genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Bignoniaceae
Bignoniaceae is a family of flowering plants in the order Lamiales commonly known as the bignonias or trumpetvines.Vernon H. Heywood, Richard K. Brummitt, Ole Seberg, and Alastair Culham. ''Flowering Plant Families of the World''. Firefly Book ...
.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s, known in
Latin America
Latin America or
* french: Amérique Latine, link=no
* ht, Amerik Latin, link=no
* pt, América Latina, link=no, name=a, sometimes referred to as LatAm is a large cultural region in the Americas where Romance languages — languages derived f ...
by the common names poui, pau d'arco, or ipê. The latter sometimes appears as epay or simply ipe (unaccented) in
English
English usually refers to:
* English language
* English people
English may also refer to:
Peoples, culture, and language
* ''English'', an adjective for something of, from, or related to England
** English national ide ...
. The large
timber
Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
species are sometimes called lapacho or guayacan, but these names are more properly applied to the species '' Handroanthus lapacho'' and ''
Handroanthus guayacan
''Handroanthus guayacan'', is a Bignoniaceae tree native to South America and the Mexican states of Campeche, Chiapas, Oaxaca, Quintana Roo, Tabasco, and Veracruz.
References
guayacan
''Guaiacum'' (''OED'' 2nd edition, 1989.name ''Handroanthus'' was established in 1970,''Handroanthus'' in International Plant Names Index. (see ''External links'' below). but was not generally accepted. In 1992, its species were included in ''
Tabebuia
''Tabebuia'' is a genus of flowering plants in the family Bignoniaceae.Eberhard Fischer, Inge Theisen, and Lúcia G. Lohmann. 2004. "Bignoniaceae". pages 9-38. In: Klaus Kubitzki (editor) and Joachim W. Kadereit (volume editor). ''The Families ...
'' in the most recent revision of that genus.Alwyn H. Gentry. 1992. "Bignoniaceae: Part II (Tribe Tecomeae)". ''Flora Neotropica'' Monograph 25(part 2):1-150. ''Handroanthus'' was resurrected in 2007 when a comparison of DNA sequences by
cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
methods showed that ''Tabebuia'', as then
circumscribed
In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius.
Not every polyg ...
, was not monophyletic.Susan O. Grose and Richard G. Olmstead. 2007. "Evolution of a Charismatic Neotropical Clade: Molecular Phylogeny of ''Tabebuia'' s.l., Crescentieae, and Allied Genera (Bignoniaceae)". ''Systematic Botany'' 32(3):650-659.Susan O. Grose and Richard G. Olmstead. 2007. "Taxonomic Revisions in the Polyphyletic Genus ''Tabebuia'' s.l. (Bignoniaceae)". ''Systematic Botany'' 32(3):660-670.
''Handroanthus'' are
indigenous
Indigenous may refer to:
*Indigenous peoples
*Indigenous (ecology), presence in a region as the result of only natural processes, with no human intervention
*Indigenous (band), an American blues-rock band
*Indigenous (horse), a Hong Kong racehorse ...
from
Central America
Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
to northern
Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
,
Paraguay
Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
and
Chile
Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, with one species, ''
Handroanthus billbergii
''Handroanthus'' is a genus of flowering plants in the family Bignoniaceae.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30 species of trees, known in Latin America by ...
'', native to northern
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
and the
Antilles
The Antilles (; gcf, label=Antillean Creole, Antiy; es, Antillas; french: Antilles; nl, Antillen; ht, Antiy; pap, Antias; Jamaican Patois: ''Antiliiz'') is an archipelago bordered by the Caribbean Sea to the south and west, the Gulf of Mex ...
. ''Handroanthus'' are frequently cultivated far from their
natural
Nature, in the broadest sense, is the physical world or universe. "Nature" can refer to the phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. Although humans are ...
range
Range may refer to:
Geography
* Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra)
** Mountain range, a group of mountains bordered by lowlands
* Range, a term used to i ...
