The Satyrini is one of the tribes of the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a distinct family, Satyridae. This group contains nearly half of the known divers ...

. It includes about 2200 species and is therefore the largest tribe in the subfamily which comprises 2500 species.

Distribution

Satyrini butterflies have a worldwide distribution, but the distribution pattern differs between

subtribes Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants Plants are predominantly Photosynthesis, photosynthetic eukaryotes of th ...

. Some subtribes are almost restricted to a single

biogeographic region An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...

, such as the

Pronophilina Pronophilina is a Neotropical subtribe of butterflies of the subfamily Satyrinae. They are a species-rich group with highest diversity in the tropical and subtropical mountains, especially the Andes. Before 1970, they were poorly studied, but re ...

, which is found only in Andean

cloud forests A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud ...

from Venezuela to

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

Biology

The larval food plants of many species in this tribe are grasses, i.e. Poaceae. It is considered that the Satyrini diversified at about the same time as the grasses did, and that the

radiation In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or through a material medium. This includes: * ''electromagnetic radiation'', such as radio waves, microwaves, infrared, visi ...

of the tribe is therefore closely related to the evolution of the grasses. In contrast, the tribe has a few genera which show uncommon feeding preferences. Three genera, ''

Euptychia ''Euptychia'' is a genus of satyrid butterflies found in the Neotropical realm. The genus was erected by Jacob Hübner Jacob Hübner (20 June 1761 – 13 September 1826, in Augsburg) was a German entomologist. He was the author of ''Sammlung E ...

'', ''

Ragadia ''Ragadia'' is a genus of brush-footed butterflies (family Nymphalidae). This genus is one of those commonly called ringlets. Species * ''Ragadia annulata'' * ''Ragadia crisilda'' - striped ringlet * ''Ragadia critias'' * ''Ragadia crito'' * ''R ...

'' and ''

Acrophtalmia Acrophtalmia is a genus of butterflies in the family Nymphalidae. Species *'' Acrophtalmia artemis'' C. & R. Felder, 1861 *'' Acrophtalmia leuce'' C. & R. Felder, 867 __NOTOC__ Year 867 (Roman numerals, DCCCLXVII) was a common year startin ...

'', feed on

Lycopsida Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching s ...

, and moreover, some species of ''Euptychia'' have been reported to feed on

mosses Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and horn ...

Neckeraceae Neckeraceae is a moss family in the order Hypnales. There are about 200 species native to temperate and tropical regions. Most grow on rocks, or other plants. Description Members of the family are usually large and glossy plants with creeping st ...

. This is interesting cases as these genera are not considered to be very closely related to each other.

Taxonomy

Speckled wood (Pararge aegeria aegeria) male

Mycalesis francisca formosana high altitude ventral view 20150705

Marbled white (Melanargia galathea) male

The taxonomy of the subfamily Satyrinae has recently undergone a significant revision. showed that most of the traditionally well known group are

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

or polyphyletic and developed a new taxonomy, based on

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

. The classification scheme transferred many genera traditionally placed in the

Elymniini The Elymniini is one of the tribes of the subfamily Satyrinae. If the subfamily Satyrinae elevate to family status, this tribe shall be treated as subfamily Elymniinae. Elymniini was formerly a large group, but recently, it is considered to be ...

to the Satyrini, and later studies have largely followed this classification. Thus, the Satyrini is now considered to include the genera previously placed in the Elymniini, such as '' Lethe'', ''

Pararge ''Pararge'' is a genus of butterflies of the family Nymphalidae. Species * ''Pararge aegeria'' – speckled wood (Linnaeus, 1758) * ''Pararge xiphia'' – Madeiran speckled wood (Fabricius, 1775) * ''Pararge xiphioides'' – Canary s ...

'' and ''

Mycalesis ''Mycalesis'', the bushbrowns, are a genus of brush-footed butterflies. They are common in the warm regions from Central Asia to Australia, and have a high diversity in South Asia and the Wallacea. They are notably polymorphic, with wet- and ...

''. However, the phylogenetic statuses of some subtribes within the tribe are still unclear and the revision of the classification is likely to continue. ---- According to , the Satyrini includes the following

and

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

: ;

Parargina Parargina is a subtribe of butterflies of the subfamily Satyrinae.Parargina
Tree of Life Web Project

Genera

Moore, 1893 :* ''

Chonala ''Chonala'' is a genus of butterflies of the family Nymphalidae found in eastern Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasi ...

