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Dinogorgon Rubidgei
''Dinogorgon'' is a genus of gorgonopsid from the Late Permian of South Africa and Tanzania. The generic name ''Dinogorgon'' is derived from Greek, meaning "terrible gorgon", while its species name ''rubidgei'' is taken from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. The type species of the genus is ''D. rubidgei''. ''Dinogorgon'' is part of the gorgonopsian subfamily Rubidgeinae, a derived group of large-bodied gorgonopsians restricted to the Late Permian (Lopingian). The Rubidgeinae subfamily first appeared in the ''Tropidostoma'' Assemblage Zone, and reached their highest diversity in the ''Cistecephalus'' and ''Daptocephalus'' assemblage zones of the Beaufort Group in South Africa. History of discovery The type species of ''Dinogorgon rubidgei'' was discovered on Wellwood farm, a farm owned by the grandfather of Bruce Rubidge, Sidney H. Rubidge, outside of G ...
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Late Permian
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * Late (album), ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his ''Pocketwatch (album), Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * Late (song), "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from ''Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * Late (The Handmaid's Tale), "Late" (''The Handmaid's Tale''), a television episode * LaTe, Laivateollisuus, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also

* * * ''Lates'', a genus of fish in the lates perch family * Later (other) ...
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Derived (phylogenetics)
In phylogenetics, a primitive (or ancestral) character, trait, or feature of a lineage or taxon is one that is inherited from the common ancestor of a clade (or clade group) and has undergone little change since. Conversely, a trait that appears ''within'' the clade group (that is, is present in any subgroup within the clade but not all) is called advanced or derived. A clade is a group of organisms that consists of a common ancestor and all its lineal descendants. A primitive trait is the original condition of that trait in the common ancestor; advanced indicates a notable change from the original condition. These terms in biology contain no judgement about the sophistication, superiority, value or adaptiveness of the named trait. "Primitive" in biology means only that the character appeared first in the common ancestor of a clade group and has been passed on largely intact to more recent members of the clade. "Advanced" means the character has evolved within a later subgroup ...
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Clelandina
''Clelandina'' is an extinct genus of rubidgeine gorgonopsian from the Late Permian of ''Cistecephalus'' Assemblage Zone of South Africa. It was first named by Broom in 1948. The type and only species is ''C. rubidgei''. It is relatively rare, with only four known specimens. Description ''Clelandina rubidgei'' has an extraordinarily small sclerotic ring relative to the size of its orbit, which implies that it was diurnal. It is the only rubidgeine with a preserved sclerotic ring, so it is unknown whether this trait was shared by other members of the subfamily. Like all rubidgeines, it was relatively large, with a skull up to 36 cm long. It had reduced dentition, with the teeth posterior to the canines being absent and replaced with a bony ridge. The skull has heavily pachyostosed, with massive rugose bosses. Classification ''Clelandina'' shares many characteristics with the contemporary ''Rubidgea'', and is currently recognized as the sister taxon of this genus. Toge ...
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Aelurognathus
''Aelurognathus'' is an extinct genus of gorgonopsian therapsids from the Permian of South Africa. Discovery The type species is ''Aelurognathus tigriceps'', originally named ''Scymnognathus tigriceps'' by South African paleontologists Robert Broom and Sydney H. Haughton in 1913, and later assigned to the new genus ''Aelurognathus'' by Haughton in 1924. ''Scymnognathus parringtoni'' von Huene, 1950, previously assigned to ''Aelurognathus'', is now classified as a species of ''Sauroctonus''. ''Aelurognathus nyasaensis'' Haughton, 1926 is not referable to the genus.Kammerer CF. (2016) Systematics of the Rubidgeinae (Therapsida: Gorgonopsia) PeerJ 4:e1608 https://doi.org/10.7717/peerj.1608 Palaeobiology A broken tooth beside the skeleton of a dicynodont from the ''Tropidostoma'' Assemblage Zone has been attributed to ''Aelurognathus'', indicating that it scavenged. The bones of the back of the skeleton are the most scattered, suggesting that the ''Aelurognathus'' individuals ...
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Reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians (tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, historically combined with that of modern amphibians, is called herpetology. The earliest known proto-reptiles originated around ...