, as ornamental trees, for their large and showy flowers. They easily become naturalized where introduced because their seeds are prolifically produced and widely scattered by the wind.George W. Staples and Derral R. Herbst. 2005. "A Tropical Garden Flora" Bishop Museum Press: Honolulu, HI, USA. Several species are important timber trees of the
American
American(s) may refer to:
* American, something of, from, or related to the United States of America, commonly known as the "United States" or "America"
** Americans, citizens and nationals of the United States of America
** American ancestry, pe ...
tropics
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referr ...
.Alwyn H. Gentry. 1992. "A Synopsis of Bignoniaceae Ethnobotany and Economic Botany". ''Annals of the Missouri Botanical Garden'' 79(1):53-64.Medicinal use has been reported, but its efficacy and
side effects
In medicine, a side effect is an effect, whether therapeutic or adverse, that is secondary to the one intended; although the term is predominantly employed to describe adverse effects, it can also apply to beneficial, but unintended, consequence ...
Chamisso
Adelbert von Chamisso (; 30 January 178121 August 1838) was a German poet and botanist, author of ''Peter Schlemihl'', a famous story about a man who sold his shadow. He was commonly known in French as Adelbert de Chamisso (or Chamissot) de Bonc ...
Handroanthus billbergii
''Handroanthus'' is a genus of flowering plants in the family Bignoniaceae.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30 species of trees, known in Latin America by ...
'' (
Bureau
Bureau ( ) may refer to:
Agencies and organizations
* Government agency
*Public administration
* News bureau, an office for gathering or distributing news, generally for a given geographical location
* Bureau (European Parliament), the administra ...
Handroanthus chrysanthus
''Handroanthus chrysanthus'' (araguaney or yellow ipê), formerly classified as ''Tabebuia chrysantha'', also known as ''araguaney'' in Venezuela, as ''guayacán'' in Colombia, Panama, and Ecuador, as ''tajibo'' in Bolivia, and as ''ipê-amarelo ...
Handroanthus guayacan
''Handroanthus guayacan'', is a Bignoniaceae tree native to South America and the Mexican states of Campeche, Chiapas, Oaxaca, Quintana Roo, Tabasco, and Veracruz.
References
guayacan
''Guaiacum'' (''OED'' 2nd edition, 1989.Seemann) S. Grose
* ''
Handroanthus heptaphyllus
''Handroanthus heptaphyllus'', commonly referred to as the pink trumpet tree or pink tab, is a Bignoniaceae tree native to tropical and subtropical regions of South America. It grows in the high forest watershed of the Paraná River, Paraguay Ri ...
Handroanthus impetiginosus
''Handroanthus impetiginosus'', the pink ipê, pink lapacho or pink trumpet tree, is a tree in the family Bignoniaceae, distributed throughout North, Central and South America, from northern Mexico south to northern Argentina. It is the national ...
Handroanthus serratifolius
''Handroanthus serratifolius'' is a species of tree, commonly known as yellow lapacho, pau d'arco, yellow poui, yellow ipe, pau d'arco amarelo, or ''ipê-amarelo''.
Description
It is a tree native to forests throughout Central and South Ame ...
'' (
Vahl Vahl is a surname. Notable people with the surname include:
* Christian-Friedrich Vahl (born 1955), German cardiac surgeon
* Emanuel Vahl (born 1938), Ukrainian-Israeli composer
*Herbert-Ernst Vahl, German SS general
*Jens Vahl (1796–1854), Danis ...
Sprague
Sprague may refer to:
Places
;Canada
* Sprague, Manitoba, a small town near the Minnesota/Manitoba border
;United States
* Sprague, Alabama, Montgomery County, Alabama
* Sprague, Connecticut
* Sprague, Missouri
* Sprague, Nebraska
* Sprague ...
Sonder
Sonder may refer to: Geography
*Mount Sonder, mountain in Australia
*Sonder, village in Minahasa Regency, Indonesia
People
*Otto Wilhelm Sonder (1812–1881), German botanist and pharmacist
Other uses
* Sonder (Dermot Kennedy album), ''Sonder'' ( ...
