Fossil Flora Of Kızılcahamam District
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fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
flora in
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
stems from at least six
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Güvem and Beşkonak villages, north of
Ankara Ankara is the capital city of Turkey and List of national capitals by area, the largest capital by area in the world. Located in the Central Anatolia Region, central part of Anatolia, the city has a population of 5,290,822 in its urban center ( ...
, north of
Kızılcahamam Kızılcahamam is a municipality and Districts of Turkey, district of Ankara Province, Turkey. Its area is 1,623 km2, and its population is 26,872 (2022). It is 70 km north of the city of Ankara, near the motorway to Istanbul. Its elevati ...
and south of the
Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
coast. They are to above sea level, extending north-south and east-west. Between six and seven million years ago, a fresh water lake existed there in a forested area with mostly broad leaved
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
tree species, dominated by sequoia and oak. The Turkish
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
Professor Baki Kasaplıgil (1918-1992) from
Çankırı Çankırı, historically known as Gangra (Greek language, Greek: Γάγγρα), is a city in Turkey, about northeast of Ankara. It is situated about 800 m (2500 ft) above sea level. It is the seat of Çankırı Province and of Çankır ...
who worked at the Biology Department,
Mills College Mills College at Northeastern University in Oakland, California is part of Northeastern University's global university system. Mills College was founded as the Young Ladies Seminary in 1852 in Benicia, California; it was relocated to Oakland in ...
,
Oakland Oakland is a city in the East Bay region of the San Francisco Bay Area in the U.S. state of California. It is the county seat and most populous city in Alameda County, with a population of 440,646 in 2020. A major West Coast port, Oakland is ...
,
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
from 1956 to 1992, made
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
collecting trips to the area around
Kızılcahamam Kızılcahamam is a municipality and Districts of Turkey, district of Ankara Province, Turkey. Its area is 1,623 km2, and its population is 26,872 (2022). It is 70 km north of the city of Ankara, near the motorway to Istanbul. Its elevati ...
-
Çerkeş Çerkeş is a town in Çankırı Province in the Central Anatolia region of Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Sou ...
road in 1964 and 1968. In the 1964 trip he collected fossil leaves of ''
Glyptostrobus ''Glyptostrobus'' is a small genus of conifers in the family Cupressaceae (formerly in the family Taxodiaceae). The sole living species, '' Glyptostrobus pensilis'', is native to subtropical southeastern China, from Fujian west to southeast Yu ...
'', '' Sequoia'', ''
Zelkova ''Zelkova'' (from the Georgian language, Georgian ''dzelkva'', 'stone pillar') is a genus of six species of deciduous trees in the elm family Ulmaceae, native to southern Europe, and southwest and eastern Asia. They vary in size from shrubs (''Ze ...
'', ''
Fagus Beech (genus ''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to subtropical (accessory forest element) and temperate (as dominant element of Mesophyte, mesophytic forests) Eurasia and North America. There are 14 accepted ...
'', ''
Carpinus Hornbeams are hardwood trees in the plant genus ''Carpinus'' in the family Betulaceae. Its species occur across much of the temperate regions of the Northern Hemisphere. Common names The common English name ''hornbeam'' derives from the hard ...
'', ''
Ulmus Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...
'', ''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
'', ''
Tilia ''Tilia'' is a genus of about 30 species of trees or bushes, native throughout most of the temperateness, temperate Northern Hemisphere. The tree is known as linden for the European species, and basswood for North American species. In Great Bri ...
'' and '' Acer''. In the 1968 trip, he collected 400 plant parts such as leaves, seeds, fruits, twigs and branches. He also collected 50 fossils of frog and fish skeletons and larvae and insects. In 1969, Wayne Fry from
University of Berkeley The University of California, Berkeley (UC Berkeley, Berkeley, Cal, or California), is a public land-grant research university in Berkeley, California, United States. Founded in 1868 and named after the Anglo-Irish philosopher George Berkeley ...
made plant fossil collections in Güvem. Daniel I. Axelrod also assisted in the research of this flora.{{Cite web, url=http://docplayer.biz.tr/6274084-Ankara-kizilcahamam-yakinindaki-guvem-koyu-civarinda-bulunan-son-tersiyer-kozalakli-yesil-yaprakli-ormani.html, title = ANKARA, KIZILCAHAMAM YAKININDAKİ GÜVEM KÖYÜ CİVARINDA BULUNAN SON TERSİYER KOZALAKLI-YEŞİL YAPRAKLI ORMANI* - PDF Free Download


