscidae Mauroniscidae is a family of Cleroidea, cleroid beetles, formerly included in the family Melyridae. There are presently five or six genera and roughly 30 described species in Mauroniscidae, all of which are native to the Americas. Almost nothing ...
Majer, 1995 *****Family
Melyridae Melyridae (common name: soft-winged flower beetles) are a family of beetles of the superfamily Cleroidea. Description Most are elongate-oval, soft-bodied beetles 10 mm long or less. Many are brightly patterned in black and brown, yellow, ...
Leach, 1815 (soft-winged flower beetles) ****Superfamily
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark ...
Latreille, 1802 *****Family † Parandrexidae Kirejtshuk, 1994 *****Family † Sinisilvanidae Hong, 2002 *****Family Boganiidae Sen Gupta and Crowson, 1966 *****Family Byturidae Gistel, 1848 *****Family Helotidae Chapuis, 1876 *****Family
Protocucujidae Protocucujidae is a family of beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most oth ...
Crowson, 1954 *****Family
Sphindidae Sphindidae is a family of beetles, in the suborder Polyphaga. They are called slime mold beetles due to their exclusive feeding on slime molds during adult and larval stages, other aspects of their life history are obscure. Palaeontological disc ...
Jacquelin du Val, 1860 *****Family Biphyllidae LeConte, 1861 *****Family
Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumscription, it contains 6 tribes ( Tritomini, Dacnini, Megalodacnini, Er ...
Latreille, 1802 *****Family
Monotomidae Monotomidae is a family of beetles in the superfamily Cucujoidea. The family is found worldwide, with approximately 240 species in 33 genera. The ecological habits of the family are diverse, with different members of the group being found under t ...
Laporte, 1840 *****Family Hobartiidae Sen Gupta and Crowson, 1966 *****Family
Cryptophagidae Cryptophagidae is a family of beetles with representatives found in all biogeographic realms. Members of this family are commonly called silken fungus beetles and both adults and larvae appear to feed exclusively on fungi although in a wide vari ...
Kirby, 1826 *****Family Agapythidae Sen Gupta and Crowson, 1969 *****Family Priasilphidae Crowson, 1973 *****Family Phloeostichidae Reitter, 1911 *****Family
Silvanidae Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. V ...
Kirby, 1837 *****Family Cucujidae Latreille, 1802 *****Family Myraboliidae Lawrence and Britton, 1991 *****Family Cavognathidae Sen Gupta and Crowson, 1966 *****Family Lamingtoniidae Sen Gupta and Crowson, 1969 *****Family
Passandridae Passandridae, the "parasitic flat bark beetles," are a family of beetles notable for being one of the very few beetle families with larvae that are, as far as known, exclusively ectoparasitic on the immature stages of other beetles and Hymenopter ...
Blanchard, 1845 *****Family
Phalacridae The Phalacridae are a family of beetles commonly called the shining flower beetles, They are often found in composite flowers. They are oval-shaped, usually tan, and about 2 mm in length. Most species feed on fungus, although a number feed ...
Leach, 1815 *****Family Propalticidae Crowson, 1952 *****Family
Laemophloeidae Laemophloeidae, lined flat bark beetles, is a beetle family in the superfamily Cucujoidea characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia. Size r ...
Ganglbauer, 1899 *****Family Tasmosalpingidae Lawrence and Britton, 1991 *****Family Cyclaxyridae Gimmel, Leschen and Ślipiński, 2009 *****Family Kateretidae Kirby, 1837 *****Family
Nitidulidae The sap beetles, also known as Nitidulidae, are a family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antennae. Some have red or yellow spots or bands. They feed mainly on decaying vegetable ma ...
Latreille, 1802 *****Family Smicripidae Horn, 1880 *****Family
Bothrideridae Bothrideridae is a family of beetles in the superfamily Coccinelloidea. They are known commonly as the cocoon-forming beetles or dry bark beetles. They occur worldwide with most native to the Old World tropics. In older literature, the family was ...