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Rubidgea
''Rubidgea'' is a genus of Gorgonopsia, gorgonopsid from the upper Permian of South Africa and Tanzania, containing the species ''Rubidgea atrox''. The generic name ''Rubidgea'' is sometimes believed to be derived from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Supergroup, Karoo Basin. However, this generic name was actually erected in honor of Rubidge's paternal grandfather, Sydney Rubidge, who was a renowned fossil hunter. Its species name ''atrox'' is derived from Latin, meaning “fierce, savage, terrible”. ''Rubidgea'' is part of the gorgonopsian subfamily Rubidgeinae, a Derived (phylogenetics), derived group of large-bodied gorgonopsians restricted to the Late Permian (Lopingian). The subfamily Rubidgeinae first appeared in the Tropidostoma Assemblage Zone, ''Tropidostoma'' Assemblage Zone. They reached their highest diversity in the ''Cistecephalus Assemblage Zone, ...
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Rubidgeinae
Rubidgeinae is an extinct subfamily of gorgonopsid therapsids known only from Africa. They were among the largest gorgonopsians, and their fossils are common in the Cistecephalus and Daptocephalus assemblage zones of the Karoo Basin. They lived during the Late Permian, and became extinct at the end of the Permian. Description Rubidgeines were large, quadrupedal carnivores of the family Gorgonopsidae. Their largest teeth are their upper canines, which were blade-like and had well-developed serrations. Their postcanine teeth were small and conical, but were also frequently serrated. Tooth replacement was rapid relative to basal therocephalians. Rubidgeines can be distinguished from other gorgonopsians by the absence of a blade-like parasphenoid bone and reduced or absent preparietal bone. The jugal bone, while narrow in most gorgonopsians, was often broadly expanded in rubidgeines. The largest rubidgeins were ''Dinogorgon'' and ''Rubidgea''. Paleobiology Rubidgeines were among ...
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James Kitching
James William Kitching (6 February 1922 – 24 December 2003) was a South African vertebrate palaeontologist and regarded as one of the world’s greatest fossil finders. Career His work in the southern hemisphere, including Antarctica, led to the establishment of one of the world's finest fossil collections, housed at the Bernard Price Institute for Palaeontological Research (BPI) in Johannesburg.University of the Witwatersrand, Johannesburg, Bernard Price Institute
He contributed greatly to the palaeontology of southern

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Graaff-Reinet
Graaff-Reinet is a town in the Eastern Cape Province of South Africa. It is the oldest town in the province. It is also the sixth-oldest town in South Africa, after Cape Town, Stellenbosch, Simon's Town, Paarl and Swellendam. The town was the centre of a short-lived republic in the late 18th century. The town was a starting point for Great Trek groups led by Gerrit Maritz and Piet Retief and furnished large numbers of the Voortrekkers in 1835–1842. Graaff-Reinet is home to more national monuments than any other town or city in South Africa. It is also known for being a flourishing market for agricultural produce, noted for its mohair industry, and sheep and ostrich farming. History Graaff-Reinet was established by the Dutch East India Company in 1786, after Cape Town in 1652, Stellenbosch in 1679, Paarl in 1687 and Swellendam in 1745. The town is named after then-governor of the Cape Colony, Cornelis Jacob van de Graaff, and his wife. The town was originally established ...
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Beaufort Group
The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlies the Stormberg Group. Based on stratigraphic position, lithostratigraphic and biostratigraphic correlations, palynological analyses, and other means of geological dating, the Beaufort Group rocks are considered to range between Middle Permian (Wordian) to Early Triassic (Anisian) in age. Background During the period when sedimentation of the Beaufort Group rocks took place, the Ecca sea had retreated to the northeastern Karoo Basin. All sediment deposition at this time took place in a terrestrial, although in a predominantly fluvial or alluvial environment that was seasonally arid. This environment covered a vast area and deposition was influenced by a retroarc foreland basin. This foreland system was caused by crustal uplift (oro ...
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Daptocephalus Assemblage Zone
The ''Daptocephalus'' Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa.Rubidge, B. S. (1995). Biostratigraphy of the Beaufort Group (Karoo Supergroup). ''Biostratigraphic series''. This biozone has outcrops located in the upper Teekloof Formation west of 24°E, the majority of the Balfour Formation east of 24°E, and the Normandien Formation in the north. It has numerous localities which are spread out from Colesberg in the Northern Cape, Graaff-Reniet to Mthatha in the Eastern Cape, and from Bloemfontein to Harrismith in the Free State. The ''Daptocephalus'' Assemblage Zone is one of eight biozones found in the Beaufort Group and is considered Late Permian (Lopingian) in age. Its contact with the overlying ''Lystrosaurus'' Assemblage Zone marks the Permian-Triassic boundary. Previously known as the ''Dicynodo ...
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