'' Moore, 1893 :* ''

Kirinia ''Kirinia'' is a genus of butterflies of the family Nymphalidae found in Europe and Asia. Species Listed alphabetically:''Kirinia''
...

'' Westwood, 1841 :* '' Lasiommata'' Tutt, 1896 :* ''

Lopinga ''Lopinga'' is a genus of butterflies of the family Nymphalidae. Species * ''Lopinga achine ''Lopinga achine'', the woodland brown, is a Palearctic butterfly in the family Nymphalidae. Description in Seitz P. ''achine'' Scop. (= ''dejanir ...

'' Moore, 1893 :* ''

Orinoma ''Orinoma'' is a genus of butterflies of the family Nymphalidae found in Asia. Species Listed alphabetically:''Orinoma''
at the ''T ...

'' Gray, 1846 :* ''

'' Hübner, 1819 :* ''

Rhaphicera ''Rhaphicera'' is a genus of butterflies of the family Nymphalidae found in the Indomalayan realm The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts ...

'' Butler, 1867 :* ''

Tatinga ''Tatinga thibetana'' is only species in the butterfly genus ''Tatinga'' of the family Nymphalidae found in Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right ...

'' Moore, 1893 :* ''

Nosea ''Nosea hainanensis'' is only species in the butterfly genus ''Nosea'' of the family Nymphalidae endemic to China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous count ...

'' Moore, 1893 ; Lethina Reuter, 1896 :* ''

Aphysoneura ''Aphysoneura'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the su ...

'' Karsch, 1894 :* '' Lethe'' Hübner, 1819 :* ''

Neope ''Neope'' is a genus of butterflies of the family Nymphalidae found in Asia.Ninguta'' Moore, 1892 :* '' Ptychandra'' Felder & Felder, 1861 :* ''

Mandarinia ''Mandarinia'' is a monotypic butterfly genus in the family Nymphalidae (Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly consi ...

'' Leech, 1892 ; Mycalesina Reuter, 1896 :* '' Lohora'' Moore, 1880 :* '' Brakefieldia'' Aduse-Poku, Lees & Wahlberg, 2016 :* ''

Telinga The land of Telugu people was referred during ancient times as ''Āndhra dēśa'' (country of Andhra) and ''Trilingadēśa'' (country of Trilinga). The word ''Telugu'' is ''believed'' to have been derived from ''trilinga'', as in ''Trilinga De ...

'' Moore, 1880 :* '' Heteropsis'' Westwood, 1850 :* '' Culapa'' Moore, 1879 :* '' Mydosama'' Moore, 1880 :* ''

'' Hübner, 1818 :* '' Devyatkinia'' Monastyrskii & Uémura, 2016 :* '' Hallelesis'' Condamin, 1961 :* ''

Bicyclus ''Bicyclus'' is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae. The species are found in the Afrotropical realm. Species *''Bicyclus abnormis'' (Dudgeon, 1909) *''Bicyclus albocincta'' (Rebel, 1914) *''Bicyclus alboplag ...

'' Kirby, 1871 ; Eritina Miller, 1968 :* '' Coelites'' Westwood, 1850 :* '' Erites'' Westwood, 1851 :* ''

Orsotriaena ''Orsotriaena medus'' is a butterfly found in south Asia, southeast Asia, and Australia. It is the only species in the genus ''Orsotriaena'', first described by Hans Daniel Johan Wallengren in 1858. Historical name The butterfly has historica ...

'' Wallengren, 1858 :* ''

Zipaetis ''Zipaetis'' is a genus of satyrid butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly colou ...

'' Hewitson, 1863 ; Ragadiina Herrich-Schäffer, 1864 :* ''

'' Felder & Felder, 1861 :* ''

Acropolis An acropolis was the settlement of an upper part of an ancient Greek city, especially a citadel, and frequently a hill with precipitous sides, mainly chosen for purposes of defense. The term is typically used to refer to the Acropolis of Athens, ...