Erichson, 1845 *****Family Cerylonidae Billberg, 1820 *****Family Alexiidae Imhoff, 1856 *****Family Discolomatidae Horn, 1878 *****Family
Endomychidae Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus '' Endomychus'', a genus ere ...
Leach, 1815 *****Family
Coccinellidae Coccinellidae () is a widespread family (biology), family of small beetles. They are commonly known as ladybugs in North America and ladybirds in the United Kingdom; "lady" refers to Mary, mother of Jesus, mother Mary. Entomologists use the ...
Latreille, 1807 (ladybirds or lady beetles) *****Family
Corylophidae Corylophidae is a family of minute hooded beetles, sometimes called minute fungus beetles, in the superfamily Coccinelloidea. There are about 18 genera and at least 120 described species in Corylophidae. They feed on microfungi such as molds, ...
LeConte, 1852 *****Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh, Whiting and Miller, 2010 *****Family
Latridiidae Latridiidae (sometimes spelled "Lathridiidae") is a family of tiny, little-known beetles commonly called minute brown scavenger beetles or fungus beetles. The number of described species currently stands at around 1050 in 29 genera but the number ...
Erichson, 1842 ****Superfamily
Tenebrionoidea The Tenebrionoidea are a very large and diverse superfamily of beetles. It generally corresponds to the Heteromera of earlier authors. Taxonomy Tenebrionoidea contains the following families: * Aderidae Winkler 1927 (ant-like leaf beetles) ...
Latreille, 1802 *****Family
Mycetophagidae The Mycetophagidae or hairy fungus beetles are a family of beetles in the superfamily Tenebrionoidea. The different species are between 1.0 and 6.5 mm in length. The larvae and adults live in decaying leaf litter, fungi, and under bark. Most ...
Leach, 1815 *****Family Archeocrypticidae Kaszab, 1964 *****Family
Pterogeniidae Pterogeniidae is a family of beetles belonging to the superfamily Tenebrionoidea. They are found in South Asia, Southeast Asia, China, New Guinea and the Solomon Islands. Adults and larvae have been found associated with the fruiting bodies of po ...
Crowson, 1953 *****Family
Ciidae The minute tree-fungus beetles, family Ciidae, are a sizeable group of beetles which inhabit Polyporales bracket fungi or coarse woody debris. Most numerous in warmer regions, they are nonetheless widespread and a considerable number of species ...
Leach, 1819 *****Family
Tetratomidae Tetratomidae is a small family of beetles sometimes called polypore fungus beetles. The family consists of several genera, most of which used to be in the family Melandryidae. Tetratomidae can be found worldwide. Their food consists of fruitin ...
Billberg, 1820 *****Family
Melandryidae Melandryidae is a family of beetles in the superfamily Tenebrionoidea. Members of the family are found worldwide, with around 420 species in 60 genera. Larvae and adults are generally associated with rotting wood and wood-decomposing fungi. Gen ...
Leach, 1815 *****Family
Mordellidae The Mordellidae are a family of beetles commonly known as tumbling flower beetles for the typical irregular movements they make when escaping predators, or as pintail beetles due to their abdominal tip which aids them in performing these tumblin ...
Latreille, 1802 (tumbling flower beetles) *****Family
Ripiphoridae Ripiphoridae (formerly spelled Rhipiphoridae) is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic p ...
Laporte, 1840 *****Family
Zopheridae Zopheridae is a family of beetles belonging to Tenebrionoidea. It has grown considerably in recent years as the members of two other families are now included (Monommatidae and Colydiidae), as subfamilies or (in the former case) even as tribe ( ...
Solier, 1834 *****Family Ulodidae Pascoe, 1869 *****Family
Promecheilidae Promecheilidae is a family of beetles in the superfamily Tenebrionoidea. Perimylopidae is considered a synonym. They are found in southern South America and associated archipelagos like South Georgia and the Falkland Islands, Falklands, New Zeala ...
