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Stramenopiles
Stramenopile is a clade of organisms distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, along with Alveolata and Rhizaria. Members of the clade are referred to as 'stramenopiles'. Stramenopiles are eukaryotes; since they are neither fungi, animals, nor plants, they are classified as protists. Most stramenopiles are single-celled, but some are multicellular algae including some brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid heliozoa, and oomycetes. T ...
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Heterokont
Heterokonts are a group of protists (formally referred to as Heterokonta, Heterokontae or Heterokontophyta). The group is a major line of eukaryotes. Most are algae, ranging from the giant multicellular kelp to the unicellular diatoms, which are a primary component of plankton. Other notable members of the Stramenopiles include the (generally) parasitic oomycetes, including ''Phytophthora'', which caused the Great Famine of Ireland, and ''Pythium'', which causes seed rot and damping off. The name "heterokont" refers to the type of motile life cycle stage, in which the flagellated cells possess two differently arranged flagella (see zoospore). History In 1899, Alexander Luther created the term "Heterokontae" for some algae with unequal flagella, today called Xanthophyceae. Later, some authors (e.g., Copeland, 1956) included other groups in Heterokonta, expanding the name's sense. The term continues to be applied in different ways, leading to Heterokontophyta being applie ...
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Heterokont
Heterokonts are a group of protists (formally referred to as Heterokonta, Heterokontae or Heterokontophyta). The group is a major line of eukaryotes. Most are algae, ranging from the giant multicellular kelp to the unicellular diatoms, which are a primary component of plankton. Other notable members of the Stramenopiles include the (generally) parasitic oomycetes, including ''Phytophthora'', which caused the Great Famine of Ireland, and ''Pythium'', which causes seed rot and damping off. The name "heterokont" refers to the type of motile life cycle stage, in which the flagellated cells possess two differently arranged flagella (see zoospore). History In 1899, Alexander Luther created the term "Heterokontae" for some algae with unequal flagella, today called Xanthophyceae. Later, some authors (e.g., Copeland, 1956) included other groups in Heterokonta, expanding the name's sense. The term continues to be applied in different ways, leading to Heterokontophyta being applie ...
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Eukaryotes
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Eu ...
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Rhizaria
The Rhizaria are an ill-defined but species-rich supergroup of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus Paulinella in the phylum Cercozoa, they are all non-photosynthethic, but many foraminifera and radiolaria have a symbiotic relationship with unicellular algae. A multicellular form, ''Guttulinopsis vulgaris'', a cellular slime mold, has also been described. This group was used by Cavalier-Smith in 2002, although the term "Rhizaria" had been long used for clades within the currently recognized taxon. Being described mainly from rDNA sequences, they vary considerably in form, having no clear morphological distinctive characters (synapomorphies), but for the most part they are amoeboids with filose, reticulose, or microtubule-supported pseudopods. In the absence of an apomorphy, the group is ill-defined, and its composition has been very fluid. Some Rhizaria possess mineral exoskeleton (thecae or loricas), which is in differ ...
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Hyphochytrid
Hyphochytrids are eukaryotic organisms in the group of Stramenopiles (Heterokonta). Characteristics They are distinguished by an anterior tinsel flagellum on their zoospores. Also they have a rhizoidal or hypha-like vegetative system (hence the prefix "Hypho-"). Classification This group may be put alternatively at the phylum, class, subclass or order level, being referred to as Hyphochytriomycota, Hyphochytriomycetes (or Hyphochytrea), Hyphochytriomycetidae (or Hyphochytridae) and Hyphochytriales, respectively. The variants Hyphochytri''di''omycota and Hyphochytri''di''omycetes are also sometimes used, presumably by analogy to the Chytridiomycetes, or due to the perpetuation of a typographical error. However, the stem is Hyphochytri- (from '' Hyphochytrium'') and not Hyphochytridi- (from ''Chytridium''). In the past the classes Hyphochytridiomycetes, Oomycetes and Chytridiomycetes were grouped together in the now obsolete taxon Mastigomycotina as fungi with flagellate spores o ...
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Flagellate
A flagellate is a cell or organism with one or more whip-like appendages called flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryotes and their means of motion. The term presently does not imply any specific relationship or classification of the organisms that possess flagella. However, the term "flagellate" is included in other terms (such as "dinoflagellate" and "choanoflagellata") which are more formally characterized. Form and behavior Flagella in eukaryotes are supported by microtubules in a characteristic arrangement, with nine fused pairs surrounding two central singlets. These arise from a basal body. In some flagellates, flagella direct food into a cytostome or mouth, where food is ingested. Flagella often support hairs, called mastigonemes, or contain rods. Their ultrastructure plays an important role in classifying eukaryotes. Among protoctists and microscopic anima ...
