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Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s,
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s and
herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
s in 146, to 156
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
with a worldwide distribution. The
APG IV system The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publishe ...
from 2016 classifies the Boraginaceae as single family of the order Boraginales within the
asterids In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade (a monophyletic group). Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total floweri ...
. Under the older Cronquist system it was included in Lamiales, but it is now clear that it is no more similar to the other families in this order than they are to families in several other asterid orders. A revision of the Boraginales, also from 2016, split the Boraginaceae in eleven distinct families: Boraginaceae '' sensu stricto'', Codonaceae, Coldeniaceae, Cordiaceae, Ehretiaceae, Heliotropiaceae, Hoplestigmataceae, Hydrophyllaceae,
Lennoaceae Lennooideae is a subfamily of parasitic flowering plants of southwestern North America and northwestern South America. The relationships of this subfamily to other plants remain uncertain. It was traditionally treated at family rank as Lennoacea ...
, Namaceae, and Wellstediaceae. These plants have alternately arranged leaves, or a combination of alternate and opposite leaves. The leaf blades usually have a narrow shape; many are linear or lance-shaped. They are smooth-edged or toothed, and some have petioles. Most species have bisexual flowers, but some taxa are
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
. Most
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds ...
is by
hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
ns, such as
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s. Most species have
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s that have a coiling shape, at least when new, called scorpioid cymes. The flower has a usually five-lobed calyx. The corolla varies in shape from rotate to bell-shaped to tubular, but it generally has five lobes. It can be green, white, yellow, orange, pink, purple, or blue. There are five
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s and one style with one or two stigmas. The fruit is a
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
, sometimes fleshy. Most members of this family have hairy leaves. The coarse character of the hairs is due to
cystolith Cystolith (Gr. "cavity" and "stone") is a botanical term for outgrowths of the epidermal cell wall, usually of calcium carbonate, formed in a cellulose matrix in special cells called lithocysts, generally in the leaf of plants. Cystoliths are pr ...
s of
silicon dioxide Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is one ...
and calcium carbonate. These hairs can induce an adverse skin reaction, including itching and rash in some individuals, particularly among people who handle the plants regularly, such as gardeners. In some species,
anthocyanin Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical compo ...
s cause the flowers to change color from red to blue with age. This may be a signal to
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
s that a flower is old and depleted of
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
and
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
.Hess, D. 2005. ''Systematische Botanik''. Well-known members of the family include: *
alkanet Alkanet is the common name of several related plants in the borage family (Boraginaceae): * Alkanet, ''Alkanna tinctoria'', the source of a red dye; this is the plant most commonly called simply "alkanet" * Various other plants of the genus '' Alk ...
(''Alkanna tinctoria'') * borage (''Borago officinalis'') *
comfrey ''Symphytum'' is a genus of flowering plants in the borage family, Boraginaceae, known by the common name comfrey (pronounced ). There are 59 recognized species.WFO (2022): Symphytum L. Published on the Internet; http://www.worldfloraonline.org/ ...
(''Symphytum'' spp.) * fiddleneck (''Amsinckia'' spp.) * forget-me-not (''Myosotis'' spp.) * geigertree (''Cordia sebestena'') *
green alkanet ''Pentaglottis'' is a monotypic genus of flowering plants in the family Boraginaceae. It is represented by a single species, ''Pentaglottis sempervirens'', commonly known as the green alkanet, evergreen bugloss or alkanet, and is a bristly, pe ...
(''Pentaglottis sempervirens'') * heliotrope (''Heliotropium'' spp.) *
hound's tongue ''Cynoglossum'' is a genus of small-flowered plants in the family Boraginaceae (borage family). ''Cynoglossum officinale'', the common hound's-tongue, is a native of Asia, Africa, and Europe. It has been introduced into North America, and it is ...
(''Cynoglossum'' spp.) *
lungwort ''Pulmonaria'' (lungwort) is a genus of flowering plants in the family Boraginaceae, native to Europe and western Asia, with one species (''P. mollissima'') east to central Asia. According to various estimates there may be between 10 and 18 spe ...
(''Pulmonaria'' spp.) * oysterplant (''Mertensia maritima'') * purple viper's bugloss/Salvation Jane (''Echium plantagineum'') * Siberian bugloss (''Brunnera macrophylla'') *
viper's bugloss ''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the borag ...