Images of the fossil plants and fossil sites

Image:Leaf fossils of Glyptostrobus europaeus and Quercus drymeya from the Pliocene of Kızılcahamam, Ankara Province, Turkey.jpg, Leaf fossils of Glyptostrobus europaeus (left) and Quercus drymeya (right) from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Kizilcahamam, found near Belpınar village by Osman Güldemir, a teacher at the
Anadolu University Anadolu University () is a public university in Eskişehir, Turkey. Anadolu University Distance Education Faculty (or Anadolu University Global Campus) serves as the national provider of distance education and is a member of the European Associ ...
in
Eskişehir Eskişehir ( , ; from 'old' and 'city') is a city in northwestern Turkey and the capital of the Eskişehir Province. The urban population of the city is 821 315 (Odunpazari + Tebebasi), with a metropolitan population of 921 630. The city is l ...
,
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
. Image:Mahalle.jpg, Güvem village with the Seyhamamı
mosque A mosque ( ), also called a masjid ( ), is a place of worship for Muslims. The term usually refers to a covered building, but can be any place where Salah, Islamic prayers are performed; such as an outdoor courtyard. Originally, mosques were si ...
. Image:Basalt Columns.JPG,
Columnar basalt Basalt (; ) is an aphanitic (fine-grained) extrusive igneous rock formed from the rapid cooling of low-viscosity lava rich in magnesium and iron (mafic lava) exposed at or very near the surface of a rocky planet or moon. More than 90% of a ...
formed by
lava Lava is molten or partially molten rock (magma) that has been expelled from the interior of a terrestrial planet (such as Earth) or a Natural satellite, moon onto its surface. Lava may be erupted at a volcano or through a Fissure vent, fractu ...
flows at Beşkonak village. Image:06890 Kızılcahamam-Ankara, Turkey - panoramio (11).jpg, Due to drying and cooling of the climate at the onset of the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages, and g ...
in Kizilcahamam, the Sequoia forests with their many exotic plant species disappeared and have now been replaced by
Scots pine ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US), Baltic pine, or European red pine is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-gr ...
(
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US), Baltic pine, or European red pine is a species of tree in the pine family Pinaceae that is native plant, native to Eurasia. It can readily be identified by its combination of fairly s ...
) forests. Image:06890 Ortaköy-Kızılcahamam-Ankara, Turkey - panoramio.jpg, Professor Baki Kasaplıgil collected many of his
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
specimens along
dirt roads A dirt road or track is a type of unpaved road not paved with asphalt, concrete, brick, or stone; made from the native material of the land surface through which it passes, known to highway engineers as subgrade material. Terminology Simi ...
that had newly been made by a
bulldozer A bulldozer or dozer (also called a crawler) is a large tractor equipped with a metal #Blade, blade at the front for pushing material (soil, sand, snow, rubble, or rock) during construction work. It travels most commonly on continuous tracks, ...
.


Fossil plant species so far described from the deposits


Pteridophyta A pteridophyte is a vascular plant (with xylem and phloem) that reproduces by means of spores. Because pteridophytes produce neither flowers nor seeds, they are sometimes referred to as " cryptogams", meaning that their means of reproduction is h ...