Lacordaire, 1859 *****Family
Chalcodryidae The Chalcodryidae are a family of beetles in the superfamily Tenebrionoidea The Tenebrionoidea are a very large and diverse superfamily of beetles. It generally corresponds to the Heteromera of earlier authors. Taxonomy Tenebrionoidea co ...
Watt, 1974 *****Family Trachelostenidae Lacordaire, 1859 *****Family
Tenebrionidae Darkling beetle is the common name for members of the beetle family Tenebrionidae, comprising over 20,000 species in a cosmopolitan distribution. Taxonomy ''Tenebrio'' is the Latin generic name that Carl Linnaeus assigned to some flour beetles ...
Latreille, 1802 (darkling beetles) *****Family
Prostomidae Prostomidae is a family of beetles with no vernacular common name, though recent authors have coined the name jugular-horned beetles. They are often found in dead wood. The family consist of two extant genera with about 20 species. '' Prostomis a ...
Thomson, 1859 *****Family
Synchroidae The Synchroidae are a small family of tenebrionoid beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguish ...
Lacordaire, 1859 *****Family
Stenotrachelidae Stenotrachelidae, commonly called false longhorn beetles is a family of beetles in the superfamily Tenebrionoidea. They are native to the Northern Hemisphere. The larvae feed on heavily decomposed wood, while the adults are likely short lived a ...
Thomson, 1859 *****Family
Oedemeridae The family Oedemeridae is a cosmopolitan group of beetles commonly known as false blister beetles, though some recent authors have coined the name pollen-feeding beetles. There are some 100 genera and 1,500 species in the family, mostly associate ...
Latreille, 1810 *****Family
Meloidae Blister beetles are beetles of the family Meloidae, so called for their defensive secretion of a blistering agent, cantharidin. About 7,500 species are known worldwide. Many are conspicuous and some are aposematically colored, announcing their ...
Gyllenhal, 1810 (blister beetles) *****Family
Mycteridae The family Mycteridae is a small group of tenebrionoid beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distingu ...
Perty, 1840 (palm beetles and flower beetles) *****Family
Boridae The Boridae are a small family of tenebrionoid beetles with no vernacular common name, though recent authors have coined the name conifer bark beetles. The family contains three genera. '' Boros'' is native to North America and northern Eurasia, ...
Thomson, 1859 *****Family
Trictenotomidae The Trictenotomidae are a small family of beetles in the superfamily Tenebrionoidea, containing fifteen species in two genera. Most species are found in the Oriental realm where they live in montane forest habitats. The family is considered, base ...
Blanchard, 1845 *****Family
Pythidae The family Pythidae is a small group of tenebrionoid beetles with no vernacular common name, though recent authors have coined the name dead log bark beetles. There are seven genera, which are largely native to the mid-high latitude regions of t ...
Solier, 1834 *****Family
Pyrochroidae Fire-coloured beetles is the common name for members of the tenebrionoid family Pyrochroidae. The family is found worldwide, and is most diverse at temperate latitudes. Adults measure ; larvae reach . Larvae of Pyrochroinae are found associated ...
Latreille, 1806 *****Family
Salpingidae Salpingidae or narrow-waisted bark beetles is a family of beetles in the superfamily Tenebrionoidea. The species are small, about 1.5 – 7 mm in length. The family is globally distributed and consists of about 45 genera and 300 species, w ...
Leach, 1815 *****Family Anthicidae Latreille, 1819 *****Family Aderidae Csiki, 1909 *****Family
Scraptiidae The family Scraptiidae is a small group of tenebrionoid beetles sometimes called false flower beetles. There are about 400 species in 30 genera with a world-wide distribution. The adults are found on flowers, sometimes in large numbers, but are a ...