'' Hemming, 1934 :* ''

'' Westwood, 1851 ; Coenonymphina Tutt, 1896 :* ''

Coenonympha ''Coenonympha'' is a butterfly genus belonging to the Coenonymphina, a subtribe of the browns (Satyrinae). The latter are a subfamily of the brush-footed butterflies (Nymphalidae). As a rule, Palearctic species are colloquially called heaths, wh ...

'' Hübner, 1819 :* ''

Sinonympha ''Sinonympha '' is a monotypic butterfly genus in the family Nymphalidae (subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were for ...

'' Lee, 1994 :* '' Oressinoma'' Doubleday, 1849 :* ''

Altiapa ''Platypthima'' is a genus of satyrid butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly co ...

'' Parsons, 1986 :* '' Argynnina'' Butler, 1867 :* ''

Argyronympha ''Argyronympha'' is a genus of satyrid butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly c ...

'' Mathew, 1886 :* ''

Argyrophenga ''Argyrophenga'' is a genus of butterflies that are endemic to New Zealand.

'' Doubleday, 1845 :* ''

Dodonidia The forest ringlet (''Dodonidia helmsii''), also known as Helms' butterfly, is a rare butterfly of the family Nymphalidae endemic to New Zealand. It is the only species in the genus ''Dodonidia''. Taxonomy The forest ringlet was first describ ...

'' Butler, 1884 :* '' Erebiola'' Fereday, 1879 :* '' Erycinidia'' Rothschild & Jordan, 1905 :* '' Geitoneura'' Butler, 1867 :* '' Harsiesis'' Fruhstorfer, 1911 :* '' Heteronympha'' Wallengren, 1858 :* ''

Hyalodia ''Hyalodia'' is a monotypic butterfly genus of the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a disti ...

'' Jordan, 1924 :* ''

Hypocysta ''Hypocysta'' is a genus of butterflies in the family Nymphalidae. Species The genus contains 12 species. * '' Hypocysta adiante'' (Hübner, 1825) – orange ringlet * '' Hypocysta angustata'' Waterhouse & Lyell, 1914 – black and white ring ...

'' Westwood, 1851 :* ''

Lamprolenis ''Lamprolenis'' is a monotypic butterfly genus of the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a d ...

'' Godman & Salvin, 1881 :* ''

Nesoxenica ''Nesoxenica'' is a monotypic butterfly genus of the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a dis ...

'' Waterhouse & Lyell, 1914 :* ''

Oreixenica ''Oreixenica'' is a genus of butterflies in the family Nymphalidae. The genus contains six species.

'' Waterhouse & Lyell, 1914 :* ''

Paratisiphone ''Paratisiphone'' is a monotypic butterfly genus of the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a ...

'' Watkins, 1928 :* ''

Percnodaimon ''Percnodaimon merula'', the black mountain ringlet, is a satyrid butterfly in the family Nymphalidae. It is currently the only recognised species in the monotypic genus ''Percnodaimon,'' endemic to New Zealand, although there may be other un ...

'' Butler, 1876 :* ''

Platypthima ''Platypthima'' is a genus of satyrid butterflies. Species Listed alphabetically:Tisiphone'' Hübner, 1819 ;

Euptychiina The butterfly subtribe Euptychiina ( Lepidoptera: Nymphalidae) is a diverse group within the tribe Satyrini, occurring throughout Central and South America, in addition to a few species known from North America. Euptychiina is a predominantly lo ...

Reuter, 1896 :* ''

'' Hübner, 1818 :* ''

Palaeonympha ''Palaeonympha'' is a monotypic butterfly genus of the subfamily Satyrinae in the family Nymphalidae. Its one species, ''Palaeonympha opalina'', is found in Taiwan

'' Butler, 1871 :* ''

Amphidecta ''Amphidecta'' is a genus of satyrid butterfly found in the Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of t ...

'' Butler, 1867 :* ''

Archeuptychia ''Archeuptychia'' is a monotypic satyrid butterfly genus described by Walter Forster in 1964. Its sole member, ''Archeuptychia cluena'' is found in Brazil. The original name of this species was ''Papilio cluena'' as described by Dru Drury i ...

'' Forster, 1964 :* ''

Amiga Amiga is a family of personal computers introduced by Commodore in 1985. The original model is one of a number of mid-1980s computers with 16- or 32-bit processors, 256 KB or more of RAM, mouse-based GUIs, and significantly improved graphi ...