(''Echium vulgare'')


Genera

According to
Kew Kew () is a district in the London Borough of Richmond upon Thames. Its population at the 2011 census was 11,436. Kew is the location of the Royal Botanic Gardens ("Kew Gardens"), now a World Heritage Site, which includes Kew Palace. Kew is a ...
; * '' Actinocarya'' * '' Adelinia'' * ''
Adelocaryum ''Adelocaryum'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is southern Arabian Peninsula and the Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, p ...
'' * '' Aegonychon'' * '' Afrotysonia'' * '' Ailuroglossum'' * ''
Alkanna ''Alkanna'' is a genus of herbaceous plants including about 60 species of the family Boraginaceae. Selected species * '' A. amana'' Rech.f. * '' A. angustifolia'' Sümbül * '' A. areolata'' Boiss. * '' A. attilae'' P.H.D ...
'' * '' Amblynotus'' * ''
Amphibologyne ''Amphibologyne'' is a genus of flowering plants belonging to the family Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distri ...
'' * ''
Amsinckia ''Amsinckia'' is a genus of flowering plants commonly known as fiddlenecks. The common name is derived from the flower stems, which curl over at the top in a manner reminiscent of the head of a fiddle. Fiddlenecks are in the family Boraginac ...
'' * '' Amsinckiopsis'' * ''
Anchusa The genus ''Anchusa'' belongs to the borage family (Boraginaceae). It includes about 35 species found growing in Europe, North Africa, South Africa and Western Asia. They are introduced in the United States. They consist of annual plants, bie ...
'' * '' Ancistrocarya'' * ''
Andersonglossum ''Andersonglossum'' is a small genus of North American plants in the borage family (Boraginaceae). They are commonly called American comfreys, wild comfreys, or hound's tongues. Taxonomy Members of this genus were formerly placed in the genus ' ...
'' * '' Anoplocaryum'' * '' Antiotrema'' * '' Antiphytum'' * ''
Arnebia ''Arnebia'' is a genus of flowering plants in the family Boraginaceae. There are about 38 species, most are located in the Mediterranean region and eastwards to the Himalayas and one species extending down to tropical Africa. The generic name " ...
'' * '' Asperugo'' * '' Auxemma'' * '' Borago'' * '' Bothriospermum'' * '' Bourreria'' * '' Brachybotrys'' * '' Brandella'' * ''
Brunnera ''Brunnera'' is a genus of flowering plants in the family Boraginaceae. They are rhizomatous perennials, native to the woodlands of Eastern Europe and North West Asia. They have hairy leaves and sprays of blue flowers in spring. Numerous cultiv ...
'' * ''
Buglossoides ''Buglossoides'' is a genus consisting of 15 species of annual or perennial herbs, native to Europe and Asia. They grow naturally in habitats ranging from sunny scrub to rocky slopes and woodland areas. These plants are covered in fine bristles ...
'' * '' Caccinia'' * ''
Cerinthe ''Cerinthe'' is a genus of flowering plants in the family Boraginaceae, known as honeyworts. The genus is characterised by a calyx made up of separate, rather than fused, sepals, a tubular corolla, and the schizocarpic fruit that divides into ...
'' * '' Chionocharis'' * '' Choriantha'' * ''
Codon The genetic code is the set of rules used by living cells to translate information encoded within genetic material ( DNA or RNA sequences of nucleotide triplets, or codons) into proteins. Translation is accomplished by the ribosome, which links ...
'' * ''
Coldenia ''Coldenia'', named after C. Colden, is a monotypic genus of flowering plants traditionally included in the borage family, Boraginaceae ''sensu lato''. It was assigned to the subfamily Ehretioideae, but molecular data revealed it to be more clos ...
'' * ''
Cordia ''Cordia'' is a genus of flowering plants in the borage family, Boraginaceae. It contains about 300 species of shrubs and trees, that are found worldwide, mostly in warmer regions. Many of the species are commonly called manjack, while ''bocote' ...
'' * '' Craniospermum'' * '' Crucicaryum'' * ''
Cryptantha ''Cryptantha'' is a genus of flowering plants in the borage family, Boraginaceae. They are known commonly as cat's eyes and popcorn flowers (the latter name is also used to refer to the closely related genus ''Plagiobothrys'',Hasenstab-Lehman, ...
