Echimyidae Phylogeny
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Echimyidae Phylogeny
Echimyidae is the family of neotropical spiny rats and their fossil relatives. This is the most species-rich family of hystricognath rodents. It is probably also the most ecologically diverse, with members ranging from fully arboreal to terrestrial to fossorial to semiaquatic habits. They presently exist mainly in South America; three members of the family also range into Central America, and the hutias are found in the Greater Antilles in the Caribbean. Species of the extinct subfamily Heteropsomyinae formerly lived on Cuba, Hispaniola, and Puerto Rico in the Antilles, probably until the arrival of Europeans. Characteristics In general form, most spiny rats resemble rats, although they are more closely related to guinea pigs and chinchillas. Most species have stiff, pointed hairs, or spines, that presumably serve for protection from predators. Many echimyids can break off their tails when attacked. This action may confuse predators long enough for the spiny rat to esc ...
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Coypu
The nutria (''Myocastor coypus''), also known as the coypu, is a large, herbivorous, semiaquatic rodent. Classified for a long time as the only member of the family Myocastoridae, ''Myocastor'' is now included within Echimyidae, the family of the spiny rats. The nutria lives in burrows alongside stretches of water, and feeds on river plant stems. Originally native to subtropical and temperate South America, it has since been introduced to North America, Europe, Asia, and Africa, primarily by fur farmers. Although it is still hunted and trapped for its fur in some regions, its destructive burrowing and feeding habits often bring it into conflict with humans, and it is considered an invasive species. Nutria also transmit various diseases to humans and animals mainly through water contamination. Etymology The genus name ''Myocastor'' derives from the two Ancient Greek words (), meaning "rat, mouse", and (), meaning "beaver". Literally, therefore, the name ''Myocastor'' means ...
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Cuba
Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbean Sea, Gulf of Mexico, and Atlantic Ocean meet. Cuba is located east of the Yucatán Peninsula (Mexico), south of both the American state of Florida and the Bahamas, west of Hispaniola ( Haiti/Dominican Republic), and north of both Jamaica and the Cayman Islands. Havana is the largest city and capital; other major cities include Santiago de Cuba and Camagüey. The official area of the Republic of Cuba is (without the territorial waters) but a total of 350,730 km² (135,418 sq mi) including the exclusive economic zone. Cuba is the second-most populous country in the Caribbean after Haiti, with over 11 million inhabitants. The territory that is now Cuba was inhabited by the Ciboney people from the 4th millennium BC with the Gua ...
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Caribbean
The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Sea and the North Atlantic Ocean) and the surrounding coasts. The region is southeast of the Gulf of Mexico and the North American mainland, east of Central America, and north of South America. Situated largely on the Caribbean Plate, the region has more than 700 islands, islets, reefs and cays (see the list of Caribbean islands). Island arcs delineate the eastern and northern edges of the Caribbean Sea: The Greater Antilles and the Lucayan Archipelago on the north and the Lesser Antilles and the on the south and east (which includes the Leeward Antilles). They form the West Indies with the nearby Lucayan Archipelago (the Bahamas and Turks and Caicos Islands), which are considered to be part of the Caribbean despite not bordering the Caribbe ...
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Greater Antilles
The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, and the Cayman Islands. Six island states share the region of the Greater Antilles, with Haiti and the Dominican Republic sharing the island of Hispaniola. Together with the Lesser Antilles, they make up the Antilles. While most of the Greater Antilles consists of independent countries, Puerto Rico is an unincorporated territory of the United States, while the Cayman Islands are a British Overseas Territory. The largest island by area is Cuba, which extends to the western end of the island group. Puerto Rico lies on the eastern end, and the island of Hispaniola, the largest island by population, is located in the middle. Jamaica lies to the south of Cuba, while the Cayman Islands are located to the west. The state of Florida is the closes ...
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Hutia
Hutias (known in Spanish as jutía) are moderately large cavy-like rodents of the subfamily Capromyinae that inhabit the Caribbean islands, with most species restricted to Cuba and Hispaniola. Twenty species of hutia have been identified, but at least half are extinct. Only Desmarest's hutia and the prehensile-tailed hutia remain common and widespread; all other extant species are considered threatened by the IUCN. The extinct giant hutias of the family Heptaxodontidae also inhabited the Caribbean, but are not thought to be closely related, with the giant hutias belonging in the superfamily Chinchilloidea. Description Most species have a head-and-body length that ranges from and weigh less than , but Desmarest's hutia has a head-and-body length of and weighs . They resemble the coypu in some respects. Tails are present, varying from vestiges to prehensile. They have stout bodies and large heads. Most species are herbivorous, though some consume small animals. Instead of burro ...
