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Holostei
Holostei is a group of ray-finned bony fish. It is divided into two major clades, the Halecomorphi, represented by a single living species, the bowfin (''Amia calva''), as well as the Ginglymodi, the sole living representatives being the gars (Lepisosteidae), represented by seven living species in two genera (''Atractosteus'', ''Lepisosteus''). The earliest members of the clade appeared during the Early Triassic, over 250 million years ago. Holostei was thought to be regarded as paraphyletic. However, a recent study provided evidence that the Holostei are the closest living relates of the Teleostei, both within the Neopterygii. This was found from the morphology of the Holostei, for example presence of a paired vomer. Holosteans are closer to teleosts than are the chondrosteans, the other group intermediate between teleosts and cartilaginous fish, which are regarded as (at the nearest) a sister group to the Neopterigii. The spiracles of holosteans are reduced to vestigial remnan ...
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Actinopterygii
Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or horny spines (rays), as opposed to the fleshy, lobed fins that characterize the class Sarcopterygii (lobe-finned fish). These actinopterygian fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the link or connection between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). By species count, actinopterygians dominate the vertebrates, and they constitute nearly 99% of the over 30,000 species of fish. They are ubiquitous throughout freshwater and marine environments from the deep sea to the highest mountain streams. Extant species can range in size from ''Paedocypris'', at , to the massive ocean sunfish, at , and the long-bodied oarfish, at . The vast majority of Actinopt ...
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Ray-finned Fish
Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or horny spines (rays), as opposed to the fleshy, lobed fins that characterize the class Sarcopterygii (lobe-finned fish). These actinopterygian fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the link or connection between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). By species count, actinopterygians dominate the vertebrates, and they constitute nearly 99% of the over 30,000 species of fish. They are ubiquitous throughout freshwater and marine environments from the deep sea to the highest mountain streams. Extant species can range in size from ''Paedocypris'', at , to the massive ocean sunfish, at , and the long-bodied oarfish, at . The vast majority of Actinoptery ...
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Neopterygii
Neopterygii (from Greek νέος ''neos'' 'new' and πτέρυξ ''pteryx'' 'fin') is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant fishes, and over half of all living vertebrate species. While living holosteans include only freshwater taxa, teleosts are diverse in both freshwater and marine environments. Many new species of teleosts are scientifically described each year. Fossil evidence for crown group neopterygians goes back at least 251 million years to the Induan stage of the Early Triassic epoch, however, one study incorporating morphological data from fossils and molecular data from nuclear and mitochondrial DNA, places this divergence date at least 284 mya (million years ago), during the Artinskian stage of the Early Permian. Another study suggests an even earlier split (360 myr ago, near the Devonian-Carboniferous boundary). Evolution and diversity Liv ...
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Amia Calva
The bowfin (''Amia calva'') is a bony fish, native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being the sole surviving species of the Halecomorphi, a group of fish that first appeared during the Early Triassic, around 250 million years ago. The bowfin is often considered a "primitive fish" because they have retained some morphological characteristics of their early ancestors. The closest living relatives of bowfins are gars, with the two groups being united in the clade Holostei. Bowfins are demersal freshwater piscivores, commonly found throughout much of the eastern United States, and in southern Ontario and Quebec. Fossil deposits indicate Amiiformes were once widespread in both freshwater and marine environments across North and South America, Europe, Asia, and Africa. Now, their range is limited to much of the eastern United States and adjacent southern Canada, including th ...
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Parasemionotiformes
Parasemionotiformes is an extinct order of neopterygian ray-finned fish that existed globally during the Triassic period. It comprises the families Parasemionotidae and Promecosominidae. Many of the included genera are monotypic and most species lived during the Early Triassic epoch.Romano et al. (2016): Marine Early Triassic Osteichthyes from Spiti, Indian Himalayas. Swiss Journal of Palaeontology 135: 275-294 https://doi.org/10.1007/s13358-015-0098-6 Parasemionotiforms were normally small to medium-sized fishes. They were predominantly marine. Evolutionary relationships Parasemionotiformes are neopterygians, which is the clade that encompasses the vast majority of living ray-finned fishes (Actinopterygii) and about half of all living species of vertebrates. Neopterygii are divided into Teleostei and Holostei. The latter represents a depauperate group today but used to be a diverse clade especially during the Mesozoic Era. The only surviving members of the Holostei are the g ...
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Bowfin
The bowfin (''Amia calva'') is a bony fish, native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being the sole surviving species of the Halecomorphi, a group of fish that first appeared during the Early Triassic, around 250 million years ago. The bowfin is often considered a "primitive fish" because they have retained some morphological characteristics of their early ancestors. The closest living relatives of bowfins are gars, with the two groups being united in the clade Holostei. Bowfins are demersal freshwater piscivores, commonly found throughout much of the eastern United States, and in southern Ontario and Quebec. Fossil deposits indicate Amiiformes were once widespread in both freshwater and marine environments across North and South America, Europe, Asia, and Africa. Now, their range is limited to much of the eastern United States and adjacent southern Canada, including th ...
