The Eudaminae are a

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

skipper butterflies Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...

( family Hesperiidae). Their original type genus ''Eudamus'' is today a

junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...

of '' Urbanus''. They are largely found in the Neotropics, with some extending into temperate North America, and one genus, ''

Lobocla ''Lobocla'' is a genus of skippers in the family Hesperiidae subfamily Eudaminae, within which they are placed in subtribe Loboclina. Unlike the rest of the Eudaminae, which have a New World distribution, the genus is endemic to Asia. Species L ...

'', endemic to East Asia. (2009): Tree of Life Web Project &ndash
Eudaminae
Version of 2009-JUN-11. Retrieved 2009-DEC-24.

Taxonomy

The Eudaminae have been recent subject to significant taxonomic revisions based on genome analysis, including by Brower & Warren 2009, and Li et al. 2019. Historically, the subfamily has been included as tribe Eudamini in subfamily Pyrginae, based on perceived similarities with two of the tribes in that subfamily, the

Celaenorrhinini The Celaenorrhinini are a tribe of spread-winged skippers in the skipper butterfly subfamily Pyrginae. When the Eudaminae were still included in the Pyrginae as a tribe, their delimitation versus the Celaenorrhinini was disputed, and there were ...

and Pyrgini. As of Li et al. 2019, the Eudaminae are divided into four tribes:

Entheini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are largely found in the Neotropics, with some extending into temperate North America ...

Phocidini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are largely found in the Neotropics, with some extending into temperate North Ameri ...

Eudamini The Eudaminae are a subfamily of skipper butterflies (family (biology), family Hesperiidae). Their original biological type, type genus ''Eudamus'' is today a junior synonym of ''Urbanus (genus), Urbanus''. They are largely found in the Neotropic ...

and

Oileidini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are largely found in the Neotropics, with some extending into temperate North America ...

Current status and subdivisions

Except where otherwise noted, the classification below follows Li et al., 2019:

Tribe Entheini

*''

Drephalys ''Drephalys'' is a Neotropical butterfly genus in the family Hesperiidae, in which it is placed in tribe Entheini. The genus was established in 1893 by Edward Yerbury Watson, who assigned ''Eudamus helixus'' (now '' Drephalys helixus'') as typ ...

'' Watson, 1893 *''

Udranomia ''Udranomia'' is a genus of Neotropical butterflies in the family Hesperiidae, in which it is placed in tribe Entheini. It was established in 1870 by Arthur Gardiner Butler. Species Per Li et al. 2019, the genus contains the following species ...

'' A. Butler, 1870 *''

Phanus ''Phanus'' is a genus of butterflies in the family Hesperiidae, in which it is placed in tribe Entheini. Species *'' Phanus albiapicalis'' Austin, 1993 Mexico *'' Phanus australis'' Miller, 1965 Brazil *'' Phanus confusis'' Austin, 1993 Mexico ...

'' Hübner,

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Hyalothyrus ''Hyalothyrus'' is a genus of skippers in the family Hesperiidae, in which it is placed in tribe Entheini. Species *'' Hyalothyrus infernalis'' (Moschler, 1877) French Guiana, Suriname *'' Hyalothyrus infa'' Evans, 1952 Colombia, Brazil (Amazon ...

'' Mabille, 1878 *''

Entheus ''Entheus'' is a Neotropical genus of skippers in the family Hesperiidae, in which it is placed in the tribe Entheini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today ...

'' Hübner,

/small> *''

Augiades ''Augiades'' is a genus of butterflies in the family Hesperiidae, in which it is placed in tribe Entheini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synon ...

'' Hübner,

/small> *''

Tarsoctenus ''Tarsoctenus'' is a genus of Neotropical butterflies in the family Hesperiidae, in which it is placed in tribe Entheini. Species *'' Tarsoctenus corytus'' (Cramer, 777 Suriname to Colombia, Brazil (Amazonas) **''T. corytus corytus'' Suriname ...

'' E. Watson, 1893

Tribe Phocidini

*'' Phocides'' Hübner,

/small> *'' Pseudonascus'' Austin, 2008 *''

Nascus ''Nascus'' is a genus of Neotropical butterflies in the family Hesperiidae, in which it is placed to tribe Phocidini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a ...

'' E. Watson, 1893 *''

Aurina ''Aurina'' is a genus of skippers in the family Hesperiidae, in which it is placed in tribe Phocidini. Taxonomy Previously considered monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately su ...