'' Nakahara, Willmott & Espeland, 2019 :* '' Caeruleuptychia'' Forster, 1964 :* '' Capronnieria'' Forster, 1964 :* '' Cepheuptychia'' Forster, 1964 :* ''

Chloreuptychia ''Chloreuptychia'' is a genus of satyrid butterfly found in the Neotropical realm. Species Listed alphabetically:Cissia'' Doubleday, 1848 :* ''

Coeruleotaygetis ''Taygetina peribaea'' is a species of butterfly in the subfamily Satyrinae. It is found in the Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the t ...

'' Forster, 1964 :* ''

Cyllopsis ''Cyllopsis'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. The genus was erected by Rudolf Felder in 1869. Species Listed alphabetically: *''Cyllopsis argentella'' (Butler & H. Druce, 1872) *''Cyllopsis caballeroi ...

'' Felder, 1869 :* '' Erichthodes'' Forster, 1964 :* '' Euptychoides'' Forster, 1964 :* ''

Forsterinaria ''Forsterinaria'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Graphita'' Nakahara, Marín & Barbosa 2016 :* ''

Godartiana ''Godartiana'' is a genus of satyrid butterfly found in the Neotropical realm. Species Listed alphabetically:

'' Forster, 1964 :* ''

Harjesia ''Harjesia'' is a genus of satyrid butterfly found in the Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the ...

'' Forster, 1964 :* ''

Hermeuptychia ''Hermeuptychia'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. They are a widespread, cryptic genus, with Cytochrome c oxidase subunit I species delineation methods implying much greater species diversity than curr ...

'' Forster, 1964 :* '' Huberonympha'' Viloria & Costa, 2016 :* ''

Inbio The Instituto Nacional de Biodiversidad (INBio) is the national institute for biodiversity and conservation in Costa Rica. Created at the end of the 1980s, and despite having national status, it is a privately run institution that works closely ...

'' Nakahara & Espeland, 2015 :* '' Magneuptychia'' Forster, 1964 :* '' Megeuptychia'' Forster, 1964 :* '' Megisto'' Hübner, 1819 :* ''

Moneuptychia ''Moneuptychia'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Neonympha ''Neonympha'' is a Nearctic and Neotropical genus of satyrid butterflies. Species Listed alphabetically: *''Neonympha areolatus'' (Smith, 1797) *''Neonympha helicta'' (Hübner, 808 *''Neonympha mitchellii'' French, 1889 *'' "Neonympha" lupita' ...

'' Hübner 1818 :* '' Nhambikuara'' Freitas, Barbosa & Zacca, 2018 :* '' Optimandes'' Marín, Nakahara & Willmott, 2019 :* '' Orotaygetis'' Nakahara & Zacca, 2018 :* '' Paramacera'' Butler, 1868 :* '' Parataygetis'' Forster, 1964 :* ''

Pareuptychia ''Pareuptychia'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Paryphthimoides'' Forster, 1964 :* ''

Pharneuptychia ''Pharneuptychia'' is a genus of Satyrinae, satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Pindis'' Felder, 1869 :* ''

Posttaygetis ''Posttaygetis'' is a monotypic butterfly genus of the subfamily Satyrinae in the family Nymphalidae. Its one species is ''Posttaygetis penelea'', which is found in the Neotropical realm The Neotropical realm is one of the eight biogeograph ...

'' Forster, 1964 :* ''

Prenda ''Prenda'' is a monotypic butterfly genus of subfamily Satyrinae in the family Nymphalidae. Its one species ''Prenda clarissa'' is found in the Neotropical realm. The generic name ''Prenda'' "designates a young woman from Rio Grande do Sul, on ...

'' Freitas & Mielke, 2011 :* ''

Pseudeuptychia ''Pseudeuptychia'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Pseudodebis ''Pseudodebis'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:

'' Forster, 1964 :* ''

Rareuptychia ''Rareuptychia'' is a monotypic butterfly genus of the subfamily Satyrinae in the family Nymphalidae. Its one species, ''Rareuptychia clio'', is found in the Neotropical realm The Neotropical realm is one of the eight biogeographic realms c ...

