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Euascomycetes
Pezizomycotina make up most of the Ascomycota fungi and include most lichenized fungi too. Pezizomycotina contains the filamentous ascomycetes and is a subdivision of the Ascomycota (fungi that form their spores in a sac-like ''ascus''). It is more or less synonymous with the older taxon Euascomycota. These fungi reproduce by fission rather than budding and this subdivision includes almost all the ascus fungi that have fruiting bodies visible to the naked eye (exception: genus ''Neolecta'', which belongs to the Taphrinomycotina). See the taxobox for a list of the classes that make up the Pezizomycotina. The old class Loculoascomycetes (consisting of all the bitunicate Ascomycota) has been replaced by the two classes Eurotiomycetes and Dothideomycetes. The rest of the Pezizomycotina also include the previously defined hymenial groups Discomycetes (now Leotiomycetes) and Pyrenomycetes (Sordariomycetes). Some important groups in Pezizomycotina include: Pezizomycetes (the operculate ...
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Ascomycota
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the " ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as ''Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomyce ...
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Neolecta
''Neolecta'' is a genus of ascomycetous fungi that have fruiting bodies in the shape of unbranched to lobed bright yellowish, orangish to pale yellow-green colored, club-shaped, smooth, fleshy columns up to about 7 cm tall. The species share the English designation "Earth tongues" along with some better-known fungi (e.g. ''Geoglossum'', ''Microglossum'') with a similar general form, but in fact they are only distantly related. ''Neolecta'' is the only genus belonging to the family Neolectaceae, which is the only family belonging to the order Neolectales. ''Neolectales'', in turn, is the only order belonging to the class Neolectomycetes, which belongs to the subdivision Taphrinomycotina of the Ascomycota. ''Neolecta'' is found in Asia, North America, Northern Europe and southern Brazil. The species all live in association with trees, and at least one, ''N. vitellina'', grows from rootlets of its host, but it is not known whether the fungus is parasitic, saprotrophic, or mutu ...
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Rhynie Chert
The Rhynie chert is a Lower Devonian sedimentary deposit exhibiting extraordinary fossil detail or completeness (a Lagerstätte). It is exposed near the village of Rhynie, Aberdeenshire, Scotland; a second unit, the Windyfield chert, is located some 700 m away. The Rhynie chert contains exceptionally preserved plant, fungus, lichen and animal material preserved in place by an overlying volcanic deposit. The bulk of the Devonian fossil bed consists of primitive plants (which had water-conducting cells and sporangia, but no true leaves), along with arthropods, lichens, algae and fungi. This fossil bed is remarkable for two reasons. First, the age of the site ( Pragian, Early Devonian, formed about ) places it at an early stage in the colonisation of land. Second, these cherts are famous for their exceptional state of ultrastructural preservation, with individual cell walls easily visible in polished specimens. Stomata have been counted and lignin remnants detected in the ...
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Early Devonian
The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first Ammonoidea, ammonoids appeared, descending from Bactritida, bactritoid Nautiloidea, nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatite, Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era. References

