Eukaryotic Cells
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Eukaryotic Cells
Eukaryotes () are organisms whose Cell (biology), cells have a cell nucleus, nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the Three-domain system, three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard (archaea), Asgard archaea. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass (ecology), biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as Flagellated cell, flagellated phagotrophs. The ...
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Orosirian
The Orosirian Period (; grc, ὀροσειρά, translit=oroseirá, meaning "mountain range") is the third geologic period in the Paleoproterozoic era (geology), Era and lasted from annum, Mya to Mya (million years ago). Instead of being based on stratigraphy, these dates are defined Absolute dating, chronometrically. The later half of the period was an episode of intensive orogeny on virtually all continents. Two of the largest known impact events on Earth occurred during the Orosirian. Early in the period, 2023 Mya, a large asteroid collision created the Vredefort impact structure. The event that created the Sudbury Basin structure occurred near the end of the period, 1850 Mya. For the time period from about 2060 to 1780 Mya, an alternative period based on stratigraphy rather than chronometry, named the Columbian, was suggested in the geological timescale review 2012 edited by Gradstein et al., but , this has not yet been officially adopted by the IUGS. Paleogeography T ...
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Hemimastigophora
Hemimastigophora is a group of single-celled eukaryotic organisms with a single family, Spironemidae, first identified in 1988. Over the next 30 years, different authors proposed placing these organisms in various branches of the eukaryotes. In 2018 Lax ''et al''. reported the first genetic information for Spironemidae, and suggest that they are from an ancient lineage of eukaryotes which constitute a separate clade from all other eukaryotic kingdoms. It is potentially related to the Telonemia. History of classification Hemimastigophora was established in 1988 by Foissner ''et al''., as a new phylum with a single family, Spironemidae. Its placement on the eukaryote tree of life was unclear, but the authors suggested that the structure of its pellicle and cell nucleus indicated a close relationship with Euglenozoa. For 30 years after the description of the group, no genetic information was available. During that time, researchers proposed that it should be classified ...
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Ancyromonadida
Ancyromonadida or Planomonadida is a small group of biflagellated protists found in the soil and in aquatic habitats, where they feed on bacteria.Cavalier-Smith, T. (2013)Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa European journal of protistology, 49(2), 115-178. Includes freshwater or marine organisms, benthic, dorsoventrally compressed and with two unequal flagellae, each emerging from a separate pocket. The apical anterior flagellum can be very thin or end in the cell membrane, while the posterior flagellum is long and is inserted ventrally or laterally. The cell membrane is supported by a thin single layer teak and the mitochondrial crests are discoidal / flat. The group's placement is doubtful, as it seems to fall outside the five supergroups of eukaryotes. Cavalier-Smith considers that they constitute a basal group to Amoebozoa and Opisthokonta and places it together w ...
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Opimoda
The Scotokaryotes (Cavalier-Smith) is a proposed basal Neokaryote clade as sister of the Diaphoretickes. Basal Scotokaryote groupings are the Metamonads, the Malawimonas and the Podiata. In this phylogeny the Discoba are sometimes seen as paraphyletic and basal Eukaryotes. An alternative to the Unikont–Bikont division was suggested by Derelle ''et al.'' in 2015, where they proposed the acronyms Opimoda–Diphoda respectively, as substitutes to the older terms. The name Opimoda is formed from the letters (shown in capitals) of OPIsthokonta and aMOebozoa. In this phylogeny Discoba belongs to the Diphoda clade. Taxonomy A proposed cladogram is See also *Diphoda A bikont ("two flagella") is any of the eukaryotic organisms classified in the group Bikonta. Many single-celled members of the group, and the presumed ancestor, have two flagella. Enzymes Another shared trait of bikonts is the fusion of two ge ... References {{Taxonbar, from=Q21399998 Eukaryote unranked ...
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Stramenopiles
Stramenopile is a clade of organisms distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, along with Alveolata and Rhizaria. Members of the clade are referred to as 'stramenopiles'. Stramenopiles are eukaryotes; since they are neither fungi, animals, nor plants, they are classified as protists. Most stramenopiles are single-celled, but some are multicellular algae including some brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid heliozoa, and oomycetes. T ...