) J.R. Mattos
* ''
Handroanthus vellosoi
''Handroanthus'' is a genus of flowering plants in the family Bignoniaceae.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30 species of trees, known in Latin America by ...
paper
Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distrib ...
that resurrected ''Handroanthus'' in 2007.
*
Tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
Wood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
is very dense.
Heartwood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin t ...
lapachol
Lapachol is a natural phenolic compound isolated from the bark of the lapacho tree.Record, Samuel J.. "Lapachol" pages 17-19. In: ''Tropical Woods'' (1925). This tree is known botanically as ''Handroanthus impetiginosus'', but was formerly known ...
foliate
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
trichome
Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a p ...
scales
Scale or scales may refer to:
Mathematics
* Scale (descriptive set theory), an object defined on a set of points
* Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original
* Scale factor, a number w ...
.
*
Inflorescence
An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
with
dichotomous branching
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
coriaceous
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
,
campanulate
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
, 5 dentate,
pubescent The adjective pubescent may describe:
* people or animals undergoing puberty
* plants that are hairy, covered in trichomes
* insects that are covered in setae
In biology, setae (singular seta ; from the Latin word for "bristle") are any of a ...
yellow
Yellow is the color between green and orange on the spectrum of light. It is evoked by light with a dominant wavelength of roughly 575585 nm. It is a primary color in subtractive color systems, used in painting or color printing. In the ...
or
magenta
Magenta () is a color that is variously defined as pinkish- purplish- red, reddish-purplish-pink or mauvish-crimson. On color wheels of the RGB (additive) and CMY (subtractive) color models, it is located exactly midway between red and blu ...
Theca
In biology, a theca (plural thecae) is a sheath or a covering.
Botany
In botany, the theca is related to plant's flower anatomy. The theca of an angiosperm consists of a pair of microsporangia that are adjacent to each other and share a comm ...
e
divaricate
Divaricate means branching, or having separation or a degree of separation. The angle between branches is wide.
In botany
In botany, the term is often used to describe the branching pattern of plants. Plants are said to be divaricating when the ...
linear
Linearity is the property of a mathematical relationship ('' function'') that can be graphically represented as a straight line. Linearity is closely related to '' proportionality''. Examples in physics include rectilinear motion, the linear ...
Ovule
In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the '' integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the ...
Fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering.
Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
Seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s thin with 2
wings
A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expre ...
. Wings thin and sharply demarcated from seed body.
''Handroanthus'' is distinguished from ''Tabebuia'' by several morphological
characters
Character or Characters may refer to:
Arts, entertainment, and media Literature
* ''Character'' (novel), a 1936 Dutch novel by Ferdinand Bordewijk
* ''Characters'' (Theophrastus), a classical Greek set of character sketches attributed to The ...
. The wood is among the hardest and heaviest known. The heartwood is distinct from the sapwood and contains large quantities of lapachol. ''Handroanthus'' has the same lepidote scales as ''Tabebuia'', but also has various types of hair. The calyx is 5-dentate and campanulate to cupular. The corolla is yellow, except in those four species where it is magenta with a yellow throat. ''Tabebuia'' has only two yellow-flowered species, ''
Tabebuia aurea
''Tabebuia aurea'' is a species of ''Tabebuia'' native to South America in Suriname, Brazil, eastern Bolivia, Peru, Paraguay, and northern Argentina. The common English name Caribbean trumpet tree is misleading, as it is not native to the Cari ...
'' and '' Tabebuia nodosa''. The fruit of ''Handroanthus'' is rarely
glabrous
Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
like that of ''Tabebuia''. It usually ranges from sparsely pubescent to densely
tomentose
Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a pl ...
.
Uses
''Handroanthus'' is widely used as ornamental tree in the tropics in
landscaping
Landscaping refers to any activity that modifies the visible features of an area of land, including the following:
# Living elements, such as flora or fauna; or what is commonly called gardening, the art and craft of growing plants with a goal ...
gardens, public squares, and boulevards due to its impressive and colorful flowering. Many flowers appear on still leafless stems at the end of the
dry season
The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The te ...