* Salviniaceae :*''
Salvinia ''Salvinia'' or watermosses is a genus of free-floating aquatic ferns in the family Salviniaceae. The genus is named in honor of 17th-century Italian naturalist Anton Maria Salvini, and the generic name was first published in 1754 by French bo ...
'' sp. *
Pteridaceae Pteridaceae is a family of ferns in the order Polypodiales, including some 1150 known species in ca 45 genera (depending on taxonomic opinions), divided over five subfamilies. The family includes four groups of genera that are sometimes recogni ...
:*'' Cryptogramma aff. crispo''


Gymnosperms The gymnosperms ( ; ) are a group of woody, perennial Seed plant, seed-producing plants, typically lacking the protective outer covering which surrounds the seeds in flowering plants, that include Pinophyta, conifers, cycads, Ginkgo, and gnetoph ...

*
Cupressaceae Cupressaceae or the cypress family is a family of conifers. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecio ...
:*†'' Glyptostrobus europaeus'' :*†'' Sequoia langsdorfi'' *
Pinaceae The Pinaceae (), or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, Pinyon_pine, piñons, larches, pines and spruces. The family is incl ...
:*'' Pinus canariensis'' :*''
Pinus morrisonicola ''Pinus morrisonicola'' (Taiwan white pine; ), is a species of conifer in the family Pinaceae. It is a large tree, up to high and in diameter. The trunk is often crooked. Needles are in bundles of five. Mature cones are large, to long and wid ...
'' :*''
Tsuga ''Tsuga'' (, from Japanese (), the name of '' Tsuga sieboldii'') is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The English-language common name "hemlock" arose from a perceived similarity in the smell of it ...
'' sp. *
Ephedraceae Ephedraceae is a family of gymnosperms belonging to Gnetophyta, it contains only a single extant genus, ''Ephedra (plant), Ephedra'', as well as a number of extinct genera from the Early Cretaceous. Taxonomy File:Eamesia chinensis.png, ''Eame ...
:*'' Ephedra'' sp.