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Halvaria
Halvaria is a grouping that includes Alveolata and Heterokonta (Stramenopiles). Analyses in 2007 and 2008 revealed that the Stramenopiles and the Alveolata are related, and form a reduced chromalveolate clade. They group together with the Rhizaria (originally one of the six major eukaryote groups) to form a clade dubbed the SAR supergroup. A phylogenomic analysis from 2016 cast doubt on Halvaria, suggesting that Alveolata is the sister group to Rhizaria (making the ''R + A clade'') through new rhizarian sequence data, and that support for Halvaria might be an artifact of low taxon sampling as well as long branch attraction. However, later analyses from 2021 support Halvaria as a solid clade. See also *Chromalveolata *Rhizaria The Rhizaria are an ill-defined but species-rich supergroup of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus Paulinella in the phylum Cercozoa, they are all non-photosynthethic, but many foramini ... ...
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Rhizaria
The Rhizaria are an ill-defined but species-rich supergroup of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus Paulinella in the phylum Cercozoa, they are all non-photosynthethic, but many foraminifera and radiolaria have a symbiotic relationship with unicellular algae. A multicellular form, ''Guttulinopsis vulgaris'', a cellular slime mold, has also been described. This group was used by Cavalier-Smith in 2002, although the term "Rhizaria" had been long used for clades within the currently recognized taxon. Being described mainly from rDNA sequences, they vary considerably in form, having no clear morphological distinctive characters (synapomorphies), but for the most part they are amoeboids with filose, reticulose, or microtubule-supported pseudopods. In the absence of an apomorphy, the group is ill-defined, and its composition has been very fluid. Some Rhizaria possess mineral exoskeleton (thecae or loricas), which is in differ ...
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Telonemia
Telonemia is a phylum of microscopic eukaryote, single-celled organisms. They were formerly classified as protists until that kingdom fell out of general use, and are suggested to have evolutionary significance in being a possible transitional form between ecologically important heterotrophic and photosynthetic species among chromalveolates. One paper places them in the SAR supergroup. Phylogenomic analyses of 127 genes place Telonemia with Centroheliozoa in a group also consisting of cryptomonads and haptophytes (see Cryptomonads-haptophytes assemblage). Although they have been studied in primarily marine environments, they have also been found in freshwater. Shalchian-Tabrizi ''et al'' say that 18S rDNA sequences in the phylum formed two major groups, Group 1 and 2, including ''T. subtilis'' and ''T. antarcticum'' respectively, and that these were further sub-divided into several statistically supported clades of sequences with restricted geographic distribution. Species of Te ...
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TSAR
Tsar ( or ), also spelled ''czar'', ''tzar'', or ''csar'', is a title used by East Slavs, East and South Slavs, South Slavic monarchs. The term is derived from the Latin word ''Caesar (title), caesar'', which was intended to mean "emperor" in the European medieval sense of the term—a ruler with the same rank as a Roman emperor, holding it by the approval of another emperor or a supreme ecclesiastical official (the Pope or the Ecumenical Patriarch)—but was usually considered by western Europeans to be equivalent to "king". It lends its name to a system of government, tsarist autocracy or tsarism. "Tsar" and its variants were the official titles of the following states: * Bulgarian Empire (First Bulgarian Empire in 681–1018, Second Bulgarian Empire in 1185–1396), and also used in Kingdom of Bulgaria, Tsardom of Bulgaria, in 1908–1946 * Serbian Empire, in 1346–1371 * Tsardom of Russia, in 1547–1721 (replaced in 1721 by ''imperator'' in Russian Empire, but still re ...
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Haptista
Haptista is a proposed group of protists made up of centrohelids and haptophytes. Phylogenomic studies indicate that Haptista, together with ''Ancoracysta twista'', forms a sister clade to the SAR+Telonemia supergroup, but it may also be sister to the Cryptista (+Archaeplastida). It is thus one of the earliest diverging Diaphoretickes. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 and Ruggiero et al. 2015. * Subphylum Centroheliozoa Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987 eliozoa Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003** Class Centrohelea Kuhn 1926 stat. n. Cavalier-Smith 1993 entroplastiales; Centrohelina Hartmann 1913; Centroplasthelida Febvre-Chevalier 1984* Subphylum Haptophytina Cavalier-Smith 2015 (Haptophyta Hibberd 1976 sensu Ruggerio et al. 2015) ** Clade Rappemonada Kim et al. 2011 *** Class Rappephyceae Rappephyceae, or Rappemonads, are a small family of protists first described in 2011, of uncertain phylog ...
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