Diaphragmus
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Diaphragmus
''Diaphragmus'' is an extinct genus of brachiopod belonging to the order Productida and family Linoproductidae. Specimens have been found in Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ... beds in North America. Classification Leighton & Maples (2002) conducted multiple phylogenetic analyses which are strongly in agreement that the four subfamilies Buxtoniinae, Echinoconchinae, Pustulinae and Juresaniinae form the family Echinoconchidae. The results of their phylogenetic analyses are displayed in the cladogram below: Species *''D. elegans'' Norwood and Pratten *''D. nivosus'' Gordon 1975 References Paleozoic animals Productida {{Brachiopod-stub ...
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Antiquatonia
''Antiquatonia'' is an extinct genus of brachiopod belonging to the order Productida and family Productidae. Specimens have been found in Carboniferous beds across many continents, suggesting the genus had a cosmopolitan distribution. Species level taxonomy of ''Antiquatonia'' is in need of revision. Description Like other members of Productinae, ''Antiquatonia'' had a deep corpus cavity, a geniculate profile with long trails, and well-developed marginal structures such as ear baffles. This genus had thick, halteroid ventral spines. The ears were flanked by a ridge of ventral spines, with complimentary internal lateral ridges. Classification Muir-Wood & Cooper (1960) placed ''Antiquatonia'' in the subfamily Dictyoclostinae, though later analyses would find it to belong in Productinae instead. Leighton & Maples (2002) conducted multiple phylogenetic analyses which are strongly in agreement that ''Antiquatonia'' forms a clade with ''Diaphragmus ''Diaphragmus'' is an extinc ...
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Pulchratia
''Pulchratia'' is an extinct genus of brachiopods which lived in marine habitats during the Upper Carboniferous period. Its fossils have been found in North America. Description Like other echinoconchids, ''Pulchratia'' had recumbent spines arranged in irregular concentric bands, with the spines of this genus being mostly uniform in size and not having strong bases. It also has long lateral ridges which diverge from the hinge line. In these conditions it differs from the related '' Parajuresania'', which has thicker posterior spines than anterior spines, and short cardinal ridges that parallel the hinge line, but otherwise the two genera are very similar. The buttress plates of ''Pulchratia'' are convergent, weak or entirely lost in adult specimens, while the lateral ridges are slightly divergent from the hinge. Distribution Fossil remains of ''Pulchratia'' are not known outside of North America, suggesting the genus may have been endemic to the continent. Remains of ''Pulchr ...
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Echinaria (brachiopod)
''Echinaria'' is an extinct genus of brachiopods which lived during the Upper Carboniferous and Lower Permian periods. Its fossils have been found in Eurasia, North America and northern South America. Description Like other echinoconchids, ''Echinaria'' had thin, recumbent spines arranged in concentric bands on both valves, as well as a corpus with a planoconvex profile and deep cavity. Members of this genus had cardinal ridges and an elongated outline which widened anteriorly. ''Echinaria'' are medium to large-sized echinoconchids, and while notably larger than ''Echinoconchus ''Echinoconchus'' is an extinct genus of brachiopods which lived during the Lower Carboniferous period. The genus was abundant and had a cosmopolitan distribution. Description Like other echinoconchids, ''Echinoconchus'' had thin, recumbent spin ...'', many similarities are observed between the two genera and they are placed in the same subfamily. Classification Leighton & Maples (2002) conducted m ...
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Echinoconchus
''Echinoconchus'' is an extinct genus of brachiopods which lived during the Lower Carboniferous period. The genus was abundant and had a cosmopolitan distribution. Description Like other echinoconchids, ''Echinoconchus'' had thin, recumbent spines arranged in concentric bands on both Valve (mollusc), valves, as well as a corpus with a planoconvex profile and deep cavity. This genus had a transversely subcircular outline, weakly concave dorsal valve and short dorsal trail. Ridges supporting the narrow Cardinal processes, cardinal process are present, as well as elongated, mildly anteriorly-raised Adductor muscles (bivalve), adductor muscle scars. Classification Leighton & Maples (2002) conducted multiple phylogenetic analyses which are strongly in agreement that the four subfamilies Buxtoniinae, Echinoconchinae, Pustulinae and Juresaniinae form the family Echinoconchidae. The cladogram results of their phylogenetic analyses are displayed in the cladogram below: References

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Pustula (brachiopod)
''Pustula'' is an extinct genus of brachiopods which lived during the Carboniferous period. It is the type genus of the subfamily Pustulinae. Its fossils have been found in Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ..., North America and northern Africa. Description Members of the genus ''Pustula'' had a subrectangular outline with a weak ventral sulcus, minimal trails and dorsal median folds. Like other echinoconchids, they had recumbent spines arranged in irregular concentric bands, with ''Pustula'' having elongated spine bases and weak spine differentiation. The dorsal face of this brachiopod was trifid, and the cardinal process was narrow. Marginal ridges and cardinal process pits are absent in this genus, while the cardinal ridges diminished towards the ears ...
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Echinoconchinae
Echinoconchinae is an extinct subfamily of brachiopods which lived during the Carboniferous and Permian periods in marine habitats. The family had a cosmopolitan distribution. Taxonomy The exact evolutionary relationships of Echinoconchinae relative to other groups of the suborder Productidina have been a matter of extensive debate throughout much of the 20th Century, primarily due to the three genera '' Buxtonia'', ''Pustula'' and '' Juresania'' (with the debate later expanded to their families and subfamilies) shifting in position repeatedly between phylogenies and classifications. The emphasis on internal versus external characters to determine the systematics of these groups has largely been responsible for this: the original ''Treatise on Invertebrate Paleontology The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by m ...
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Pustulinae
Pustulinae is an extinct subfamily of medium to large-sized brachiopods which lived during the Early Carboniferous period in marine habitats. It is named after the type genus ''Pustula''. Taxonomy The exact evolutionary relationships of Pustulinae relative to other groups of the suborder Productidina have been a matter of extensive debate throughout much of the 20th Century, primarily due to the three genera '' Buxtonia'', ''Pustula'' and '' Juresania'' (with the debate later expanded to their families and subfamilies) shifting in position repeatedly between phylogenies and classifications. The emphasis on internal versus external characters to determine the systematics of these groups has largely been responsible for this: the original ''Treatise on Invertebrate Paleontology'' published in 1965 placed emphasis on internal characters (including the cardinal process) as diagnostic, whereas the 2000 revision primarily used external features and shell shape, resulting in differing ...
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Juresaniinae
Juresaniinae is an extinct subfamily of brachiopods which lived during the Carboniferous and Permian periods in marine habitats. Taxonomy The exact evolutionary relationships of Juresaniinae relative to other groups of the suborder Productidina have been a matter of extensive debate throughout much of the 20th Century, primarily due to the three genera '' Buxtonia'', ''Pustula'' and '' Juresania'' (with the debate later expanded to their families and subfamilies) shifting in position repeatedly between phylogenies and classifications. The emphasis on internal versus external characters to determine the systematics of these groups has largely been responsible for this: the original ''Treatise on Invertebrate Paleontology'' published in 1965 placed emphasis on internal characters (including the cardinal process) as diagnostic, whereas the 2000 revision primarily used external features and shell shape, resulting in differing classification of these clades. More recently, Leighton ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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