Handroanthus impetiginosus
''Handroanthus impetiginosus'', the pink ipê, pink lapacho or pink trumpet tree, is a tree in the family Bignoniaceae, distributed throughout North, Central and South America, from northern Mexico south to northern Argentina. It is the national ...
tropics
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referr ...
. ''
Handroanthus chrysanthus
''Handroanthus chrysanthus'' (araguaney or yellow ipê), formerly classified as ''Tabebuia chrysantha'', also known as ''araguaney'' in Venezuela, as ''guayacán'' in Colombia, Panama, and Ecuador, as ''tajibo'' in Bolivia, and as ''ipê-amarelo ...
'', ''
Handroanthus guayacan
''Handroanthus guayacan'', is a Bignoniaceae tree native to South America and the Mexican states of Campeche, Chiapas, Oaxaca, Quintana Roo, Tabasco, and Veracruz.
References
guayacan
''Guaiacum'' (''OED'' 2nd edition, 1989.Handroanthus serratifolius
''Handroanthus serratifolius'' is a species of tree, commonly known as yellow lapacho, pau d'arco, yellow poui, yellow ipe, pau d'arco amarelo, or ''ipê-amarelo''.
Description
It is a tree native to forests throughout Central and South Ame ...
Handroanthus vellosoi
''Handroanthus'' is a genus of flowering plants in the family Bignoniaceae.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30 species of trees, known in Latin America by ...
Anthony Huxley
Anthony Julian Huxley (2 December 1920 – 26 December 1992) was a British botanist. He edited ''Amateur Gardening'' from 1967 to 1971, and was vice-president of the Royal Horticultural Society in 1991.
He was the son of Julian Huxley. He was ...
, Mark Griffiths, and Margot Levy (1992). ''The New Royal Horticultural Society Dictionary of Gardening''. The Macmillan Press, Limited: London. The Stockton Press: New York. (set).
''
Handroanthus heptaphyllus
''Handroanthus heptaphyllus'', commonly referred to as the pink trumpet tree or pink tab, is a Bignoniaceae tree native to tropical and subtropical regions of South America. It grows in the high forest watershed of the Paraná River, Paraguay Ri ...
'', ''Handroanthus serratifolius'', ''Handroanthus guayacan'', ''Handroanthus chrysanthus'', and ''
Handroanthus billbergii
''Handroanthus'' is a genus of flowering plants in the family Bignoniaceae.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30 species of trees, known in Latin America by ...
'' are important timber trees of the
Neotropics
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone.
Definition
In bioge ...
. The wood of ''Handroanthus billbergii'' is valued for
carving
Carving is the act of using tools to shape something from a material by scraping away portions of that material. The technique can be applied to any material that is solid enough to hold a form even when pieces have been removed from it, and ...
. Indigenous peoples of the Amazon made hunting bows from the wood, which is the source of the common name ''pau d'arco'', "bow stick".
Much of the
lumber
Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, wi ...
from ''Handroanthus'' is
export
An export in international trade is a good produced in one country that is sold into another country or a service provided in one country for a national or resident of another country. The seller of such goods or the service provider is an ...
ed. The wood is durable outdoors, where it is usually used for furniture and decking. It is increasingly popular as a decking material due to its insect resistance and durability. ''Handroanthus'' and the unrelated ''
Guaiacum
''Guaiacum'' (''OED'' 2nd edition, 1989.Entry "guaiacum" in Zygophyllaceae
Zygophyllaceae is a family of flowering plants that contains the bean-caper and caltrop. The family includes around 285 species in 22 genera.
Plants in the family Zygophyllaceae may be trees, shrubs, or herbs. They are often found in dry habit ...
) produce the hardest, heaviest, and most durable wood of the American tropics. Dead trees of ''Handroanthus guayacan'' remain standing after they were killed by flooding of their
habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
during construction of the
Panama Canal
The Panama Canal ( es, Canal de Panamá, link=no) is an artificial waterway in Panama that connects the Atlantic Ocean with the Pacific Ocean and divides North and South America. The canal cuts across the Isthmus of Panama and is a condui ...
.
