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Carpinus
Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbeam'' derives from the hardness of the woods (likened to horn) and the Old English ''beam'' "tree" (cognate with Dutch ‘’Boom’’ and German ''Baum''). The American hornbeam is also occasionally known as blue-beech, ironwood, or musclewood, the first from the resemblance of the bark to that of the American beech ''Fagus grandifolia'', the other two from the hardness of the wood and the muscled appearance of the trunk and limbs. The botanical name for the genus, ''Carpinus'', is the original Latin name for the European species, although some etymologists derive it from the Celtic for a yoke. Taxonomy Formerly some taxonomists segregated them with the genera ''Corylus'' (hazels) and ''Ostrya'' (hop-hornbeams) in a separate family, Coryla ...
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Carpinus Betulus 001
Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbeam'' derives from the hardness of the woods (likened to horn) and the Old English ''beam'' "tree" (cognate with Dutch ‘’Boom’’ and German ''Baum''). The American hornbeam is also occasionally known as blue-beech, ironwood, or musclewood, the first from the resemblance of the bark to that of the American beech ''Fagus grandifolia'', the other two from the hardness of the wood and the muscled appearance of the trunk and limbs. The botanical name for the genus, ''Carpinus'', is the original Latin name for the European species, although some etymologists derive it from the Celtic for a yoke. Taxonomy Formerly some taxonomists segregated them with the genera ''Corylus'' (hazels) and ''Ostrya'' (hop-hornbeams) in a separate family, Coryla ...
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Coryloideae
Coryloideae is a subfamily in the woody angiosperm family Betulaceae, commonly known as the birch family, and consists of four extant genera - ''Corylus'' L., ''Ostryopsis'' Decne., ''Carpinus'' L., and ''Ostrya'' Scop. These deciduous trees and shrubs are primarily distributed in the boreal and cool temperate zones of the Northern Hemisphere, with the majority occurring in Asia, many occurring in North America and a few species occurring as far south as South America.Stults, D. Z. & Axsmith, B. J. 2009. Betulaceae From The Pliocene And Pleistocene Of Southwest Alabama, Southeastern United States. ''Review Of Palaeobotany And Palynology'', 155, 25-31. Synapomorphies such reduced staminate flowers, advanced wood anatomy features, and the presence of spermidines in pollen define the Coryloideae.Chen, Z. D., Manchester, S. R. & Sun, H. Y. 1999. Phylogeny And Evolution Of The Betulaceae As Inferred From Dna Sequences, Morphology, And Paleobotany. ''American Journal of Botany'', 86, 1 ...
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Carpinus Perryae
''Carpinus perryae'' is an extinct species of hornbeam known from fossil fruits found in the Klondike Mountain Formation deposits of northern Washington state, dated to the early Eocene Ypresian stage. Based on described features, ''C. perryae'' is the oldest definite species in the genus ''Carpinus''. History and classification ''Carpinus perryae'' was described from two type specimens, the holotype and paratype fertile bracts, numbered UWBM 71171, specimen A and B respectively. Both compression fossil specimens are preserved together on a single rock slab that is part of the Burke Museum paleobotanical collections. Found at the Klondike Mountain Formations UWBM site B2737, which is designated the type locality, the fossils were described by paleobotanists Pigg, Manchester, and Wehr (2003) along with ''Corylus johnsonii'' and '' Palaeocarpinus barksdaleae''. The specific epithet ''perryae'' was coined as a matronym recognizing Madilane Perry for her work founding the Stonero ...
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American Hornbeam
''Carpinus caroliniana'', the American hornbeam, is a small hardwood tree in the genus ''Carpinus''. American hornbeam is also known as blue-beech, ironwood, musclewood and muscle beech. It is native to eastern North America, from Minnesota and southern Ontario east to Maine, and south to eastern Texas and northern Florida. It also grows in Canada (southwest Quebec and southeast Ontario). It occurs naturally in shaded areas with moist soil, particularly near the banks of streams or rivers, and is often a natural constituent understory species of the riverine and maritime forests of eastern temperate North America. Description American hornbeam is a small tree reaching heights of , rarely , and often has a fluted and crooked trunk. The bark is smooth and greenish-grey, becoming shallowly fissured in all old trees. The leaves are alternate, long, with prominent veins giving a distinctive corrugated texture, and a serrated margin. The male and female catkins appear in spring at t ...