Angiosperms Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. T ...
, dicots

* Cercidiphyllaceae :*†''
Cercidiphyllum ''Cercidiphyllum'' is a genus containing two species of plants, both commonly called katsura. They are the sole members of the monotypic family (biology), family Cercidiphyllaceae. The genus is native plant, native to Japan and China and unrelate ...
crenatum'' *
Magnoliaceae The Magnoliaceae () are a flowering plant family, the magnolia family, in the order Magnoliales. It consists of two genera: '' Magnolia'' and ''Liriodendron'' (tulip trees). Unlike most angiosperms, whose flower parts are in whorls (rings), ...
:*'' Magnolia sprengeri'' *
Menispermaceae Menispermaceae (botanical Latin: 'moonseed family' from Greek ''mene'' 'crescent moon' and ''sperma'' 'seed') is a family (biology), family of flowering plants. The alkaloid tubocurarine, a neuromuscular blocker and the active ingredient in the ...
:*'' Menispermum'' sp. *
Lauraceae Lauraceae, or the laurels, is a plant Family (biology), family that includes the bay laurel, true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genus (biology), genera worldwide. They are dicotyled ...
:*''
Persea indica ''Persea indica'' is a large, evergreen tree in the laurel family (Lauraceae), native to humid uplands on Madeira and the Canary Islands in the North Atlantic. It belongs to the genus '' Persea'', a group of evergreen trees including the avocado. ...
'' *
Platanaceae Platanaceae, the plane family, is a family (biology), family of flowering plants in the order Proteales. The family consists of only a single extant taxon, extant genus ''Platanus'', with twelve known species. The plants are tall trees, native ...
:*''
Platanus ''Platanus'' ( ) is a genus consisting of a small number of tree species native to the Northern Hemisphere. They are the sole living members of the family Platanaceae. All mature members of ''Platanus'' are tall, reaching in height. The type ...
'' sp. *
Juglandaceae The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have ...
:*†''
Platycarya ''Platycarya'' is a genus of flowering plants in the family Juglandaceae native to eastern Asia in China, Korea, and Japan.Flora of China''Platycarya''/ref> The genus was formerly treated as comprising a single species '' Platycarya strobilacea ...
miocenica'' :*''
Pterocarya fraxinifolia ''Pterocarya fraxinifolia'' is a species of tree in the Juglandaceae family. It is commonly known as the Caucasian wingnut or Caucasian walnut. It is native to the Caucasian region Armenia, Azerbaijan, Georgia, Iran, Russia, Ukraine and Turk ...
'' *
Salicaceae The Salicaceae are the willow family of flowering plants. The traditional family (Salicaceae ''sensu stricto'') includes the willows, poplars. Genetic studies summarized by the Angiosperm Phylogeny Group (APG) have greatly expanded the circumsc ...
:*''
Salix Willows, also called sallows and osiers, of the genus ''Salix'', comprise around 350 species (plus numerous hybrids) of typically deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions. Most species are known ...
'' sp. :*''
Populus tremula ''Populus tremula'' (commonly called aspen, common aspen, Eurasian aspen, European aspen, or quaking aspen) is a species of poplar native to cool temperate regions of the Old World. Description It is a substantial deciduous tree growing to t ...
'' *
Myricaceae Myricaceae is a small family of dicotyledonous shrubs and small trees in the order Fagales with its type genus ''Myrica'', the sweet gales. There are three genera in the family, although some botanists separate many species from Myrica into a fo ...
:*'' Comptonia'' sp. :*''
Myrica ''Myrica'' is a genus of about 35–50 species of small trees and shrubs in the family Myricaceae, order Fagales. The genus has a wide distribution, including Africa, Asia, Europe, North America, and South America, and missing only from Antar ...
banksiaefolia'' *
Fagaceae The Fagaceae (; ) are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with around 1,000 or more species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species ...
:*''
Castanea The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. The name also refers to the edible nuts they produce. They are native to temperate regions of the Northern Hemisphere. Description C ...
'' sp. :*''
Castanopsis ''Castanopsis'', commonly called chinquapin or chinkapin, is a genus of evergreen trees belonging to the beech family, Fagaceae. The genus contains about 140 species, which are today restricted to tropical and subtropical eastern Asia. The Engli ...
'' sp. :*''
Fagus Beech (genus ''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to subtropical (accessory forest element) and temperate (as dominant element of Mesophyte, mesophytic forests) Eurasia and North America. There are 14 accepted ...
'' sp. :*†''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
drymeja'' :*†''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
heidingeri'' :*†''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
kubinyi'' :*†''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
aff. lonchitis'' :*'' Quercus petroela'' aff. :*'' Quercus sclerophyllina'' :*'' Quercus semecarpifolia'' :*†''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
seyfriedii'' (very similar to ''
Quercus phellos ''Quercus phellos'', the willow oak, is a North American species of a deciduous tree in the red oak group of oaks. It is native to the south-central and eastern United States. Description It is a medium-sized tree growing to tall (exceptiona ...
'' from eastern and central
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) :*†''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
sosnoisky'' :*'' Quercus hartwissiana'' :*'' Quercus trojana'' aff. *