'' Forster, 1964 :* '' Satyrotaygetis'' Forster, 1964 :* '' Sepona'' Freitas & Barbosa, 2016 :* '' Splendeuptychia'' Forster, 1964 :* '' Stegosatyrus'' Zacca, Mielke & Pyrcz, 2013 :* '' Stevenaria'' Viloria, Costa, Neild & Nakahara, 2016 :* ''

Taydebis ''Taydebis'' is a monotypic butterfly genus in the subfamily Satyrinae in the family Nymphalidae. The genus was established by André Victor Lucci Freitas in 2003. Its one species, ''Taydebis peculiaris'', was found in Brazil Brazil ( pt, ...

'' Freitas, 2003 :* ''

Taygetina ''Taygetina'' is a monotypic butterfly genus of the subfamily Satyrinae in the family Nymphalidae. The species in '' Coeruleotaygetis'' have been included in ''Taygetina''. Its one species, ''Taygetina banghaasi'', is found in the Neotropical rea ...

'' Forster, 1964 :* ''

Taygetis ''Taygetis'' is a genus of satyrid butterflies found in the Neotropical realm. Taxonomy and evolution The genus appeared as polyphyletic,, 2013: Systematics and evolutionary history of butterflies in the “''Taygetis'' clade” (Nymphalidae: S ...

'' Hübner, 1819 :* ''

Taygetomorpha ''Taygetomorpha'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Yphthimoides ''Yphthimoides'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:Zischkaia ''Zischkaia'' is a genus of satyrid butterflies found in the Neotropical realm. Species Listed alphabetically:

'' Forster, 1964 ;

Erebiina The Satyrini is one of the tribes of the subfamily Satyrinae. It includes about 2200 species and is therefore the largest tribe in the subfamily which comprises 2500 species. Distribution Satyrini butterflies have a worldwide distribution, bu ...

Tutt, 1896 :* ''

Erebia ''Erebia'' is a Holarctic genus of brush-footed butterflies, family Nymphalidae. Most of the about 90–100 species (see also below) are dark brown or black in color, with reddish-brown to orange or more rarely yellowish wing blotches or band ...

'' Elwes, 1899 ; Maniolina Grote, 1897 :* ''

Aphantopus ''Aphantopus'' is a butterfly genus of the Satyrinae. The genus is confined to the Palearctic. Species *''Aphantopus arvensis'' (Oberthür, 1876) (western China) **''Aphantopus arvensis arvensis'' **''Aphantopus arvensis campana'' Leech, 1892 (c ...

'' Wallengren, 1853 :* ''

Cercyonis ''Cercyonis'' is a genus of butterflies of the subfamily Satyrinae in the family Nymphalidae found in North America. They are commonly called wood-nymphs or wood nymphs. Species Listed alphabetically:Hyponephele'' Muschamp, 1915 :* ''

Maniola ''Maniola'' is a genus of butterflies within the family Nymphalidae.

'' Schrank, 1801 :* ''

Pyronia ''Pyronia'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae. Description These butterflies have vestigial forelegs that cannot be used for walking. Male forelegs exhibit 2 tarsal joints, while female forelegs h ...

'' Hübner, 1819 ; Melanargiina Wheeler, 1903 :* ''

Melanargia ''Melanargia'' is a genus of butterflies belonging to the family Nymphalidae and the subfamily Satyrinae (formerly family Satyridae). This genus, described by Johann Wilhelm Meigen in 1828, is the only genus in the subtribe Melanargiina, Wheel ...

'' Meigen, 1828 ;

Reuter, 1896 :* '' Altopedaliodes'' Forster, 1964 :* '' Antopedaliodes'' Forster, 1964 :* '' Apexacuta'' Pyrcz, 2004 :* '' Arhuaco'' Adams & Bernard, 1977 :* '' Cheimas'' Thieme, 1907 :* '' Corades'' Doubleday, 1849 :* '' Corderopedaliodes'' Forster, 1964 :* '' Daedalma'' Hewitson, 1858 :* '' Dangond'' Adams & Bernard, 1979 :* ''

Drucina ''Drucina'' is a Neotropical butterfly genus in the family Nymphalidae.

