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Codonaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single family of the order Boraginales within the asterids. Under the older Cronquist system it was included in Lamiales, but it is now clear that it is no more similar to the other families in this order than they are to families in several other asterid orders. A revision of the Boraginales, also from 2016, split the Boraginaceae in eleven distinct families: Boraginaceae '' sensu stricto'', Codonaceae, Coldeniaceae, Cordiaceae, Ehretiaceae, Heliotropiaceae, Hoplestigmataceae, Hydrophyllaceae, Lennoaceae, Namaceae, and Wellstediaceae. These plants have alternately arranged leaves, or a combination of alternate and opposite leaves. The leaf blades usually have a narrow shape; many are linear or lance-shaped. They are smooth-edged or toothed, and some hav ...
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Boraginales
Boraginales is an order of flowering plants in the asterid clade. It includes the Boraginaceae and a number of other families, with a total of about 125 genera and 2,700 species. Its herbs, shrubs, trees and lianas (vines) have a worldwide distribution. Taxonomy History The classification of plants now known as Boraginales dates to the ''Genera plantarum'' (1789) when Antoine Laurent de Jussieu named a group of plants Boragineae, to include the genus ''Borago'', now the type genus. However, since the first valid description was by Friedrich von Berchtold and Jan Svatopluk Presl (1820), the botanical authority is given as Juss. ex Bercht. & J.Presl, where the ''ex'' refers to the prior authority of Jussieu. Lindley (1853) changed the name to the modern Boraginaceae. Jussieu divided the Boragineae into five groups. Since then Boraginaceae has been treated either as a large family with several subfamilies, or as a smaller family with several closely related families. The famil ...
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APG IV System
The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published in 2016, seven years after its predecessor the APG III system was published in 2009, and 18 years after the first APG system was published in 1998. In 2009, a linear arrangement of the system was published separately; the APG IV paper includes such an arrangement, cross-referenced to the 2009 one. Compared to the APG III system, the APG IV system recognizes five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, making a total of 64 angiosperm orders and 416 families. In general, the authors describe their philosophy as "conservative", based on making changes from APG III only where "a well-supported need" has been demonstrated. This has sometimes resulted in placements that a ...
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Asterids
In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade (a monophyletic group). Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total flowering plant species. Well-known plants in this clade include the common daisy, forget-me-nots, nightshades (including potatoes, eggplants, tomatoes, chili peppers and tobacco), the common sunflower, petunias, yacon, morning glory, sweet potato, coffee, lavender, lilac, olive, jasmine, honeysuckle, ash tree, teak, snapdragon, sesame, psyllium, garden sage, table herbs such as mint, basil, and rosemary, and rainforest trees such as Brazil nut. Most of the taxa belonging to this clade had been referred to as Asteridae in the Cronquist system (1981) and as Sympetalae in earlier systems. The name asterids (not necessarily capitalised) resembles the earlier botanical name but is intended to be the name of a clade rather than a formal ranked name ...
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Pollination
Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, and bats; water; wind; and even plants themselves, when self-pollination occurs within a closed flower. Pollination often occurs within a species. When pollination occurs between species, it can produce hybrid offspring in nature and in plant breeding work. In angiosperms, after the pollen grain (gametophyte) has landed on the stigma, it germinates and develops a pollen tube which grows down the style until it reaches an ovary. Its two gametes travel down the tube to where the gametophyte(s) containing the female gametes are held within the carpel. After entering an ovum cell through the micropyle, one male nucleus fuses with the polar bodies to produce the endosperm tissues, while the other fuses with the ovule to produce the embr ...
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Dioecy
Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is biparental reproduction. Dioecy has costs, since only about half the population directly produces offspring. It is one method for excluding self-fertilization and promoting allogamy (outcrossing), and thus tends to reduce the expression of recessive deleterious mutations present in a population. Plants have several other methods of preventing self-fertilization including, for example, dichogamy, herkogamy, and self-incompatibility. Dioecy is a dimorphic sexual system, alongside gynodioecy and androdioecy. In zoology In zoology, dioecious species may be opposed to hermaphroditic species, meaning that an individual is either male or female, in which case the synonym gonochory is more often used. Most animal species are dioecious (gon ...