Betulaceae Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams, numbering a total of 167 species. They are mostly natives of ...
:*''
Alnus Alders are trees of the genus ''Alnus'' in the birch family Betulaceae. The genus includes about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few species ext ...
'' sp. :*'' Betula luminifera'' aff. :*''
Carpinus Hornbeams are hardwood trees in the plant genus ''Carpinus'' in the family Betulaceae. Its species occur across much of the temperate regions of the Northern Hemisphere. Common names The common English name ''hornbeam'' derives from the hard ...
miocenica'' *
Sapindaceae The Sapindaceae are a family (biology), family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1,858 accepted species. Examples include Aesculus, horse chestnut, maples, ackee and lychee. The ...
:*†'' Acer angustilobum'' :*†'' Acer tribolatum'' *
Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of the Anacardiaceae bear fruits that are drupes and in some cases produce ...
:*''
Astronium ''Astronium'' is a genus of flowering plants in the cashew family, Anacardiaceae. It is native to Central and South America. ''Astronium'' is a genus of dioecious trees. Leaves are deciduous, alternate, and odd-pinnate. Species , Plants of the ...
'' sp. *
Ebenaceae The Ebenaceae are a family of flowering plants belonging to order Ericales. The family includes ebony and persimmon among about 768 species of trees and shrubs. It is distributed across the tropical and warmer temperate regions of the world. It i ...
:*''
Diospyros ''Diospyros'' is a genus of over 700 species of deciduous and evergreen trees and shrubs. The majority are native to the tropics, with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark timb ...
aff. miokaki'' *
Aquifoliaceae ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen or ...
:*''
Ilex ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen or ...
gracilis'' (close to '' Ilex serrata'') *
Rosaceae Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...
:*''
Sorbus aucuparia ''Sorbus aucuparia'', commonly called rowan (, also ) and mountain-ash, is a species of deciduous tree or shrub in the rose family. The tree has a slender trunk with smooth bark, a loose and roundish crown, and its leaves are pinnate in pairs o ...
'' aff. *
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
:*''
Cercis ''Cercis'' is a genus of about 10 species in the subfamily Cercidoideae of the pea family (biology), family Fabaceae. It contains small deciduous trees or large shrubs commonly known as redbuds in the USA. They are characterised by simple, rou ...
'' sp. :*†''
Sophora ''Sophora'' is a genus of about 45 species of small trees and shrubs in the pea family Fabaceae. The species have a pantropical distribution. The generic name is derived from ''sophera'', an Arabic name for a pea-flowered tree. The genus formerl ...
'' sp. *
Malvaceae Malvaceae (), or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include Theobroma cacao, cacao, Cola (plant), cola, cotton, okra, Hibiscus sabdariffa, ...
:*''
Tilia platyphyllos ''Tilia platyphyllos'', the large-leaved lime or large-leaved linden, is a species of flowering plant in the family (biology), family Malvaceae (Tiliaceae). It is a deciduous tree, native plant, native to much of continental Europe as well as sou ...
'' aff, *
Ulmaceae The Ulmaceae () are a family (biology), family of flowering plants that includes the elms (genus ''Ulmus''), and the zelkovas (genus ''Zelkova''). Members of the family are widely distributed throughout the Temperate, north temperate zone, and ha ...
:*''
Ulmus Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...
'' sp. :*†''
Zelkova ''Zelkova'' (from the Georgian language, Georgian ''dzelkva'', 'stone pillar') is a genus of six species of deciduous trees in the elm family Ulmaceae, native to southern Europe, and southwest and eastern Asia. They vary in size from shrubs (''Ze ...
ungeri'' *
Berberidaceae The Berberidaceae are a family (biology), family of 18 genera of flowering plants commonly called the barberry family. This family is in the order (biology), order Ranunculales. The family contains about 700 known species, of which the majority ...
:*'' Berberis chinensis'' aff. *
Moraceae Moraceae is a family of flowering plants comprising about 48 genera and over 1100 species, and is commonly known as the mulberry or fig family. Most are widespread in tropical and subtropical regions, less so in temperate climates; however, their ...
:*''
Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family (biology), family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few spe ...
'' sp. *
Altingiaceae Altingiaceae is a small Family (biology), family of flowering plants in the Order (biology), order Saxifragales,Peter F. Stevens (2001 onwards). "Altingiaceae". At: Angiosperm Phylogeny Website. At: Missouri Botanical Garden Website. (see ''Exter ...
:*†''
Liquidambar ''Liquidambar'', commonly called sweetgum (star gum in the UK), gum, redgum, satin-walnut, styrax or American storax, is the only genus in the flowering plant family Altingiaceae and has 15 species. They were formerly often treated as a part of ...
europaeum'' *
Simaroubaceae The Simaroubaceae, also known as the quassia family, are a small, mostly tropical, family in the order Sapindales. In recent decades, it has been subject to much taxonomic debate, with several small families being split off. A molecular phyloge ...
:*''
Ailanthus altissima ''Ailanthus altissima'' ( ), commonly known as tree of heaven or ailanthus tree, is a deciduous tree in the quassia family. It is native to northeast and central China, and Taiwan. Unlike other members of the genus ''Ailanthus'', it is found ...
'' aff.