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Xanthophyceae
Yellow-green algae or the Xanthophyceae (xanthophytes) are an important group of heterokont algae. Most live in fresh water, but some are found in marine and soil habitats. They vary from single-celled flagellates to simple colonial and filamentous forms. Xanthophyte chloroplasts contain the photosynthetic pigments chlorophyll ''a'', chlorophyll ''c'', β-carotene, and the carotenoid diadinoxanthin. Unlike other heterokonts, their chloroplasts do not contain fucoxanthin, which accounts for their lighter colour. Their storage polysaccharide is chrysolaminarin. Xanthophyte cell walls are produced of cellulose and hemicellulose. They appear to be the closest relatives of the brown algae. Classifications The species now placed in the Xanthophyceae were formerly included in the Chlorophyceae. In 1899, Lüther created the group Heterokontae for green algae with unequal flagella. Pascher (1914) included the Heterokontae in the Chrysophyta. In 1930, Allorge renamed the group as Xant ...
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Protists
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exclusion of other eukaryotes means that protists do not form a natural group, or clade. Therefore, some protists may be more closely related to animals, plants, or fungi than they are to other protists. However, like the groups ''algae'', ''invertebrates'', and ''protozoans'', the biological category ''protist'' is used for convenience. Others classify any unicellular eukaryotic microorganism as a protist. The study of protists is termed protistology. History The classification of a third kingdom separate from animals and plants was first proposed by John Hogg in 1860 as the kingdom Protoctista; in 1866 Ernst Haeckel also proposed a third kingdom Protista as "the kingdom of primitive forms". Originally these also included prokaryotes, but ...
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Brown Algae
Brown algae (singular: alga), comprising the class Phaeophyceae, are a large group of multicellular algae, including many seaweeds located in colder waters within the Northern Hemisphere. Brown algae are the major seaweeds of the temperate and polar regions. They are dominant on rocky shores throughout cooler areas of the world. Most brown algae live in marine environments, where they play an important role both as food and as a potential habitat. For instance, ''Macrocystis'', a kelp of the order Laminariales, may reach in length and forms prominent underwater kelp forests. Kelp forests like these contain a high level of biodiversity. Another example is ''Sargassum'', which creates unique floating mats of seaweed in the tropical waters of the Sargasso Sea that serve as the habitats for many species. Many brown algae, such as members of the order Fucales, commonly grow along rocky seashores. Some members of the class, such as kelps, are used by humans as food. Between 1,500 and ...
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Chrysophyte
The Chrysophyceae, usually called chrysophytes, chrysomonads, golden-brown algae or golden algae are a large group of algae, found mostly in freshwater. Golden algae is also commonly used to refer to a single species, ''Prymnesium parvum'', which causes fish kills. The Chrysophyceae should not be confused with the Chrysophyta, which is a more ambiguous taxon. Although "chrysophytes" is the anglicization of "Chrysophyta", it generally refers to the Chrysophyceae. Members Originally they were taken to include all such forms of the diatoms and multicellular brown algae, but since then they have been divided into several different groups (e.g., Haptophyceae, Synurophyceae) based on pigmentation and cell structure. Some heterotrophic flagellates as the bicosoecids and choanoflagellates were sometimes seen as related to golden algae too. They are now usually restricted to a core group of closely related forms, distinguished primarily by the structure of the flagella in motile cells, ...
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Heliozoa
Heliozoa, commonly known as sun-animalcules, are microbial eukaryotes (protists) with stiff arms ( axopodia) radiating from their spherical bodies, which are responsible for their common name. The axopodia are microtubule-supported projections from the amoeboid cell body, and are variously used for capturing food, sensation, movement, and attachment. They are similar to Radiolaria, but they are distinguished from them by lacking central capsules and other complex skeletal elements, although some produce simple scales and spines. They may be found in both freshwater and marine environments. Classification Originally the heliozoa were treated together as a formal taxon Heliozoa or Heliozoea, with the rank of class or phylum, but it has been realised that they are polyphyletic, as the various orders show notable differences and are no longer believed to be descended from a single common ancestor. Instead, "heliozoa" is regarded as a descriptive term applying to various lines of proti ...
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Choanoflagellate
The choanoflagellates are a group of free-living unicellular and colonial flagellate eukaryotes considered to be the closest living relatives of the animals. Choanoflagellates are collared flagellates, having a funnel shaped collar of interconnected microvilli at the base of a flagellum. Choanoflagellates are capable of both asexual and sexual reproduction. They have a distinctive cell morphology characterized by an ovoid or spherical cell body 3–10  µm in diameter with a single apical flagellum surrounded by a collar of 30–40 microvilli (see figure). Movement of the flagellum creates water currents that can propel free-swimming choanoflagellates through the water column and trap bacteria and detritus against the collar of microvilli, where these foodstuffs are engulfed. This feeding provides a critical link within the global carbon cycle, linking trophic levels. In addition to their critical ecological roles, choanoflagellates are of particular interest to evolutionary ...
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