'' Evans, 1937 *'' Emmelus'' O. Mielke & Casagrande , 2016 *''

Porphyrogenes ''Porphyrogenes'' is a Neotropical genus of spread-winged skippers in the family Hesperiidae, in which they are placed to tribe Phocidini. Species *'' Porphyrogenes boliva'' Evans, 1952 - Venezuela *'' Porphyrogenes calathana'' (Hewitson, 1868) ...

'' E. Watson, 1893 *''

Nicephellus ''Nicephellus'' is a monotypic genus of Neotropical butterflies in the family Hesperiidae, in which it is placed in tribe Phocidini. The sole species of the genus is ''Nicephellus nicephorus'', also called the two-spotted scarlet-eye, and was or ...

'' Austin, 2008 *''

Salatis ''Salatis'' is a Neotropical genus of spread-winged skippers in the family Hesperiidae, in which they are placed to tribe Phocidini. Species of the genus ''Salatis'' are native to Central America and South America, including in the Amazon River ...

'' Evans, 1952 *'' Salantoia'' Grishin, 2019 *''

Sarmientoia ''Sarmientoia'' is a genus of skippers in the family Hesperiidae, in which they are placed to tribe Phocidini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junio ...

'' Berg, 1897 *'' Bungalotis'' E. Watson, 1893 *''

Euriphellus ''Euriphellus'' is a genus of Neotropical butterflies in the family Hesperiidae, in which it is placed in the tribe Phocidini. The genus was separated from ''Dyscophellus'' by George Traut Austin in 2008, and originally contained a single species ...

'' Austin, 2008 *''

Dyscophellus ''Dyscophellus'' is a genus of largely Neotropical butterflies in the family Hesperiidae. The genus has been the subject of recent revisions based on genome analysis. It is currently placed in tribe (biology), tribe Phocidini within the subfamily ...

'' Godman & Salvin, 1893 *'' Phareas'' Westwood, 1852 *'' Ornilius'' Grishin, 2022 *'' Fulvatis'' Grishin, 2022 *''

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'' Grishin, 2022

Tribe Eudamini

Subtribe Eudamina

*''

Cecropterus ''Cecropterus'' is a genus of skipper (butterfly), skippers in the family Hesperiidae, within which it is placed in subtribe Eudamina. It has three subgenera: ''Cecropterus'', ''Thorybes'' and ''Murgaria''. Species As of 2019, the genus holds th ...

'' Herrich-Schäffer, 1869 *'' Spicauda'' Grishin, 2019 *'' Urbanus'' Hübner,

807 __NOTOC__ Year 807 ( DCCCVII) was a common year starting on Friday (link will display the full calendar) of the Julian calendar. Events By place Abbasid Empire and Byzantine Empire * Emperor Nikephoros I is forced to sue for peace, on ...

/small> *'' Telegonus'' Hübner,

/small> *'' Autochton'' Hübner, 1823 *''

Spathilepia ''Spathilepia'' is a butterfly genus in the family Hesperiidae (Eudaminae). The genus is monotypic containing the single species ''Spathilepia clonius'' found in Texas, Central America and Colombia to Brazil and Argentina. They are native to Mexi ...

'' A. Butler, 1870 *''

Astraptes ''Astraptes'', commonly known as the flasher butterflies, is a genus of skipper butterflies in the subfamily Eudaminae. They are found in the Nearctic and Neotropical ecozones. Species The following species are recognised in the genus ''Astrapte ...

'' Hübner,

/small> *''

Narcosius ''Narcosius'' is a genus of butterflies in the family Hesperiidae (Eudaminae The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are ...

'' Steinhauser, 1986 *''

Proteides ''Proteides'' is a Neotropical genus of spread-winged skipper butterflies in the family Hesperiidae. Species The following two species are recognised in the genus ''Proteides'' *'' Proteides maysi'' (Lucas, 1857) Cuba *'' Proteides mercurius'' ...

'' Hübner,

/small> *''

Epargyreus ''Epargyreus'' is the genus of flasher butterflies, in the subfamily Eudaminae of skipper (butterfly), skipper butterflies. The species are found in the Nearctic realm, Nearctic and Neotropical realms. Taxonomy Etymology The genus name is derive ...

'' Hübner,

/small> *''

Chioides ''Chioides'' is a mainly Neotropical genus of skipper butterflies in the family Hesperiidae (Eudaminae) They range from the south-western United States to Argentina and there are several Antillean endemic species. They are robust and have hindw ...

'' Lindsey, 1921

Subtribe Loboclina

*''

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'' R. Williams, 1927 *'' Zeutus'' Grishin, 2019 *''

'' Moore, 1884 *'' Lobotractus'' Grishin, 2019 *''

Codatractus ''Codatractus'' is a Neotropical and Nearctic genus in the family Hesperiidae (Eudaminae). Species The following species are recognised in the genus ''Codatractus'': * ''Codatractus cyledis'' (Dyar, 1912) - Mexico * ''Codatractus bryaxis'' (Hew ...