'' * '' Cynoglossopsis'' * ''
Cynoglossum ''Cynoglossum'' is a genus of small-flowered plants in the family Boraginaceae (borage family). ''Cynoglossum officinale'', the common hound's-tongue, is a native of Asia, Africa, and Europe. It has been introduced into North America, and it is ...
'' * '' Cynoglottis'' * '' Cystostemon'' * '' Dasynotus'' * '' Decalepidanthus'' * '' Draperia'' * '' Echiochilon'' * '' Echium'' * ''
Ehretia ''Ehretia'' is a genus of flowering plants in the borage family, Boraginaceae. It contains about 50 species. The generic name honors German botanical illustrator Georg Dionysius Ehret (1708–1770). Species Accepted species and other notable t ...
'' * ''
Ellisia ''Ellisia'' is a genus of flowering plants in the forget-me-not family (Boraginaceae), containing the sole species ''Ellisia nyctelea''. It is native to North America, where it is also known as Aunt Lucy, false baby blue eyes, and waterpod. The ...
'' * ''
Embadium ''Embadium'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is Central and Southern Australia. Species: *''Embadium jobnstonii'' *''Embadium stagnense'' *''Embadium uncinatum'' References

{{Taxo ...
'' * '' Emmenanthe'' * '' Eremocarya'' * ''
Eriodictyon ''Eriodictyon'' is a genus of plants known by the common name yerba santa within the Hydrophylloideae subfamily of the borage family, Boraginaceae. They are distributed throughout the southwestern United States and Mexico. Description Most spe ...
'' * ''
Eritrichium ''Eritrichium'' (alpine forget-me-not) is a genus of flowering plants in the family Boraginaceae. It contains 78 species. Notable members include '' Eritrichium howardii'' and ''Eritrichium nanum''. Its native range stretches from temperate Euras ...
'' * '' Eucrypta'' * ''
Euploca ''Euploca'' is an almost cosmopolitan genus of plants with around 100 species. It was first described by Thomas Nuttall in 1837. While part of the broadly defined Boraginaceae in the APG IV system from 2016, a revision of the order Boraginales fr ...
'' * '' Gastrocotyle'' * ''
Glandora ''Glandora'' is a genus of flowering plants in the family Boraginaceae, native to the western and central Mediterranean region; Morocco, Algeria, Portugal, Spain, France, Italy and Greece. It was split from ''Lithodora ''Lithodora'' is a genus ...
'' * '' Greeneocharis'' * '' Gyrocaryum'' * ''
Hackelia ''Hackelia'' (stickseeds) is a genus of plants in the borage family, Boraginaceae. It includes 54 species found in North America, western South America, temperate Eurasia, and Australia. 12 species are native to California California is a ...
'' * '' Halacsya'' * ''
Halgania ''Halgania'' is a genus of small shrubs in the family Boraginaceae. The genus comprises about 20 species that are endemic to Australia. ''Halgania'' is named for Emmanuel Halgan, a vice-admiral in the French Navy. Features ''Halgania'' species ...
'' * '' Harpagonella'' * '' Heliocarya'' * '' Heliotropium'' * '' Hesperochiron'' * '' Heterocaryum'' * '' Huynhia'' * ''
Hydrophyllum ''Hydrophyllum'' is a genus of herbaceous perennial plants in the waterleaf family. It consists of nine species, all of which are native to North America. All ''Hydrophyllum'' are found in areas of mesic or moist soil. Species * ''Hydrophyllum ...
'' * '' Iberodes'' * '' Ivanjohnstonia'' * ''
Ixorhea ''Ixorhea'' is a genus of flowering plants belonging to the family Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution. ...
'' * '' Johnstonella'' * ''
Lappula ''Lappula'' is a genus of flowering plants in the borage family known generally as stickseeds. They are native to the northern hemisphere. These are annual herbs producing funnel-shaped flowers and prickly fruits. One of the best known species is ...
'' * ''
Lasiocaryum ''Lasiocaryum'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the T ...
'' * ''
Lennoa ''Lennoa'' is a monotypic genus of flowering plants belonging to the family Boraginaceae. It only contains one known species, Lennoa madreporoides Lex. It is within the subfamily of Lennoaceae. Its native range is Mexico to Venezuela. It is foun ...