Angiosperms Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. T ...
,
monocots Monocotyledons (), commonly referred to as monocots, ( Lilianae '' sensu'' Chase & Reveal) are flowering plants whose seeds contain only one embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but with various ranks a ...

* Smilaceae :*''
Smilax aspera ''Smilax aspera'', with common names common smilax, rough bindweed, sarsaparille,Potamogetonaceae The Potamogetonaceae, commonly referred to as the pondweed family, is an aquatic family of monocotyledonous flowering plants. The roughly 110 known species are divided over five genera. The largest genus in the family by far is '' Potamogeton'', ...
:*''
Potamogeton ''Potamogeton'' is a genus of aquatic, mostly freshwater, plants of the family Potamogetonaceae. Most are known by the common name pondweed, although many unrelated plants may be called pondweed, such as Canadian pondweed (''Elodea canadensis' ...
'' sp. *
Hydrocharitaceae Hydrocharitaceae is a flowering plant family which includes 14 accepted genera and a total of ca 135 known species (Christenhusz & Byng 2016). The family holds a number of species of aquatic plants, including tape-grass, the well-known Canadian ...
:*''
Egeria densa ''Elodea densa'', the large-flowered waterweed or Brazilian waterweed, is a species of '' Elodea'' native to warm temperate South America in southeastern Brazil, Argentina, Chile and Uruguay. It is considered a problematic invasive species due t ...
'' aff. :*'' Najas'' sp. *
Typhaceae The Typhaceae () are a family of flowering plants, sometimes called the cattail family. The botanical name for the family has been recognized by most taxonomists. Description Members can be recognized as large marsh herbs with alternate two-rank ...
:*''
Typha ''Typha'' is a genus of about 30 species of monocotyledonous flowering plants in the family Typhaceae. These plants have a variety of common names, in British English as bulrushStreeter D, Hart-Davies C, Hardcastle A, Cole F, Harper L. 2009. ' ...
'' sp.