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Klondike Mountain Formation
The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington state. The formation, named for the type location designated in 1962, Klondike Mountain north of Republic, Washington, is composed of volcanic rocks in the upper unit and volcanics plus lacustrine (lakebed) sedimentation in which a lagerstätte with exceptionally well-preserved plant and insect fossils has been found, along with fossil epithermal hot springs. The formation is the youngest in a group of formations which belong to the Challis Sequence rocks. The formation unconformably overlies rocks of the Eocene Sanpoil Volcanics and much older Triassic and Permian formations. The formation is bounded on its edges by a series of high-angle strike slip faults, which have contained the Klondike Mountain Formation in a series of graben structures, such as the Republic Graben. Public access to a fossiliferous outcrop at the north end of Republic is ...
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Betulaceae
Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams numbering a total of 167 species. They are mostly natives of the temperate Northern Hemisphere, with a few species reaching the Southern Hemisphere in the Andes in South America. Their typical flowers are catkins and often appear before leaves. In the past, the family was often divided into two families, Betulaceae (''Alnus'', ''Betula'') and Corylaceae (the rest). Recent treatments, including the Angiosperm Phylogeny Group, have described these two groups as subfamilies within an expanded Betulaceae: Betuloideae and Coryloideae. Betulaceae flowers are monoecious, meaning that they have both male and female flowers on the same tree. Their flowers present as catkins and are small and inconspicuous, often with reduced perianth parts. These flowers have large feathery stamen and produce a high volume of ...
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Hazel
The hazel (''Corylus'') is a genus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan . though some botanists split the hazels (with the hornbeams and allied genera) into a separate family Corylaceae. The fruit of the hazel is the hazelnut. Hazels have simple, rounded leaves with double-serrate margins. The flowers are produced very early in spring before the leaves, and are monoecious, with single-sex catkins. The male catkins are pale yellow and long, and the female ones are very small and largely concealed in the buds, with only the bright-red, 1-to-3 mm-long styles visible. The fruits are nuts long and 1–2 cm diameter, surrounded by an involucre (husk) which partly to fully encloses the nut. ...
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Corylus
The hazel (''Corylus'') is a genus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan . though some botanists split the hazels (with the hornbeams and allied genera) into a separate family Corylaceae. The fruit of the hazel is the hazelnut. Hazels have simple, rounded leaves with double-serrate margins. The flowers are produced very early in spring before the leaves, and are monoecious, with single-sex catkins. The male catkins are pale yellow and long, and the female ones are very small and largely concealed in the buds, with only the bright-red, 1-to-3 mm-long styles visible. The fruits are nuts long and 1–2 cm diameter, surrounded by an involucre (husk) which partly to fully encloses the nut ...
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Republic, Washington
Republic is a city in Ferry County, Washington, United States. The population was 1,073 at the 2010 census, a 12.5% increase over the 2000 census. It is the county seat of Ferry County. It was the largest mining camp in the Republic Mining District, and home to the "Hot Air Line" railway. Geography Republic is located at (48.648159, −118.734947). According to the United States Census Bureau, the city has a total area of , all of it land. Republic is located near the source of the Sanpoil River in a long graben valley bordered by the Okanagan Highlands to the west and the Kettle mountain range to the east. Curlew Lake, long (Elev: ), provides fishing and boating to summer visitors northeast of Republic. Swan Lake is small mountain lake to the south of Republic and also serves to be a popular lake for local residents and tourists to visit. Republic is surrounded by the Colville National Forest and to the south is the Colville Indian Reservation. History Republic Mining ...
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Early Eocene
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian is consistent with the lower Eocene. Events The Ypresian Age begins during the throes of the Paleocene–Eocene Thermal Maximum (PETM). The Fur Formation in Denmark, the Messel shales in Germany, the Oise amber of France and Cambay amber of India are of this age. The Eocene Okanagan Highlands are an uplands subtropical to temperate series of lakes from the Ypresian. Stratigraphic definition The Ypresian Stage was introduced in scientific literature by Belgian geologist André Hubert Dumont in 1850. The Ypresian is named after the Flemish city of Ypres in Belgium (spelled ''Ieper'' in Dutch). The definitions of the original stage were totally different from the modern ones. The Ypresian shares its name with the Belgian Ieper Group (French: ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root to be ...
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