The wood of ''Handroanthus'' brings a high
price
A price is the (usually not negative) quantity of payment or compensation given by one party to another in return for goods or services. In some situations, the price of production has a different name. If the product is a "good" in the ...
. The wood of other species is sometimes fraudulently sold as ''Handroanthus''. By 2007, FSC-certified ipê wood had become readily available on the market, although certificates are occasionally forged.
Much of the ipê imported into the United States is used for decking. Starting in the late 1960s, importing companies targeted large
boardwalk
A boardwalk (alternatively board walk, boarded path, or promenade) is an elevated footpath, walkway, or causeway built with wooden planks that enables pedestrians to cross wet, fragile, or marshy land. They are also in effect a low type of brid ...
projects to sell ipê, beginning with the
New York City Department of Parks and Recreation
The New York City Department of Parks and Recreation, also called the Parks Department or NYC Parks, is the department of the government of New York City responsible for maintaining the city's parks system, preserving and maintaining the ecolog ...
, which maintains the city's
boardwalk
A boardwalk (alternatively board walk, boarded path, or promenade) is an elevated footpath, walkway, or causeway built with wooden planks that enables pedestrians to cross wet, fragile, or marshy land. They are also in effect a low type of brid ...
s, including along the
beach
A beach is a landform alongside a body of water which consists of loose particles. The particles composing a beach are typically made from rock, such as sand, gravel, shingle, pebbles, etc., or biological sources, such as mollusc sh ...
of Coney Island. The city began using ipê around that time and has since converted the entire boardwalk—over long—to ipê. The ipê lasted about 25 years, at which time (1994) the department began replacing it with new ipê. ''
La Sultana
''La Sultana'' is a yacht that has been refitted from a Soviet era ferry. She was launched in 1962, under the name ''Aji-Petri'' and was the fifth in a series of 12 ships that were originally built for the Russian fleet. Her purpose was to trans ...
'', a yacht refashioned from a Soviet spy vessel, was fitted with an ipê deck during its restoration.La Sultana Superyacht: The Spy Ship You Can Sunbathe On Billionaire. 14 September 2015. Tara Loader Wilkinson. 24 December 2015. In 2008-2009,
Wildwood, New Jersey
Wildwood is a city in Cape May County in the U.S. state of New Jersey. It is part of the Ocean City metropolitan statistical area and is a popular summer resort destination along the Jersey Shore. As of the 2020 U.S. census, the city's year-ro ...
, rebuilt a section of its boardwalk using ipê. The town had pledged to use domestic
black locust
''Robinia pseudoacacia'', commonly known in its native territory as black locust, is a medium-sized hardwood deciduous tree, belonging to the tribe Robinieae of the legume family Fabaceae. It is endemic to a few small areas of the United States ...
, but it was not available in time.
Given that ipê trees typically grow in
densities
Density (volumetric mass density or specific mass) is the substance's mass per unit of volume. The symbol most often used for density is ''ρ'' (the lower case Greek language, Greek letter Rho (letter), rho), although the Latin letter ''D'' ca ...
of only one or two trees per , large areas of
forest
A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
must be searched and cut down to create paths to harvest the trees to fill orders for boardwalks and to a lesser extent, homeowner decks.
Medical claims for ''Handroanthus'' are numerous, but so far, have always originated from unreliable sources. For this reason, the extensive literature on this subject is not reviewed here.
The bark of several species of ''Handroanthus'' is sold in South American markets. Similar-looking bark is often fraudulently passed off as ''Handroanthus''. It is used in various ways to relieve certain symptoms of certain
cancer
Cancer is a group of diseases involving abnormal cell growth with the potential to invade or spread to other parts of the body. These contrast with benign tumors, which do not spread. Possible signs and symptoms include a lump, abnormal b ...
s. No evidence shows that it prevents the disease or slows its progression, as is often claimed.
The bark is dried, shredded, and then boiled to make a bitter or sour-tasting brownish-colored tea. Tea from the inner bark of pink ipê (''
Handroanthus impetiginosus
''Handroanthus impetiginosus'', the pink ipê, pink lapacho or pink trumpet tree, is a tree in the family Bignoniaceae, distributed throughout North, Central and South America, from northern Mexico south to northern Argentina. It is the national ...
