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Pteridaceae
Pteridaceae is a family of ferns in the order Polypodiales, including some 1150 known species in ca 45 genera (depending on taxonomic opinions), divided over five subfamilies. The family includes four groups of genera that are sometimes recognized as separate families: the adiantoid, cheilanthoid, pteridoid, and hemionitidoid ferns. Relationships among these groups remain unclear, and although some recent genetic analyses of the Pteridales suggest that neither the family Pteridaceae nor the major groups within it are all monophyletic, as yet these analyses are insufficiently comprehensive and robust to provide good support for a revision of the order at the family level. Description Members of Pteridaceae have creeping or erect rhizomes. The leaves are almost always compound and have linear sori that are typically on the margins of the leaves and lack a true indusium, typically being protected by a false indusium formed from the reflexed margin of the leaf. Taxonomy Tradi ...
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Cryptogrammoideae
Cryptogrammoideae is a subfamily of ferns in the family Pteridaceae. The subfamily contains three genera and about 23 species. Taxonomy In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), Cryptogrammoideae is one of the five subfamilies of the family Pteridaceae. Although the subfamily Cryptogrammoideae is similar to the family Cryptogrammaceae proposed by Pichi Sermolli in 1963, that group contained the morphologically similar genus '' Onychium'' (now in the subfamily Pteridoideae) instead of the less morphologically similar genus ''Coniogramme ''Coniogramme'' is one of three genera in the subfamily Cryptogrammoideae of the fern family Pteridaceae. A cultivated species, '' Coniogramme pilosa'', is known as "bamboo fern." Species *'' Coniogramme affinis'' ( C. Presl) Wall. ex Hieron ...''. In 2006, Smith ''et al.'' included Cryptogrammaceae as part of the family Pteridaceae, and in 2011, Christenhusz ''et al.'' listed its three genera in Cryptogrammoideae, ...
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Vittarioideae
Vittarioideae is a subfamily of the fern family Pteridaceae, in the order Polypodiales. The subfamily includes the previous families Adiantaceae (adiantoids or maidenhair ferns) and Vittariaceae (vittarioids or shoestring ferns). Description The subfamily includes two distinct groups of ferns: the adiantoids, consisting of the single genus ''Adiantum'', and the vittarioids, several genera, including ''Vittaria'', which typically have highly reduced leaves, usually entire, and an epiphytic habit. The ferns historically considered as ''Adiantum'' include both petrophilic and terrestrial plants. The vittarioid ferns are primarily epiphytic in tropical regions and all have simple leaves with sori that follow the veins and lack true indusia; the sori are most often marginal with a false indusium formed from the reflexed leaf margin. The family also includes a species, '' Vittaria appalachiana'', that is highly unusual in that the sporophyte stage of the life cycle is absent. This spe ...
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Adiantaceae
Vittarioideae is a subfamily of the fern family Pteridaceae, in the order Polypodiales. The subfamily includes the previous families Adiantaceae (adiantoids or maidenhair ferns) and Vittariaceae (vittarioids or shoestring ferns). Description The subfamily includes two distinct groups of ferns: the adiantoids, consisting of the single genus ''Adiantum'', and the vittarioids, several genera, including ''Vittaria'', which typically have highly reduced leaves, usually entire, and an epiphytic habit. The ferns historically considered as ''Adiantum'' include both petrophilic and terrestrial plants. The vittarioid ferns are primarily epiphytic in tropical regions and all have simple leaves with sori that follow the veins and lack true indusia; the sori are most often marginal with a false indusium formed from the reflexed leaf margin. The family also includes a species, '' Vittaria appalachiana'', that is highly unusual in that the sporophyte stage of the life cycle is absent. This spe ...
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Polypodiales
The order Polypodiales encompasses the major lineages of polypod ferns, which comprise more than 80% of today's fern species. They are found in many parts of the world including tropical, semitropical and temperate areas. Description Polypodiales are unique in bearing sporangia with a vertical annulus interrupted by the stalk and stomium. These sporangial characters were used by Johann Jakob Bernhardi to define a group of ferns he called the "Cathetogyratae"; the Pteridophyte Phylogeny Group has suggested reviving this name as the informal term cathetogyrates, to replace the ambiguously circumscribed term "polypods" when referring to the Polypodiales. The sporangia are born on stalks 1–3 cells thick and are often long-stalked. (In contrast, the Hymenophyllales have a stalk composed of four rows of cells.) The sporangia do not reach maturity simultaneously. Many groups in the order lack indusia, but when present, they are attached either along the edge of the indusium or in its ...
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Pteridoideae
''Pteridoideae'' is one of the five subfamilies of the fern family Pteridaceae. This subfamily contains about 14 genera and around 400 species. Taxonomy Phylogeny The following diagram shows a likely phylogenic relationship between Pteridoideae and the other the Pteridaceae subfamilies. The cladogram below shows one hypothesis for the evolutionary relationships among the genera of the Pteridoideae, based on a maximum likelihood analysis using six plastid markers. The authors of the study identified four major clades. The ''Pteris'' and JAPSTT clades are found worldwide; the GAPCC clade is pantropical; and the ''Actiniopteris''+''Onychium'' clade is restricted to the Old World. Genera The Pteridophyte Phylogeny Group classification of 2016 (PPG I) recognized 13 genera. Shortly afterwards, the genus ''Gastoniella'' was created for three species formerly placed in ''Anogramma'' which a molecular phylogenetic analysis had shown to be distinct. *'' Actiniopteris'' Link *'' Anogra ...