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Petiole (botany)
In botany, the petiole () is the stalk that attaches the leaf blade to the stem, and is able to twist the leaf to face the sun. This gives a characteristic foliage arrangement to the plant. Outgrowths appearing on each side of the petiole in some species are called stipules. Leaves with a petiole are said to be petiolate, while leaves lacking a petiole are called sessile or apetiolate. Description The petiole is a stalk that attaches a leaf to the plant stem. In petiolate leaves, the leaf stalk may be long, as in the leaves of celery and rhubarb, or short. When completely absent, the blade attaches directly to the stem and is said to be sessile. Subpetiolate leaves have an extremely short petiole, and may appear sessile. The broomrape family Orobanchaceae is an example of a family in which the leaves are always sessile. In some other plant groups, such as the speedwell genus '' Veronica'', petiolate and sessile leaves may occur in different species. In the grasses (Poaceae), ...
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Lennoaceae
Lennooideae is a subfamily of parasitic flowering plants of southwestern North America and northwestern South America. The relationships of this subfamily to other plants remain uncertain. It was traditionally treated at family rank as Lennoaceae, and placed in different orders by different authors, including Lamiales (in the Cronquist system) and Solanales (Dahlgren system). More recently, molecular phylogenetic publications grouped it within the clade "Euasterids I", and most recently, it was demoted to a subfamily of the family Boraginaceae in the APG II system. This subfamily has a disjunct distribution, occurring in Colombia as well as a separate area in southwestern North America, covering parts of California, Arizona and Mexico. It consists of up to three genera, '' Ammobroma'', ''Lennoa'' and ''Pholisma'', which among them hold around five species, including the desert Christmas tree, ''Pholisma arenarium'', and sandfood, ''Pholisma sonorae''. Members of this subfamily a ...
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Hoplestigmataceae
''Hoplestigma'' is a genus of flowering plants in the family Hoplestigmataceae.Vernon H. Heywood, Richard K. Brummitt, Ole Seberg, and Alastair Culham. 2007. ''Flowering Plant Families of the World''. Firefly Books: Ontario, Canada. . One study of pollen suggested that ''Hoplestigma'' might be related to the family Ehretiaceae.Joan W. Nowicke and James S. Miller. 1989. "Pollen morphology and the relationships of Hoplestigmataceae". ''Taxon'' 38(1):12-16. In a comparison of chloroplast DNA sequences in 2014, ''Hoplestigma'' formed a strongly supported clade with ''Coldenia'' and genera that have always been placed in Cordiaceae.Maximilian Weigend, Federico Luebert, Marc Gottschling, Thomas L.P. Couvreur, Hartmut H. Hilger and James S. Miller. 2014. "From capsules to nutlets — phylogenetic relationships in the Boraginales". ''Cladistics'' 30(5):508-518. . The authors of that study recommended that ''Hoplestigma'' and ''Coldenia'' be included in Cordiaceae. However, it was traditi ...
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Heliotropiaceae
Heliotropiaceae is a cosmopolitan family of flowering plants with approximately 450 species worldwide, though it is concentrated especially in the tropics and subtropics. There are the following four genera in the family: * ''Heliotropium (''incl. ''Tournefortia'' L.) * ''Euploca'' Nutt. * ''Ixorhea'' Fenzl * '' Myriopus'' Small ''Ixorhea'' is sister to ''Euploca'' and ''Myriopus''. Together they form a clade sister to ''Heliotropium'', which comprises four major clades: ''Heliotropium'' sect. ''Heliothamnus'' I.M.Johnst., Old World ''Heliotropium'', ''Heliotropium'' sect. ''Cochranea'' (Miers) Post & Kuntze, and the ''Tournefortia''-clade, the latter comprising ''Tournefortia'' sect. ''Tournefortia'' and all remaining New World species of ''Heliotropium''. History Prior to a 2016 revision, Heliotropiaceae was considered a subfamily of the Boraginaceae: Heliotropioideae. Even before that, however, there was already some indication in the field that Heliotropiaceae deserved to b ...
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Ehretiaceae
Ehretioideae is a subfamily of the flowering plant family Boraginaceae. Genera *''Bourreria'' P.Browne *'' Cortesia'' Cav. *''Ehretia'' P.Browne *''Halgania'' Gaudich. *''Ixorhea'' Fenzl *''Lepidocordia'' Ducke *'' Menais'' Loefl. *'' Patagonula'' L. *''Rochefortia'' Sw. *''Rotula'' Lour. *''Tiquilia'' Pers. Christiaan Hendrik Persoon (1 February 1761 – 16 November 1836) was a German mycologist who made additions to Linnaeus' mushroom taxonomy. Early life Persoon was born in South Africa at the Cape of Good Hope, the third child of an imm ... References Asterid subfamilies {{Asterid-stub ...
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