'' Butler, 1872 :* '' Druphila'' Pyrcz, 2004 :* '' Eretris'' Thieme, 1905 :* '' Eteona'' Doubleday, 1848 :* '' Foetterleia'' Viloria, 2004 :* '' Junea'' Hemming, 1964 :* ''

Lasiophila ''Lasiophila'' is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae found in South America. Species Listed alphabetically:Lymanopoda ''Lymanopoda'' is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae found in north-western South America. Species Listed alphabetically:Mygona'' Thieme, 1907 :* '' Neopedaliodes'' Viloria, Miller & Miller 2004 :* '' Oxeoschistus'' Butler, 1867 :* '' Panyapedaliodes'' Forster, 1964 :* '' Paramo'' Adams & Bernard, 1977 :* '' Parapedaliodes'' Forster, 1964 :* ''

Pedaliodes ''Pedaliodes'' is a genus of butterflies from the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a distin ...

'' Butler, 1867 :* '' Pherepedaliodes'' Forster, 1964 :* '' Physcopedaliodes'' Forster, 1964 :* '' Praepedaliodes'' Forster, 1964 :* '' Praepronophila'' Forster, 1964 :* '' Proboscis'' Thieme, 1907 :* '' Pronophila'' Doubleday, 1849 :* '' Protopedaliodes'' Viloria & Pyrcz, 1994 :* '' Pseudomaniola'' Röber, 1889 :* '' Punapedaliodes'' Forster, 1964 :* '' Steremnia'' Thieme, 1905 :* '' Steroma'' Westwood, 1850 :* '' Steromapedaliodes'' Forster, 1964 :* '' Thiemeia'' Weymer, 1912 :* ''

Diaphanos ''Diaphanos'' is a genus of butterflies in the family Nymphalidae. There are three recognised species in the Neotropics, and one undescribed subspecies of ''D. huberi''.Gerardo Lamas 2004 ''Atlas of Neotropical Lepidoptera. CheckList: Part 4A Hes ...

'' Adams & Bernard, 1981 :* ''

Ianussiusa ''Ianussiusa'' is a monotypic butterfly genus in the family Nymphalidae. It contains only one species, ''Ianussiusa maso'', which is found in Colombia and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, li ...

'' Pyrcz & Viloria, 2004 :* '' Idioneurula'' Strand, 1932 :* '' Manerebia'' Staudinger, 1897 :* ''

Neomaniola ''Neomaniola'' is a Neotropical butterfly genus in the family Nymphalidae. The genus is monotypic containing the single species ''Neomaniola euripides'', which is found in Bolivia and Argentina Argentina (), officially the Argentine Repu ...

'' Hayward, 1949 :* '' Argyrophorus'' Blanchard, 1852 :* '' Auca'' Hayward, 1953 :* '' Chillanella'' Herrera, 1966 :* '' Cosmosatyrus'' Felder & Felder, 1867 :* '' Elina'' Blanchard, 1852 :* ''

Faunula ''Faunula'' is a Neotropical genus of butterflies in the nymphalid subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly co ...

'' Felder & Felder, 1867 :* '' Haywardella'' Herrera, 1966 :* '' Homoeonympha'' Felder & Felder, 1867 :* '' Nelia'' Hayward, 1953 :* '' Neomaenas'' Wallengren, 1858 :* '' Neosatyrus'' Wallengren, 1858 :* ''

Pampasatyrus ''Pampasatyrus'' is a Neotropical genus of butterflies in the family Nymphalidae.Quilaphoestosus'' Herrera, 1966 :* '' Spinantenna'' Hayward, 1953 :* '' Tetraphlebia'' Felder & Felder, 1867 :* '' Gyrocheilus'' Butler, 1867 ; Satyrina Boisduval, 1833 :* '' Argestina'' Riley, 1923 :* ''

Davidina ''Davidina '' is a genus of butterflies in the family Nymphalidae (subfamily Satyrinae). (central China: Shaanxi Shaanxi (alternatively Shensi, see #Name, § Name) is a landlocked Provinces of China, province of China. Officially part of No ...

'' Oberthür, 1879 :* '' Berberia'' de Lesse, 1951 :* '' Hipparchia'' Fabricius, 1807 :* '' Kanetisa'' Moore, 1893 :* '' Aulocera'' Butler, 1867 :* ''

Karanasa Karanasa is an Old World genus of subfamily Satyrinae (family Nymphalidae).