'') is known as pau d'arco,
lapacho
Lapacho or taheebo is herbal tea made from the inner bark of the pau d'arco tree ''Handroanthus impetiginosus''.
Lapacho is used in the herbal medicine of several South and Central American indigenous peoples to treat a number of ailments inclu ...
, or taheebo. Its main
active ingredient
An active ingredient is any ingredient that provides biologically active or other direct effect in the diagnosis, cure, mitigation, treatment, or prevention of disease or to affect the structure or any function of the body of humans or animals. The ...
s are
lapachol
Lapachol is a natural phenolic compound isolated from the bark of the lapacho tree.Record, Samuel J.. "Lapachol" pages 17-19. In: ''Tropical Woods'' (1925). This tree is known botanically as ''Handroanthus impetiginosus'', but was formerly known ...
,
quercetin
Quercetin is a plant flavonol from the flavonoid group of polyphenols. It is found in many fruits, vegetables, leaves, seeds, and grains; capers, red onions, and kale are common foods containing appreciable amounts of it. It has a bitter flavor ...
, and other
flavonoids
Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans.
Chemically, flavonoids ...
. A
dehydrated
In physiology, dehydration is a lack of total body water, with an accompanying disruption of metabolic processes. It occurs when free water loss exceeds free water intake, usually due to exercise, disease, or high environmental temperature. Mil ...
tea is also available in
pill
Pill or The Pill may refer to:
Drugs
* Pill (pharmacy), referring to anything small for a specific dose of medicine
* "The Pill", a general nickname for the combined oral contraceptive pill
Film and television
* ''The Pill'' (film), a 2011 fil ...
form. Taheebo has been used for years in
Central America
Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
disease
A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that a ...
s including
Eczema
Dermatitis is inflammation of the skin, typically characterized by itchiness, redness and a rash. In cases of short duration, there may be small blisters, while in long-term cases the skin may become thickened. The area of skin involved can ...
,
Candidiasis
Candidiasis is a fungal infection due to any type of '' Candida'' (a type of yeast). When it affects the mouth, in some countries it is commonly called thrush. Signs and symptoms include white patches on the tongue or other areas of the mouth ...
cancer
Cancer is a group of diseases involving abnormal cell growth with the potential to invade or spread to other parts of the body. These contrast with benign tumors, which do not spread. Possible signs and symptoms include a lump, abnormal b ...
. The worth and use of Taheebo extract has been related to the importance of
quinine
Quinine is a medication used to treat malaria and babesiosis. This includes the treatment of malaria due to '' Plasmodium falciparum'' that is resistant to chloroquine when artesunate is not available. While sometimes used for nocturnal le ...
, which is taken from the bark of the South American Cinchona tree and is a medicinally accepted treatment for
malaria
Malaria is a mosquito-borne infectious disease that affects humans and other animals. Malaria causes symptoms that typically include fever, tiredness, vomiting, and headaches. In severe cases, it can cause jaundice, seizures, coma, or death. S ...
. The
herbal remedy
Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remedies ...
is typically used during
flu
Influenza, commonly known as "the flu", is an infectious disease caused by influenza viruses. Symptoms range from mild to severe and often include fever, runny nose, sore throat, muscle pain, headache, coughing, and fatigue. These symptom ...
and
cold
Cold is the presence of low temperature, especially in the atmosphere. In common usage, cold is often a subjective perception. A lower bound to temperature is absolute zero, defined as 0.00K on the Kelvin scale, an absolute thermodynamic ...
season
A season is a division of the year based on changes in weather, ecology, and the number of daylight hours in a given region. On Earth, seasons are the result of the axial parallelism of Earth's tilted orbit around the Sun. In temperate and ...
and for easing smoker's cough. It apparently works as an
expectorant
Mucoactive agents are a class of chemical agents that aid in the clearance of mucus or sputum from the upper and lower airways, including the lungs, bronchi, and trachea. Mucoactive drugs include expectorants, mucolytics, mucoregulators, and muco ...
