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Sinoceratops Zhuchengensis
''Sinoceratops'' is an extinct genus of ceratopsian dinosaur that lived approximately 73 million years ago during the latter part of the Cretaceous Period in what is now Shandong province in China. It was named in 2010 by Xu Xing ''et al.'' for three skulls from Zhucheng, China. The name of its type species ''Sinoceratops zhuchengensis'' means "Chinese horned face from Zhucheng", after the location of its discovery. ''Sinoceratops'' was a medium-sized, averagely-built, ground-dwelling, quadrupedal herbivore. It could grow up to an estimated in length and weigh up to . It was the first ceratopsid dinosaur discovered in China, and the only ceratopsid known from Asia. All other centrosaurines, and all chasmosaurines, are known from fossils discovered in North America, except for possibly ''Turanoceratops''. ''Sinoceratops'' is also significant because it is one of the largest known centrosaurines, and is much larger than any other known basal members of this group. ''Sinocer ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Chasmosaurines
Chasmosaurinae is a subfamily of ceratopsid dinosaurs. They were one of the most successful groups of herbivores of their time. Chasmosaurines appeared in the early Campanian, and became extinct, along with all other non-avian dinosaurs, during the Cretaceous–Paleogene extinction event. Broadly, the most distinguishing features of chasmosaurines are prominent brow horns and long frills lacking long spines; centrosaurines generally had short brow horns and relatively shorter frills, and often had long spines projecting from their frills. Chasmosaurines evolved in western North America (Laramidia). They are currently known definitively from rocks in western Canada, the western United States, and northern Mexico. They were highly diverse and among the most species-rich groups of dinosaurs, with new species frequently described. This high diversity of named species is likely a result of the frill. The distinctive shape of the frill with the hornlets on its edges (epoccipitals) make ...
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Shandong
Shandong ( , ; ; alternately romanized as Shantung) is a coastal province of the People's Republic of China and is part of the East China region. Shandong has played a major role in Chinese history since the beginning of Chinese civilization along the lower reaches of the Yellow River. It has served as a pivotal cultural and religious center for Taoism, Chinese Buddhism and Confucianism. Shandong's Mount Tai is the most revered mountain of Taoism and a site with one of the longest histories of continuous religious worship in the world. The Buddhist temples in the mountains to the south of the provincial capital of Jinan were once among the foremost Buddhist sites in China. The city of Qufu is the birthplace of Confucius and was later established as the center of Confucianism. Confucianism developed from what was later called the Hundred Schools of Thought from the teachings of the Chinese philosopher Confucius. Shandong's location at the intersection of ancient and modern n ...
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Wangshi Group
The Wangshi Group () is a geological Group in Shandong, China whose strata date back to the Coniacian to Campanian stages of the Late Cretaceous.Wangshi Group
in the
Dinosaur remains are among the fossils that have been recovered from the group.Weishampel et al., 2004, pp.593-600