'' * ''
Lepechiniella ''Lepechiniella'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is Iran and Western Himalaya, to Xinjiang (in China). It is also found in the countries of Afghanistan, Kazakhstan, Kyrgyzstan, Pakistan, Taji ...
'' * ''
Lepidocordia ''Lepidocordia'' is a genus of flowering plants belonging to the family Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribut ...
'' * '' Lindelofia'' * ''
Lithodora ''Lithodora'' is a genus of flowering plants in the family Boraginaceae, native to southwestern Europe, southern Greece, Turkey and Algeria. They are low-growing, evergreen shrubs and subshrubs, producing 5-lobed blue or white flowers. The Greek ...
'' * ''
Lithospermum ''Lithospermum'' is a genus of plants belonging to the family Boraginaceae. The genus is distributed nearly worldwide, but most are native to the Americas and the center of diversity is in the southwestern United States and Mexico. Species are ...
'' * '' Lobostemon'' * '' Maharanga'' * ''
Mairetis ''Mairetis'' is a monotypic genus of flowering plants belonging to the family Boraginaceae. It only contains one known species, ''Mairetis microsperma'' (Boiss.) I.M.Johnst. It is native to the Canary Islands and Morocco. The genus name of ''Ma ...
'' * '' Mattiastrum'' * '' Megacaryon'' * '' Melanortocarya'' * '' Memoremea'' * ''
Mertensia ''Mertensia'' is a genus of flowering plants in the family Boraginaceae. They are perennial herbaceous plants with blue or sometimes white flowers that open from pink-tinged buds. Such a change in flower color is common in Boraginaceae and is ca ...
'' * '' Microcaryum'' * '' Microparacaryum'' * '' Microula'' * ''
Mimophytum ''Mimophytum'' is a genus of flowering plants in the family Boraginaceae. The species occur in Northeastern Mexico and adjacent areas of Texas, United States. They are similar to the closely related genus ''Omphalodes'' but a distinct group. Mo ...
'' * '' Moltkia'' * ''
Moltkiopsis ''Moltkiopsis'' is a monotypic genus of flowering plants belonging to the family Boraginaceae. The only species is ''Moltkiopsis ciliata''. Its native range is northern Africa to Sahel and the Arabian Peninsula. It is found in the regions of Alg ...
'' * ''
Moritzia ''Moritzia'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is Costa Rica to Southern Tropical America. It is found in Brazil, Colombia, Costa Rica, Ecuador, Panamá, Peru and Venezuela. The genus name of ...
'' * ''
Myosotidium ''Myosotidium'' is a genus of plants belonging to the family Boraginaceae. This genus is represented by the single species ''Myosotidium hortensia'', the Chatham Islands lily, giant forget-me-not or Chatham Islands forget-me-not, which is endemi ...
'' * '' Myosotis'' * ''
Myriopus ''Myriopus'' is a genus of flowering plants in the borage and forget-me-not family Boraginaceae (or Heliotropiaceae Heliotropiaceae is a cosmopolitan family of flowering plants with approximately 450 species worldwide, though it is concentrated ...
'' * '' Nama'' * '' Neatostema'' * '' Nemophila'' * '' Nesocaryum'' * ''
Nihon Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
'' * ''
Nogalia ''Nogalia'' is a monotypic genus of flowering plants belonging to the family Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distrib ...
'' * ''
Nonea ''Nonea'' is a genus of flowering plants in the family Boraginaceae. Sometimes known as monkswort, these are herbaceous perennials or annual plants, native to Europe, Asia and Africa. The leaves are grayish-green, and the plants are hairy all ov ...
'' * '' Ogastemma'' * ''
Omphalodes ''Omphalodes'' (navelwort) is a genus of flowering plants in the family Boraginaceae, widely distributed in the temperate Northern Hemisphere. In spring they produce blue or white flowers similar to forget-me-nots. Both the Greek ''Omphalodes'' ...
'' * '' Omphalolappula'' * ''
Omphalotrigonotis ''Omphalotrigonotis'' is a genus of flowering plants belonging to the family Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide dist ...
'' * '' Oncaglossum'' * ''
Onosma ''Onosma'' is a genus of flowering plants in the family Boraginaceae. They are native to the Mediterranean and western Asia. They grow in dry, sunny habitats with rocky, sandy substrates.Koyuncu, O., et al. (2013)A new ''Onosma'' (Boraginaceae) s ...