{{Geological history Early Devonian, Geological epochs Devonian geochronology, *01 ...
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Paleopyrenomycites
''Paleopyrenomycites'' is a Devonian genus of fungus of uncertain phylogenetic affinity within the Pezizomycotina total group, known from the Rhynie chert The Rhynie chert is a Lower Devonian sedimentary deposit exhibiting extraordinary fossil detail or completeness (a Lagerstätte). It is exposed near the village of Rhynie, Aberdeenshire, Scotland; a second unit, the Windyfield chert, is locate .... {{Taxonbar, from=Q16987177 Ascomycota Prehistoric fungi Pragian life ...
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Laboulbeniomycetes
The Laboulbeniomycetes are a unique group of fungi that are obligatorily associated with arthropods, either as external parasites ( Herpomycetales and Laboulbeniales) or for dispersal ( Pyxidiophorales). Herpomycetales and Laboulbeniales fungi are minute; their fruiting bodies, referred to as ''thalli'', commonly measure less than one millimeter. They live on the antennae, the mouthparts or other body regions of their arthropod hosts. Although several species of Laboulbeniomycetes have more or less extensive, root-like hyphal systems (haustoria In botany and mycology, a haustorium (plural haustoria) is a rootlike structure that grows into or around another structure to absorb water or nutrients. For example, in mistletoe or members of the broomrape family, the structure penetrates th ...) inside their hosts, as a group these fungi are relatively harmless to their hosts. These fungi occur usually only on adult hosts; apparently immature arthropods eliminate them during e ...
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Sordariomycetes
Sordariomycetes is a class of fungi in the subdivision Pezizomycotina (Ascomycota), consisting of 28 orders, 90 families, 1344 genera. Sordariomycetes is from the Latin sordes (filth) because some species grow in animal feces, though growth habits vary widely across the class. Sordariomycetes generally produce their asci in perithecial fruiting bodies. Sordariomycetes are also known as Pyrenomycetes, from the Greek πυρἠν - 'the stone of a fruit' - because of the usually somewhat tough texture of their tissue. Sordariomycetes possess great variability in morphology, growth form, and habitat. Most have perithecial (flask-shaped) fruiting bodies, but ascomata can be less frequently cleistothecial (like in the genera '' Anixiella'', ''Apodus'', '' Boothiella'', ''Thielavia'', '' Zopfiella''),. Fruiting bodies may be solitary or gregarious, superficial, or immersed within stromata or tissues of the substrates and can be light to bright or black. Members of this group can grow ...
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Leotiomycetes
The Leotiomycetes are a class of ascomycete fungi. Many of them cause serious plant diseases. Systematics The class Leotiomycetes contains numerous species with an anamorph placed within the ''fungi imperfecti'' (deuteromycota), that have only recently found their place in the phylogenetic system. The older classifications placed Leotiomycetes into the Discomycetes clade ( inoperculate Discomycetes). Molecular studies have recently shed some new light to the still obscure systematics. Most scholars consider Leotiomycetes a sister taxon to Sordariomycetes in the phylogenetic tree of Pezizomycotina. Its division into subclasses have received strong support by the molecular data, but the overall monophyly of Leotiomycetes is dubious. The order Lichinodiales and family Lichinodiaceae, newly circumscribed in 2019 to contain the genus cyanolichen genus '' Lichinodium'', is the first known group of lichen-forming fungi in the Leotiomycetes. Characteristics *Most ''Leotiomycetes'' g ...
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Dothideomycetes
Dothideomycetes is the largest and most diverse class of ascomycete fungi. It comprises 11 orders 90 families, 1300 genera and over 19,000 known species. Traditionally, most of its members were included in the loculoascomycetes, which is not part of the currently accepted classification. This indicates that several traditional morphological features in the class are not unique and DNA sequence comparisons are important to define the class. The designation loculoascomycetes was first proposed for all fungi which have ascolocular development. This type of development refers to the way in which the sexual structure, bearing the sexual spores (ascospores) forms. Dothideomycetes mostly produce flask-like structures referred to as pseudothecia, although other shape variations do exist (e.g. see structures found in Hysteriales). During ascolocular development pockets (locules) form first within the vegetative cells of the fungus and then all the subsequent structures form. These includ ...
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Eurotiomycetes
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. It contains most of the fungi previously known morphologically as "Plectomycetes".


Systematics and phylogeny


Internal relationships

The class Eurotiomycetes was circumscription (taxonomy), circumscribed in 1997 by Sweden, Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time ...

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Taphrinomycotina
The Taphrinomycotina are one of three subdivisions constituting the Ascomycota (fungi that form their spores in a sac-like ascus) and is more or less synonymous with the slightly older invalid name Archiascomycetes (sometimes spelled Archaeascomycetes; archea = ancient). Recent molecular studies suggest that the group is monophyletic and basal to the rest of the Ascomycota. The major taxa are Schizosaccharomycetes, Taphrinomycetes, Neolectomycetes, and Pneumocystis. The Schizosaccharomycetes are the yeasts (e.g. ''Schizosaccharomyces'') that reproduce by fission rather than budding, unlike most other yeasts, many of which are in the subdivision Saccharomycotina. The Taphrinomycetes are dimorphic plant parasites (e.g. ''Taphrina'') with both a yeast state and a filamentous (hyphal) state in infected plants. They characteristically infect leaves, catkins, and branches, not roots. Taphrinomycetes form asci but no ascomata. The Neolectomycetes are species in a single genus, ''N ...
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Aleuria Aurantia
''Aleuria aurantia'' (orange peel fungus) is a widespread ascomycete fungus in the order Pezizales. The brilliant orange, cup-shaped ascocarps often resemble orange peels strewn on the ground, giving this species its common name. Taxonomy Christiaan Hendrik Persoon described the orange peel as ''Peziza aurantia'' in 1800. The specific epithet is the Latin word ''aurantia'' "orange". Karl Wilhelm Gottlieb Leopold Fuckel placed it the genus '' Aleuria'' in 1870. Description The orange fruiting body is 2–10 cm wide, cup-shaped, often misshapen due to crowding from other fruiting bodies. The spores are colorless and scatter in visible clouds when disturbed. It is generally regarded as edible, though difficult to collect intact and not necessarily choice, with no particularly notable North American lookalikes. In Europe, the orange peel may be confused with species of '' Otidea'' or '' Caloscypha'' which are poisonous or of unknown edibility. Similar species include ''Caloscypha ...
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