'' Lindsey, 1921 *''

Zestusa ''Zestusa'' is a genus of skippers in the family Hesperiidae subfamily Eudaminae. Species of the genus occur in Mexico and the western United States. Species The following species are recognised in the genus ''Zestusa'': * ''Zestusa dorus '' ...

'' Lindsey, 1925 *''

Ridens ''Ridens'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species *''Ridens allyni'' Freeman, 1979 - Mexico, Guatemala *''Ridens bidens'' Austin, 1998 - Brazil (Rondônia). *''Ridens biolleyi'' (Mabille, 1900) - Cos ...

'' Evans, 1952 *''

Venada ''Venada'' is a genus of skipper (butterfly), skippers in the family Hesperiidae. Species ''Venada'' consists of the following species: *''Venada advena'' *''Venada cacao'' *''Venada daneva'' *''Venada lamella'' Burns ''in'' Burns, Janzen, Hal ...

'' Evans, 1952

Subtribe Cephisina

*''

Cephise ''Cephise'' is a Neotropical genus of skipper butterflies of the subfamily Eudaminae, within which it is placed in subtribe Cephisina. Species Per Li et al. 2019, the genus contains the following species: *''Cephise aelius'' (Plötz, 1880) – ...

'' Evans, 1952

Subtribe Telemiadina

*''

Ectomis ''Ectomis'' is a genus of Neotropical and Nearctic butterflies in the family Hesperiidae (Eudaminae). Species The following species are recognised in the genus ''Ectomis'': * ''Ectomis'' (''Asina'') ** ''Ectomis gyges'' Evans, 1952 - Venezuela ...

'' Mabille, 1878 *''

Telemiades ''Telemiades'' is a genus of Neotropical butterflies in the family Hesperiidae (subfamily Eudaminae). Species The following species are recognised in the genus ''Telemiades'': *''Telemiades amphion'' (Carl Geyer, Geyer, 1832) **''T. amphion amph ...

'' Hübner,

/small> *''

Polygonus Polygonus may refer to: * Polygonus (genus), a genus of Nearctic and Neotropical spread-winged skippers * Polygonus (mythology) In Greek mythology, Polygonus (Ancient Greek: Πολύγονον means 'prolific') was the Thracian son of the sea go ...

'' Hübner,

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/small>

Tribe Oileidini

Subtribe Oileidina

*'' Flattoides'' Grishin, 2019 *''

Oileides ''Oileides'' is a genus of Neotropical spread-winged skippers in the family Hesperiidae. Species The following species are recognised in the genus ''Oileides'': *'' Oileides fenestratus '' (Gmelin, 790 - French Guiana *'' Oileides guyanensis'' ...

'' Hübner,

/small>

Subtribe Typhedanina

*''

Typhedanus ''Typhedanus'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species The following species are recognosed in the genus ''Typhedanus'': * ''Typhedanus ampyx'' (Godman & Salvin,

893 __FORCETOC__ Year 893 ( DCCCXCIII) was a common year starting on Monday (link will display the full calendar) of the Julian calendar. Events By place Europe * Vladimir, ruler (''khan'') of the Bulgarian Empire, is dethroned by his fat ...

- Venezuela to Mexico, Amazon ...

'' A. Butler, 1870 *''

Oechydrus ''Oechydrus'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species There are two species recognised in the genus ''Oechydrus'': * '' Oechydrus chersis'' (Herrich-Schäffer, 1869) **''O. c. chersis'' (Herrich-Sc ...

'' E. Watson, 1893 *''

Cogia ''Cogia'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species The following species are recognised in the genus ''Cogia'': *''Cogia abdul'' Hayward, 1947 *''Cogia azila'' Evans, 1953 *''Cogia aziris'' (Hewitson, ...

'' A. Butler, 1870 *''

Nerula ''Nerula'' is a genus of dicot skippers in the butterfly family Hesperiidae (Eudaminae The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus'' ...

'' Mabille, 1888 *''

Marela Marela is a town and sub-prefecture in the Faranah Prefecture in the Faranah Region of Guinea Guinea ( ),, fuf, 𞤘𞤭𞤲𞤫, italic=no, Gine, wo, Gine, nqo, ߖߌ߬ߣߍ߫, bm, Gine officially the Republic of Guinea (french: Rép ...

'' Mabille, 1903

Footnotes and references

Butterfly subfamilies Taxa named by Paul Mabille {{Hesperiidae-stub