, by promoting the lungs to cough up and free deeply embedded
mucus
Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It ...
and contaminants. However, lapachol is rather
toxic
Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a subs ...
and therefore a more
topical
A topical medication is a medication that is applied to a particular place on or in the body. Most often topical medication means application to body surfaces such as the skin or mucous membranes to treat ailments via a large range of classes ...
Handroanthus serratifolius
''Handroanthus serratifolius'' is a species of tree, commonly known as yellow lapacho, pau d'arco, yellow poui, yellow ipe, pau d'arco amarelo, or ''ipê-amarelo''.
Description
It is a tree native to forests throughout Central and South Ame ...
additive
Additive may refer to:
Mathematics
* Additive function, a function in number theory
* Additive map, a function that preserves the addition operation
* Additive set-functionn see Sigma additivity
* Additive category, a preadditive category with f ...
to the
entheogen
Entheogens are psychoactive substances that induce alterations in perception, mood, consciousness, cognition, or behavior for the purposes of engendering spiritual development or otherwiseRätsch, Christian, ''The Encyclopedia of Psychoac ...
ic drink
Ayahuasca
AyahuascaPronounced as in the UK and in the US. Also occasionally known in English as ''ayaguasca'' ( Spanish-derived), ''aioasca'' (Brazilian Portuguese-derived), or as ''yagé'', pronounced or . Etymologically, all forms but ''yagé'' desce ...
plant
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...
sac fungus
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The de ...
was first discovered on a ''Handroanthus'' tree, known at that time as ''Tabebuia''. The
taxonomy
Taxonomy is the practice and science of categorization or classification.
A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of
Mycosphaerella
''Mycosphaerella'' is a genus of ascomycota. With more than 10,000 species, it is the largest genus of plant pathogen fungi.
The following introduction about the fungal genus ''Mycosphaerella'' is copied (with permission) from the dissertation o ...
is in much confusion and the recognition of this name is questionable.
Symbolism
Because of its popularity, ''Handroanthus'' has often been adopted as a symbol or
emblem
An emblem is an abstract or representational pictorial image that represents a concept, like a moral truth, or an allegory, or a person, like a king or saint.
Emblems vs. symbols
Although the words ''emblem'' and '' symbol'' are often us ...
for
nations
A nation is a community of people formed on the basis of a combination of shared features such as language, history, ethnicity, culture and/or society. A nation is thus the collective identity of a group of people understood as defined by t ...
or other
political division
Administrative division, administrative unit,Article 3(1). country subdivision, administrative region, subnational entity, constituent state, as well as many similar terms, are generic names for geographical areas into which a particular, ind ...
national tree
This is a list of national trees, most official, but some unofficial.
National trees
See also
* National emblem
* Floral emblem
* List of U.S. State and territory trees
References
{{DEFAULTSORT:List Of National Trees
N
Trees
In ...
is sometimes not entirely clear. Gentry (1992) gives the following information without making that distinction.
*
Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
- ''Handroanthus serratifolius''
*
Ecuador
Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
- ''Handroanthus chrysanthus''
*
Paraguay
Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
- ''Handroanthus heptaphyllus''
*
Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
- ''Handroanthus billbergii''
History
The genus ''Handroanthus'' was erected by Joáo Rodrigues de Mattos in 1970.Joáo Rodrigues de Mattos. 1970. "''Handroanthus'', Um novo gênero para os "ipês" do Brasil". ''Loefgrenia'' 50: 1-4. It was named for the Brazilianbotanist
Oswaldo Handro
file:Oswaldo Handro.jpg, Oswaldo Handro.
Oswaldo Handro (1908–1986) was a Brazilian botanist who specialized in pteridophytes and spermatophytes. The genus ''Handroanthus'' was Genus#Generic name, named for him.
References
1908 bi ...
. "Anthus" is
derived
Derive may refer to:
* Derive (computer algebra system), a commercial system made by Texas Instruments
* ''Dérive'' (magazine), an Austrian science magazine on urbanism
*Dérive, a psychogeographical concept
See also
*
*Derivation (disambiguatio ...
from a
Greek
Greek may refer to:
Greece
Anything of, from, or related to Greece, a country in Southern Europe:
*Greeks, an ethnic group.