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Cheilanthoideae
Cheilanthoideae is one of the five subfamilies of the fern family Pteridaceae. The subfamily is thought to be monophyletic, but some of the genera into which it has been divided are not, and the taxonomic status of many of its genera and species remains uncertain, with radically different approaches in use . Phylogenic relationships The following phylogram shows a likely relationship between Cheilanthoideae and the other Pteridaceae subfamilies. Although subfamily Cheilanthoideae itself is thought to be monophyletic, many of the genera into which it has been divided (including ''Cheilanthes'', ''Doryopteris'', ''Notholaena'', and ''Pellaea'') have been shown to be polyphyletic. Genera The division of the subfamily Cheilanthoideae into genera and species remains uncertain . Christenhusz et al. (2011), the Pteridophyte Phylogeny Group classification of 2016 (PPG I), and the November 2019 version of the ''Checklist of Ferns and Lycophytes of the World'' (''World Ferns'' 8.11) ...
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Parkerioideae
Parkerioideae, Synonym (taxonomy), synonym Ceratopteridoideae, is one of the five subfamilies in the fern family Pteridaceae. It includes only the two genera ''Acrostichum'' and ''Ceratopteris''. The following diagram shows a likely phylogenic relationship between the two Parkerioideae genera and the other Pteridaceae subfamilies. References

Pteridaceae Plant subfamilies {{Pteridaceae-stub ...
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Argyrochosma
''Argyrochosma'' is a genus of ferns known commonly as false cloak ferns. The genus is included in the Cheilanthoideae subfamily of the Pteridaceae. Species now in this genus were previously treated as members of related genera ''Notholaena'' or ''Pellaea'' but were segregated into their own genus in 1987. These ferns, of which there are about 20 species, are mostly native to the Americas, from North to South and including the Caribbean, while one species, ''A. connectens'', is known from Sichuan, China. They are commonly found growing in cracks between rocks. Their leaves are generally shorter than 40 centimeters and have rounded bluish or grayish green segments. Often the lower surface of the segments is coated in a white dust, and the sporangia A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point ...
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Acrostichum
''Acrostichum'' is a fern genus in the Parkerioideae subfamily of the Pteridaceae. It was one of the original pteridophyte genera delineated by Linnaeus. It was originally drawn very broadly, including all ferns that had sori apparently "acrostichoid", or distributed in a uniform mass across the back of the frond, rather than organized in discrete sori. This led Linnaeus to include such species as ''Asplenium platyneuron'' in the genus, because the specimen he received had sori so crowded that it appeared acrostichoid. Since '' Acrostichum aureum'' is regarded as the type for the genus, it is now narrowly circumscribed only to the natural genus of three species, that are allied to the genus '' Ceratopteris''. They are collectively known as the leather ferns or leather swamp ferns, genus members commonly being found in swamps. The species of ''Acrostichum'' are massive ferns, with fronds up to 12 feet (3.5 meters) tall, that depend on a semi-aquatic existence. They do not ...
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Onychium (plant)
''Onychium'' is a genus of ferns in the subfamily Pteridoideae of the family Pteridaceae Pteridaceae is a family of ferns in the order Polypodiales, including some 1150 known species in ca 45 genera (depending on taxonomic opinions), divided over five subfamilies. The family includes four groups of genera that are sometimes recogni .... Species , the ''Checklist of Ferns and Lycophytes of the World'' recognized the following species and hybrids: *'' Onychium cryptogrammoides'' Christ *'' Onychium divaricatum'' (Poir.) Alston *'' Onychium japonicum'' (Thunb.) Kunze *'' Onychium kholianum'' Fraser-Jenk. & S.Matsumoto *'' Onychium lucidum'' (D.Don) Spreng. * ''Onychium'' × ''matsumotoi'' Fraser-Jenk. & Kandel *'' Onychium moupinense'' Ching *'' Onychium plumosum'' Ching *'' Onychium siliculosum'' (Desv.) C.Chr. *'' Onychium tenuifrons'' Ching *'' Onychium vermae'' Fraser-Jenk. & Khullar References Pteridaceae Fern genera {{Pteridaceae-stub ...
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Astrolepis
''Astrolepis'' is a small genus of ferns in the family Pteridaceae. It was formed in 1992 from species previously placed in ''Cheilanthes'' and ''Notholaena''. The name is derived from the Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ... words ἄστρον (), meaning "star," and λεπίς (), meaning "scale," referring to the star-like scales on adaxial blade surfaces. Members of the genus are commonly known as star-scaled cloak ferns and are native to the Americas. Species , the ''Checklist of Ferns and Lycophytes of the World'' recognized the following species: *'' Astrolepis cochisensis'' (Goodd.) D.M.Benham & Windham – Cochise scaly cloakfern *'' Astrolepis crassifolia'' (Houlston & T.Moore) D.M.Benham & Windham *'' Astrolepis deltoidea'' (Baker) J.B.Beck & Windham ...
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Pteris
''Pteris'' (brake) is a genus of about 300 species of ferns in the subfamily Pteridoideae of the family Pteridaceae. They are native to tropical and subtropical regions, southward to New Zealand, Australia, and South Africa, north to Japan and North America. 78 species (35 endemic) are found in China. Some species of Pteris have considerable economic and ecological value, such as Pteris multifida, Pteris ensiformis, Pteris vittata can be used for ornamental purposes; as a hyperaccumulator, Pteris multifida and Pteris vittata can be used to control soil pollution. Many of them have linear frond segments, and some have sub-palmate division. Like other members of the Pteridaceae, the frond margin is reflexed over the marginal sori. The outermost layer is the single layered epidermis without stomata. The cortex is differentiated into outer and inner cortical region. The vascular cylinder is an amphiphloic siphonostele. The term "brake", used for members of this genus, is ...
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