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Teleostei
Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest class (biology), infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all neontology, extant species of fish. Teleosts are arranged into about 40 order (biology), orders and 448 family (biology), families. Over 26,000 species have been described. Teleosts range from giant oarfish measuring or more, and ocean sunfish weighing over , to the minute male anglerfish ''Photocorynus spiniceps'', just long. Including not only torpedo-shaped fish built for speed, teleosts can be flattened vertically or horizontally, be elongated cylinders or take specialised shapes as in anglerfish and seahorses. The difference between teleosts and other bony fish lies mainly in their jaw bones; teleosts have a movable premaxilla and corresponding modifications in the jaw musculature which make it possible for them to cranial kinesi ...
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Halecomorphi
Halecomorphi is a taxon of ray-finned bony fish in the clade Neopterygii. The sole living Halecomorph is the bowfin (''Amia calva''), but the group contains many extinct species in several families (including Amiidae, Caturidae, Liodesmidae, Sinamiidae) in the order Amiiformes, as well as the extinct orders Ionoscopiformes, Panxianichthyiformes, and Parasemionotiformes. The fossil record of halecomorphs goes back at least to the Early Triassic epoch. The Halecomorphi exhibit a combination of ancestral features, such as most heavily mineralized scales, but also by more derived or "modern" features, particularly in the structure of the skull (e.g. position and shape of preopercles). Unique derived traits (synapomorphies) of the Halecomorphi include: *Unique jaw articulation in which the quadrate and symplectic participate in the joint. *Lengthened dorsal fins (in some species) *Two biconcave vertebrae per segment in the posterior body region (a condition known as dip ...
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Ginglymodi
Ginglymodi is a clade of ray-finned fish containing modern-day gars (Lepisosteidae) and their extinct relatives, including the family Lepidotidae and the orders Semionotiformes and Kyphosichthyiformes, and various other extinct taxa. Ginglymodi is one of the two major subgroups of the infraclass Holostei, the other one being Halecomorphi, which contains the bowfin and its fossil relatives. Fossil record The fossil record of ginglymodians goes back at least to the Anisian stage of the Triassic period, over 240 million years ago. '' Eosemionotus'' is one of the earliest ginglymodians. '' Acentrophorus'', another taxon from the Wuchiapingian stage of the late Permian, and '' Paracentrophorus'' from the Early Triassic epoch, could be even earlier members of the group. Ginglymodi was diverse and widespread during the Mesozoic era, but they represent a depauperate lineage today. The group first evolved in marine environments, but several lineages made separate transitions into fre ...
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Amiiformes
The Amiiformes order of fish has only one extant species, the bowfin (''Amia calva''). These Amiiformes are found in the freshwater systems of North America, in the United States and parts of southern Canada. They live in freshwater streams, rivers, and swamps. Bowfins are not on the endangered list. They have the ability to go to the surface to breathe air if the water level is too low. Characteristics of Amiiformes are a cylindrical body with a long dorsal fin, single gular plate, heterocercal caudal fin, 10 to 13 flattened branchiostegal rays, maxilla included in gape, and prominent ocellus near upper base of caudal fin. Evolution and diversity The extinct species of the Amiiformes can be found as fossils in Asia and Europe, but the bowfin is the last living species in the order. Amiiformes is therefore the last surviving order of Halecomorphi, the clade to which the bowfin and its fossil relatives belong. Other orders, such as the Parasemionotiformes, are all extinct. ...
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Spiracle (vertebrates)
Spiracles () are openings on the surface of some animals, which usually lead to respiratory systems. The spiracle is a small hole behind each eye that opens to the mouth in some fish. In the jawless fish, the first gill opening immediately behind the mouth is essentially similar to the other gill opening. With the evolution of the jaw in the early jawed vertebrates, this gill slit was caught between the forward gill-rod (now functioning as the jaw) and the next rod, the hyomandibular bone, supporting the jaw hinge and anchoring the jaw to the skull proper. The gill opening was closed off from below, the remaining opening was small and hole-like, and is termed a spiracle. In many species of sharks and all rays the spiracle is responsible for the intake of water into the buccal space before being expelled from the gills. The spiracle is often located towards the top of the animal allowing breathing even while the animal is mostly buried under sediments. As sharks adapted a faster ...
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Semionotiformes
Semionotiformes is an order of primitive, ray-finned, primarily freshwater fish from the Middle Triassic (Anisian) to the Late Cretaceous (Maastrichtian). The best-known genus is ''Semionotus'' of Europe and North America. Their closest living relatives are gars (Lepisosteidae), with both groups belonging to the clade Ginglymodi within the Holostei. Classification * Order †Semionotiformes Arambourg & Bertin 1958 sensu López-Arbarello 2012 ** Genus ?†''Orthurus'' Kner 1866 ** Genus †'' Sangiorgioichthys'' Tintori & Lombardo 2007 ** Genus †''Luoxiongichthys'' Wen et al. 2011 ** Genus †''Aphanepygus'' Bassani 1879 ** Genus †'' Placidichthys'' Brito 2000 ** Family † Pleurolepididae Lütken 1871 *** Genus †'' Pleurolepis'' Agassiz 1863 non Quenstedt 1852 ** Family †Macrosemiidae Wagner 1860a corrig. Cope 1889 sensu Murray & Wilson 2009 acrosemii Wagner 1860a*** Genus †'' Eusemius'' Vetter 1881 *** Genus †'' Blenniomoeus'' Costa 1850 Calignathus.html"_;"title="' ...
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