*Greek language, a branch of the Indo-European language family.
**Proto-Greek language, the assumed last common ancestor ...
word for "flower".
Most botanists at that time did not agree with the separation of ''Handroanthus'' from ''Tabebuia''. Alwyn H. Gentry objected strenuously and warned against "succumbing to further paroxysms of unwarranted splitting".Alwyn H. Gentry. 1972. "''Handroanthus'' (Bignoniaceae): A critique". ''Taxon'' 21(1):113-114.
In 1992, Gentry published a full taxonomictreatment of ''Tabebuia'', in which he described 99 species and one
hybrid
Hybrid may refer to:
Science
* Hybrid (biology), an offspring resulting from cross-breeding
** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species
** Hybridity, the property of a hybrid plant which is a union of two dif ...
for the genus. These consist of the 67 species and one hybrid that remain in ''Tabebuia'', the two species transferred to '' Roseodendron'', and the 30 species that are now placed in ''Handroanthus''. Gentry divided ''Tabebuia'' into 10 species groups. ''Handroanthus'', as it is currently
circumscribed
In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius.
Not every polyg ...
, is composed of Gentry's groups 3, 4, and 5. Gentry believed group 5 to be
natural
Nature, in the broadest sense, is the physical world or universe. "Nature" can refer to the phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. Although humans are ...
sister
A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a family, familial relationship, it is sometimes used endearingly to r ...
to either of the others. Thus ''Tabebuia'' was shown to be
polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...
. One of these clades consisted of the two species that constitute the genus '' Roseodendron''. Another contained the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
for ''Tabebuia'', and consequently retained that name. The name ''Handroanthus'' was resurrected for the third clade, which contained its type species, '' Handroanthus albus''.
''Handroanthus'' is sister to a clade consisting of '' Spirotecoma'', '' Parmentiera'', ''
Crescentia
''Crescentia'' (calabash tree, huingo, krabasi, or kalebas) is a genus of six species of flowering plants in the family Bignoniaceae, native to southern North America, the Caribbean, Central America northern South America. The species are mode ...
'', and ''
Amphitecna
''Amphitecna'' is a genus of plants in the family Bignoniaceae.
Species include:
*'' Amphitecna apiculata'' A.H.Gentry
*'' Amphitecna breedlovei'' A.H.Gentry
*'' Amphitecna costata'' A.H.Gentry
*'' Amphitecna donnell-smithii'' (Sprague) L.O.Will ...
''. It had for a long time been placed in the
tribe
The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confli ...
Tecomeae
Tecomeae is a tribe with 44 genera of trees, shrubs, and vines in the family Bignoniaceae.
Genera
* '' Argylia''
* '' Astianthus''
* '' Campsidium''
* '' Campsis''
* '' Catalpa''
* '' Catophractes''
* '' Chilopsis''
* '' Cybistax''
* '' Del ...
, but that tribe has been greatly reduced to only 11 or 12 genera and no longer includes ''Handroanthus''. ''Handroanthus'' is one of the 12 to 14 genera that make up a
group
A group is a number of persons or things that are located, gathered, or classed together.
Groups of people
* Cultural group, a group whose members share the same cultural identity
* Ethnic group, a group whose members share the same ethnic ide ...
informally known as the ''Tabebuia'' alliance.Richard G. Olmstead, Michelle L. Zjhra, Lúcia G. Lohmann, Susan O. Grose, and Andrew J. Eckert. 2009. "A molecular phylogeny and classification of Bignoniaceae". ''American Journal of Botany'' 96(9): 1731–1743. This group has not been assigned to any
taxonomic rank
In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family, order, class, phylum, kingdom, domain. While ...
, and neither has Crescentiina, the smallest group that it is a member of.
Cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analysis of DNA data has strongly supported ''Handroanthus'', but sampling of taxa and DNA has not been sufficient to strongly support any relationships within the genus.