Vertebrate paleofauna


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Fossil eggs

The following fossil eggs were recovered from the Jingangkou Format ...
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Braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skullcap. The remainder of the skull is the facial skeleton. In comparative anatomy, neurocranium is sometimes used synonymously with endocranium or chondrocranium. Structure The neurocranium is divided into two portions: * the membranous part, consisting of flat bones, which surround the brain; and * the cartilaginous part, or chondrocranium, which forms bones of the base of the skull. In humans, the neurocranium is usually considered to include the following eight bones: * 1 ethmoid bone * 1 frontal bone * 1 occipital bone * 2 parietal bones * 1 sphenoid bone * 2 temporal bones The ossicles (three on each side) are usually not included as bones of the neurocranium. There may variably also be extra sutural bones present. Below the ne ...
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Skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium ( facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the front ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Sinoceratops Skull
''Sinoceratops'' is an extinct genus of ceratopsian dinosaur that lived approximately 73 million years ago during the latter part of the Cretaceous Period in what is now Shandong province in China. It was named in 2010 by Xu Xing ''et al.'' for three skulls from Zhucheng, China. The name of its type species ''Sinoceratops zhuchengensis'' means "Chinese horned face from Zhucheng", after the location of its discovery. ''Sinoceratops'' was a medium-sized, averagely-built, ground-dwelling, quadrupedal herbivore. It could grow up to an estimated in length and weigh up to . It was the first ceratopsid dinosaur discovered in China, and the only ceratopsid known from Asia. All other centrosaurines, and all chasmosaurines, are known from fossils discovered in North America, except for possibly ''Turanoceratops''. ''Sinoceratops'' is also significant because it is one of the largest known centrosaurines, and is much larger than any other known basal members of this group. ''Sinocerato ...
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Zhuchengtyrannus
''Zhuchengtyrannus'' (meaning "Zhucheng tyrant") is a genus of tyrannosaurid theropod dinosaur known from the Campanian stage of the Late Cretaceous of Shandong Province, China. It belongs to the subfamily Tyrannosaurinae, and contains a single species, ''Zhuchengtyrannus magnus''. Discovery and naming ''Zhuchengtyrannus'' was first described and named by David W. E. Hone, Kebai Wang, Corwin Sullivan, Xijin Zhao, Shuqing Chen, Dunjin Li, Shuan Ji, Qiang Ji and Xing Xu in 2011 and the type species is ''Zhuchengtyrannus magnus''. The generic name is derived from the word ''Zhucheng'', which refers to the type locality, and ''tyrant'' in reference to its phylogenetic position as a tyrannosaurid. The specific name ''magnus'' meaning "great" in Latin refers to the relatively large size of ''Zhuchengtyrannus''. ''Zhuchengtyrannus'' is known solely from the holotype ZCDM V0031, a nearly complete right maxilla and associated left dentary (lower jaw, both with teeth) discovered around 2 ...
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Zhuchengtitan
''Zhuchengtitan'' (meaning "Zhucheng titan") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous of Shandong, China. It contains a single species, ''Z. zangjiazhuangensis'', named by Mo Jinyou and colleagues in 2017 from a single humerus. ''Zhuchengtitan'' can be identified by the extreme width of the top end of its humerus, as well as the expansion of the deltopectoral crest on its humerus; both of these characteristics indicate that it was likely closely related to ''Opisthocoelicaudia''. However, it differs from the latter by the flatter bottom articulating surface of its humerus. ''Zhuchengtitan'' lived in a floodplain environment alongside ''Shantungosaurus'', ''Zhuchengtyrannus'', and ''Sinoceratops''. Discovery and naming ''Zhuchengtitan'' is known from a single humerus that was discovered in the Wangshi Group within the region of Zhucheng, Shandong, China. The primary fossil localities within this region are the Longgujian, Kugou, and Zangjiazhuang Quarr ...
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Zhuchengceratops
''Zhuchengceratops'' is a genus of extinct leptoceratopsid ceratopsian that lived during the Upper Cretaceous of modern-day China. It was first described in 2010, by Xu ''et al.'', who created the binomial ''Zhuchengceratops inexpectus''. The name is derived from the location of Zhucheng, the Latinized-Greek ''ceratops'', or "horned face", and the unexpected articulated nature of the holotype. The skeleton was found in the Wangshi Group, which is of Late Cretaceous age, and most fossils are only disarticulated bones of ''Shantungosaurus''. ''Zhuchengceratops'' shares many features with Leptoceratopsidae as well as other ceratopsian groups such as Ceratopsidae. The overall size of the taxon was similar to ''Leptoceratops'', although slightly larger. ''Zhuchengceratops'' was analyzed to be in a group with ''Leptoceratops'' and ''Udanoceratops'', although internal relationships of this triplet were unresolved. Discovery and naming ''Zhuchengceratops'' is a derived leptoceratopsid ...
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Shantungosaurus
''Shantungosaurus'' (meaning "''Shandong Lizard''") is a genus of very large saurolophine hadrosaurid dinosaur found in the Late Cretaceous Wangshi Group of the Shandong Peninsula in China, containing a single species, ''Shantungosaurus giganteus''. The stratigraphic interval of ''Shantungosaurus'' ranges from the top of the Xingezhuang Formation to the middle of the Hongtuya Formation, middle to late Campanian in age. ''Shantungosaurus'' is so far the largest hadrosauroid taxon in the world, reaching between to in length and to in body mass. History of discovery First described in 1973, ''Shantungosaurus'' is known from over five incomplete skeletons. Chinese scientist Xing Xu and his colleagues indicate that ''Shantungosaurus'' is very similar to and shares many unique characters with ''Edmontosaurus'', forming a node of an ''Edmontosaurus''–''Shantungosaurus'' clade between North America and Asia, based on the new materials recovered in Shandong. Remains of several i ...
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