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Central America
Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. Central America consists of eight countries: Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, and Panama. Within Central America is the Mesoamerican biodiversity hotspot, which extends from northern Guatemala to central Panama. Due to the presence of several active geologic faults and the Central America Volcanic Arc, there is a high amount of seismic activity in the region, such as volcanic eruptions and earthquakes which has resulted in death, injury, and property damage. In the pre-Columbian era, Central America was inhabited by the indigenous peoples of Mesoamerica to the north and west and the Isthmo-Colombian peoples to the south and east. Following the Spanish expedition of Christopher Columbus' ...
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South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southern subregion of a single continent called America. South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean; North America and the Caribbean Sea lie to the northwest. The continent generally includes twelve sovereign states: Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Guyana, Paraguay, Peru, Suriname, Uruguay, and Venezuela; two dependent territories: the Falkland Islands and South Georgia and the South Sandwich Islands; and one internal territory: French Guiana. In addition, the ABC islands of the Kingdom of the Netherlands, Ascension Island (dependency of Saint Helena, Ascension and Tristan da Cunha, a British Overseas Territory), Bouvet Island ( dependency of Norway), Pa ...
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Semiaquatic
In biology, semiaquatic can refer to various types of animals that spend part of their time in water, or plants that naturally grow partially submerged in water. Examples are given below. Semiaquatic animals Semiaquatic animals include: * Vertebrates ** Amphibious fish; also several types of normally fully aquatic fish such as the grunion and plainfin midshipman that spawn in the intertidal zone ** Some amphibians such as newts and salamanders, and some frogs such as fire-bellied toads and wood frogs. ** Some reptiles such as crocodilians, turtles, water snakes and marine iguanas. ** Penguins. ** Some rodents such as beavers, muskrats and capybaras. **Some insectivorous mammals such as desmans, water shrews and platypuses. ** Some carnivoran mammals, including seals, polar bears and otters. ** Hippopotamuses. * Semiterrestrial echinoderms of the intertidal zone, such as the "cliff-clinging" sea urchin ''Colobocentrotus atratus'' and the starfish '' Pisaster ochraceus'' ...
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Fossorial
A fossorial () animal is one adapted to digging which lives primarily, but not solely, underground. Some examples are badgers, naked mole-rats, clams, meerkats, and mole salamanders, as well as many beetles, wasps, and bees. Prehistoric evidence The physical adaptation of fossoriality is widely accepted as being widespread among many prehistoric phyla and taxa, such as bacteria and early eukaryotes. Furthermore, fossoriality has evolved independently multiple times, even within a single family. Fossorial animals appeared simultaneously with the colonization of land by arthropods in the late Ordovician period (over 440 million years ago). Other notable early burrowers include ''Eocaecilia'' and possibly ''Dinilysia''. The oldest example of burrowing in synapsids, the lineage which includes modern mammals and their ancestors, is a cynodont, ''Thrinaxodon liorhinus'', found in the Karoo of South Africa, estimated to be 251 million years old. Evidence shows that this ...
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Terrestrial Animal
Terrestrial animals are animals that live predominantly or entirely on land (e.g. cats, dogs, ants, spiders), as compared with aquatic animals, which live predominantly or entirely in the water (e.g. fish, lobsters, octopuses), and amphibians, which rely on a combination of aquatic and terrestrial habitats (e.g. frogs and newts). Some groups of insects are terrestrial, such as ants, butterflies, earwigs, cockroaches, grasshoppers and many others, while other groups are partially aquatic, such as mosquitoes and dragonflies, which pass their larval stages in water. Terrestrial animals tend to be more developed and intelligent than aquatic animals. Terrestrial classes The term "terrestrial" is typically applied to species that live primarily on the ground, in contrast to arboreal species, which live primarily in trees. There are other less common terms that apply to specific groups of terrestrial animals: *Saxicolous creatures are rock dwelling. "Saxicolous" is derived from t ...
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Arboreal
Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose numerous mechanical challenges to animals moving through them and lead to a variety of anatomical, behavioral and ecological consequences as well as variations throughout different species.Cartmill, M. (1985). Climbing. In ''Functional Vertebrate Morphology'', eds. M. Hildebrand D. M. Bramble K. F. Liem and D. B. Wake, pp. 73–88. Cambridge: Belknap Press. Furthermore, many of these same principles may be applied to climbing without trees, such as on rock piles or mountains. Some animals are exclusively arboreal in habitat, such as the tree snail. Biomechanics Arboreal habitats pose numerous mechanical challenges to animals moving in them, which have been solved in diverse ways. These challenges include moving on n ...
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