'' Moore, 1893 :* ''

Oeneis ''Oeneis'' (the Arctics or graylings) is a butterfly genus of the Satyrinae. All but one of its members are Arctic, sub-Arctic or high-altitude alpine in distribution. Some of the members of the genus are among the butterflies that can get along ...

'' Hübner, 1819 :* '' Paroeneis'' Moore, 1893 :* ''

Arethusana ''Arethusana'' is a butterfly genus from the subfamily Satyrinae of the brush-footed butterfly family (Nymphalidae). It is composed of only one species, ''Arethusana arethusa'', the false grayling. Subspecies Subspecies include:

'' de Lesse, 1951 :* ''

Brintesia ''Brintesia'' is a monotypic butterfly genus in the family Nymphalidae and subfamily Satyrinae. Its one species is ''Brintesia circe'', the great banded grayling. Description ''Brintesia circe'' reaches on average wingspan of . Its wings are mai ...

'' Fruhstorfer, 1911 :* '' Minois'' Hübner, 1819 :* ''

Chazara ''Chazara'' is a butterfly genus from the subfamily Satyrinae of the brush-footed butterfly family (Nymphalidae). The most well-known is the hermit butterfly ('' C. briseis''). ''C. briseis'' can be found as far west as Morocco and as far east ...

'' Moore, 1893 :* '' Pseudochazara'' de Lesse, 1951 :* '' Satyrus'' Latreille, 1810 ; Ypthimina Reuter, 1896 :* ''

Cassionympha ''Cassionympha'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae. Species *'' Cassionympha camdeboo'' (Dickson, 1981) *'' Cassionympha cassius'' (Godart, 824 __NOTOC__ Year 824 ( DCCCXXIV) was a leap year s ...

'' van Son, 1955 :* '' Coenyra'' Hewitson, 1865 :* ''

Coenyropsis ''Coenyropsis'' is a genus of butterflies from the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a disti ...

'' van Son, 1958 :* ''

Mashuna ''Mashuna'' is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae. Species *''Mashuna mashuna ''Mashuna mashuna'', the Mashuna ringlet, is a butterfly in the family Nymphalidae. It is found in Zimbabwe Zimbabw ...

'' van Son, 1955 :* '' Mashunoides'' Mendes & Bivar de Sousa, 2009 :* '' Melampias'' Hübner, 1819 :* '' Neita'' van Son, 1955 :* '' Neocoenyra'' Butler, 1886 :* ''

Pseudonympha ''Pseudonympha'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae.

'' Wallengren, 1857 :* ''

Paternympha ''Pseudonympha'' is a genus of butterflies from the subfamily Satyrinae in the family Nymphalidae.

'' Henning & Henning, 1997 :* ''

Physcaeneura ''Physcaeneura'' is a butterfly genus from the subfamily Satyrinae The Satyrinae, the satyrines or satyrids, commonly known as the browns, are a subfamily of the Nymphalidae (brush-footed butterflies). They were formerly considered a distinct ...

'' Wallengren, 1857 :* '' Strabena'' Mabille, 1877 :* '' Stygionympha'' van Son, 1955 :* ''

Ypthima ''Ypthima'' is a species rich genus of Old World butterflies in the family Nymphalidae.Austroypthima'' Holloway, 1974 :* '' Ypthimomorpha'' van Son, 1955 ; subtribe uncertain :* '' Calisto'' Hübner, 1823 :* '' Paralasa'' Moore, 1893 :* '' Callerebia'' Butler, 1867 :* '' Proterebia'' Roos & Amschied, 1980 :* ''

Loxerebia ''Loxerebia'' is a butterfly genus of the subfamily Satyrinae. The genus is confined to China (including Tibet), Mongolia, Burma, Thailand and Laos. Most species are endemic to China. Species * '' Loxerebia albipuncta'' (Leech, 1890) * '' Loxere ...

'' Watkins, 1925 :* '' Dyndirus'' Capronnier, 1874

Notes

References

Citations

Sources

* * * * * * * * ** * * **

External links

Satyrinae
at NYMPHALIDAE.net (There is no individual page for Satyrini as it is split into pages for each subtribes.)
Boisduval 1833
at TREE OF LIFE web project

at Pteron World (in Japanese) {{Taxonbar, from=Q2717036 Satyrinae Butterfly tribes