Images of some extant close relatives of the fossil species

Image:Redwood slope.jpg, '' Sequoia'', giant redwood forests were common at higher altitudes of
Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
in the
Tertiary Tertiary (from Latin, meaning 'third' or 'of the third degree/order..') may refer to: * Tertiary period, an obsolete geologic period spanning from 66 to 2.6 million years ago * Tertiary (chemistry), a term describing bonding patterns in organic ch ...
and early
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
. Image:Redwoods in Richardson Grove State Park 1.jpg, '' Sequoia'' Image:Sequoia sempervirens BigSur.jpg, '' Sequoia'' Image:Sequoia sempervirens foliage cones Mount Tamalpais 1.jpg, '' Sequoia'' branch of seed cones. Image:Sequoia sempervirens - foliage.jpg, '' Sequoia'' Image:Sequoia sempervirens2.jpg, '' Sequoia'' Image:Glyptostrobus pensilis 2007.06.28 10.10.35-p6280031.jpg, '' Glyptostrobus europaeus'' may have survived in some refugia pockets of
wetlands A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
along Turkey's
Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
coast until the early
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
'' Glyptostrobus europaeus''
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
from early
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
deposits prove such a recent existence of this species in
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
. So recent fossil pollen have not been found in other countries bordering the
Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
. Image:Glyptostrobus pensilis Villa Taranto.jpg, ''
Glyptostrobus ''Glyptostrobus'' is a small genus of conifers in the family Cupressaceae (formerly in the family Taxodiaceae). The sole living species, '' Glyptostrobus pensilis'', is native to subtropical southeastern China, from Fujian west to southeast Yu ...
'' Image:Pinares canarios de El Hierro 14.jpg, '' Pinus canariensis'' Image:Pinares canarios de El Hierro 02.jpg, '' Pinus canariensis'' Image:Pinus canariensis (Barlovento) 02 ies.jpg, '' Pinus canariensis'' Image:Persea indica.jpg_''
Persea indica ''Persea indica'' is a large, evergreen tree in the laurel family (Lauraceae), native to humid uplands on Madeira and the Canary Islands in the North Atlantic. It belongs to the genus '' Persea'', a group of evergreen trees including the avocado. ...
'' Image:Menispermum dauricum leaves.jpg, '' Menispermum dauricum'' leaves File:Menispermum-dauricum-foliage.jpg, '' Menispermum dauricum'' foliage Image:Castanopsis cuspidata.JPG, ''
Castanopsis ''Castanopsis'', commonly called chinquapin or chinkapin, is a genus of evergreen trees belonging to the beech family, Fagaceae. The genus contains about 140 species, which are today restricted to tropical and subtropical eastern Asia. The Engli ...
'' Image:Castanopsis sieboldii1.jpg, ''
Castanopsis ''Castanopsis'', commonly called chinquapin or chinkapin, is a genus of evergreen trees belonging to the beech family, Fagaceae. The genus contains about 140 species, which are today restricted to tropical and subtropical eastern Asia. The Engli ...
'' Image:Castanopsis sieboldii (Makino) Hatus. — スダジイ by urasimaru.jpg, ''
Castanopsis ''Castanopsis'', commonly called chinquapin or chinkapin, is a genus of evergreen trees belonging to the beech family, Fagaceae. The genus contains about 140 species, which are today restricted to tropical and subtropical eastern Asia. The Engli ...
''
acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...
s. Image:Castanopsis sieboldii2.jpg, ''
Castanopsis ''Castanopsis'', commonly called chinquapin or chinkapin, is a genus of evergreen trees belonging to the beech family, Fagaceae. The genus contains about 140 species, which are today restricted to tropical and subtropical eastern Asia. The Engli ...
'' Image:Ailanthus altissima-1.jpg, ''
Ailanthus ''Ailanthus'' (; derived from ''ailanto,'' an Ambonese Malay, Ambonese word probably meaning "tree of the gods" or "tree of heaven") is a genus of trees belonging to the family Simaroubaceae, in the order Sapindales (formerly Rutales or Geranial ...
'' Image:Ailanthus altissima1.jpg, ''
Ailanthus ''Ailanthus'' (; derived from ''ailanto,'' an Ambonese Malay, Ambonese word probably meaning "tree of the gods" or "tree of heaven") is a genus of trees belonging to the family Simaroubaceae, in the order Sapindales (formerly Rutales or Geranial ...
'' seeds Image:Comptonia peregrina.jpg, '' Comptonia'' Image:Comptonia peregrina BotGardBln1105HabitusFall.JPG, '' Comptonia'' Image:Comptonia peregrina HabitusLeavesFall BotGardBln0906.JPG, '' Comptonia'' Image:Platycarya strobilacea 2.JPG, ''
Platycarya ''Platycarya'' is a genus of flowering plants in the family Juglandaceae native to eastern Asia in China, Korea, and Japan.Flora of China''Platycarya''/ref> The genus was formerly treated as comprising a single species '' Platycarya strobilacea ...
'' Image:Platycarya strobilacea 1.JPG, ''
Platycarya ''Platycarya'' is a genus of flowering plants in the family Juglandaceae native to eastern Asia in China, Korea, and Japan.Flora of China''Platycarya''/ref> The genus was formerly treated as comprising a single species '' Platycarya strobilacea ...
'' Image:Platycarya strobilacea4.jpg, ''
Platycarya ''Platycarya'' is a genus of flowering plants in the family Juglandaceae native to eastern Asia in China, Korea, and Japan.Flora of China''Platycarya''/ref> The genus was formerly treated as comprising a single species '' Platycarya strobilacea ...
'' Image:Astronium balansae.jpg, ''
Astronium ''Astronium'' is a genus of flowering plants in the cashew family, Anacardiaceae. It is native to Central and South America. ''Astronium'' is a genus of dioecious trees. Leaves are deciduous, alternate, and odd-pinnate. Species , Plants of the ...
'' Image:Magnolia sprengeri 01 by Line1.JPG, '' Magnolia sprengeri'' Image:Magnolia sprengeri 'Diva' Leaves 3008px.jpg, '' Magnolia sprengeri''


References

Pliocene Paleobotany Paleontological sites Ankara Province Fossils of Turkey