Gistel, 1848 ****Clade Phytophaga *****Superfamily
Chrysomeloidea The Chrysomeloidea are an enormous superfamily of beetles, with tens of thousands of species. The largest families are Cerambycidae, long-horned beetles, with more than 35,000 species, and Chrysomelidae, leaf beetles, with more than 13,000 speci ...
Latreille, 1802 ******Family Oxypeltidae Lacordaire, 1868 ******Family Vesperidae Mulsant, 1839 ******Family Disteniidae Thomson, 1861 ******Family
Cerambycidae The longhorn beetles (Cerambycidae), also known as long-horned or longicorns (whose larvae are often referred to as roundheaded borers), are a large family of beetles, with over 35,000 species described. Most species are characterized by anten ...
Latreille, 1802 (longhorn beetles) ******Family Megalopodidae Latreille, 1802 ******Family Orsodacnidae Thomson, 1859 ******Family
Chrysomelidae The beetle family Chrysomelidae, commonly known as leaf beetles, includes over 37,000 (and probably at least 50,000) species in more than 2,500 genera, making it one of the largest and most commonly encountered of all beetle families. Numerous s ...
Latreille, 1802 (leaf beetles) *****Superfamily Curculionoidea Latreille, 1802 ******Family
Nemonychidae Nemonychidae is a small family of weevils, placed within the ''primitive weevil'' group because they have straight rather than geniculate (elbowed) antennae. They are often called pine flower weevils. As in the Anthribidae, the labrum appears ...
Bedel, 1882 ******Family
Anthribidae Anthribidae is a family of beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other i ...
Billberg, 1820 ******Family † Ulyanidae Zherikhin, 1993 ******Family
Belidae Belidae is a family of weevils, called belids or primitive weevils because they have straight antennae, unlike the "true weevils" or Curculionidae which have geniculate (elbowed) antennae. They are sometimes known as " cycad weevils", but this pr ...
Schönherr, 1826 ******Family
Caridae Caridae is a small Gondwanan family of weevils. They are considered part of the primitive weevil group, because they have straight rather than geniculate (elbowed) antennae. The insertion of the antennae on the rostrum cannot be seen from above ...
Thompson, 1992 ******Family
Attelabidae The Attelabidae is a widespread family of weevils. They are among the primitive weevils, because of their straight antennae, which are inserted near the base of the rostrum. The prothorax is much narrower than the base of the elytra on the abdo ...
Billberg, 1820 ******Family
Brentidae Brentidae is a cosmopolitan distribution, cosmopolitan family of primarily xylophagy, xylophagous beetles also known as straight-snouted weevils; they have sometimes been called "primitive weevils", but this name also applies to taxa such as the ...
Billberg, 1820 ******Family
Brachyceridae Brachyceridae is a family (biology), family of weevils. There are at least 150 genera in the family Brachyceridae. It has been treated as a subfamily of Curculionidae. Cladogram See also * List of Brachyceridae genera References * Bisby F ...
Billberg, 1820 ******Family
Curculionidae The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae. Th ...
Latreille, 1802 (snout beetles, weevils, and bark beetles)


References

* Lawrence, J.F., Newton, A.F. Jr. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references, and data on family-group names), pp. 779–1006. In: Pakaluk, J., Slipinski, S.A. (eds.), ''Biology, phylogeny, and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson''. Muzeum i Instytut Zoologii PAN, Warszawa. * Ross H. Arnett, Jr. and Michael C. Thomas, ''
American Beetles ''American Beetles'' is a comprehensive description of the beetles of North America north of the tropical area of Mexico. It was started by Ross H. Arnett, Jr. as an update of his classic ''The Beetles of the United States''; along with Michael ...
'' (
CRC Press The CRC Press, LLC is an American publishing group that specializes in producing technical books. Many of their books relate to engineering, science and mathematics. Their scope also includes books on business, forensics and information technol ...
, 2001–2002) {{Coleoptera *