'' * '' Oreocarya'' * '' Paracaryum'' * '' Paramoltkia'' * '' Pardoglossum'' * ''
Pectocarya ''Pectocarya'' is a plant genus of about 15 species in the family Boraginaceae. Plants in this genus are known generally as combseeds. They are small annual plants which bear tiny white flowers no more than 3 millimeters in diameter. Their fruits ...
'' * ''
Pentaglottis ''Pentaglottis'' is a monotypic genus of flowering plants in the family Boraginaceae. It is represented by a single species, ''Pentaglottis sempervirens'', commonly known as the green alkanet, evergreen bugloss or alkanet, and is a bristly, pe ...
'' * '' Phacelia'' * ''
Pholisma The genus ''Pholisma'' (Nutt. ex Hook.) consists of three to five species of desert-dwelling, primarily subterranean plants. Most notable of the genus is '' Pholisma sonorae'', native to the Southwestern United States and Mexico. The species is ...
'' * ''
Pholistoma ''Pholistoma'' is a small genus of flowering plants in the borage family known generally as fiestaflowers. There are three species, all native to a section of western North America between Oregon and Baja California Baja California (; 'Lower ...
'' * '' Plagiobothrys'' * '' Podonosma'' * '' Pontechium'' * '' Pseudoheterocaryum'' * '' Pseudolappula'' * '' Pulmonaria'' * ''
Rindera ''Rindera'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is north-western Africa (Algeria), south-eastern and eastern Europe (Greece, East European Russia, Romania, Ukraine and Yugoslavia) to western and c ...
'' * ''
Rochefortia ''Rochefortia'' is a genus of flowering plants in family Boraginaceae. Its species are native to North America, the Caribbean, and South America. Species Species include: * '' Rochefortia acanthophora'' (syn: ''Ehretia ''Ehretia'' is a gen ...
'' * ''
Rochelia ''Rochelia'' is a genus of flowering plants belonging to the family Boraginaceae. It is also in subtribe ''Eritrichiinae''. Its wide native range extends from Europe (within Albania, Austria, Baltic States, Belarus, Belgium, Bulgaria, Central Eur ...
'' * '' Romanzoffia'' * ''
Rotula A ''rotulus'' (plural ''rotuli'') or ''rotula'' (pl. ''rotulae'') is often referred to as a "vertical roll," is a long and narrow strip of writing material, historically papyrus or parchment, that is wound around a wooden axle or rod. Rotuli are ...
'' * '' Sauria'' * ''
Selkirkia ''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they compr ...
'' * '' Simpsonanthus'' * ''
Sinojohnstonia ''Sinojohnstonia'' is a genus of flowering plants belonging to the family Boraginaceae. It is native to China (within north-central, south-central, south-east and Inner Mongolia regions). The genus name of ''Sinojohnstonia'' is in honour of Ivan ...
'' * '' Solenanthus'' * '' Stenosolenium'' * '' Stephanocaryum'' * '' Suchtelenia'' * '' Symphytum'' * '' Thaumatocaryum'' * '' Thyrocarpus'' * '' Tianschaniella'' * ''
Tiquilia ''Tiquilia'' is a genus of flowering plants in the borage family, Boraginaceae. The 27 species in this genus are known by the common name crinklemat. They are native to the Western Hemisphere and are mostly found in desert regions. Selected spe ...
'' * '' Tournefortia'' * '' Trachelanthus'' * ''
Trachystemon ''Trachystemon'' is a genus of flowering plants belonging to the family Boraginaceae. Its native range is Bulgaria to Turkey. It is a monotypic genus, with the only member being ''Trachystemon orientalis''(L.) D.Don. Trachystemon orientalis ' ...
'' * ''
Trichodesma ''Trichodesma'' is a genus of flowering plants in the borage family, Boraginaceae. There are about 40 to 45 species distributed in tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tr ...
'' * '' Trigonocaryum'' * '' Trigonotis'' * '' Turricula'' * '' Varronia'' * '' Valentiniella'' * ''
Wellstedia ''Wellstedia'' is a genus of flowering plants traditionally included in the family Boraginaceae ''s.l.'', but placed in its own family, Wellstediaceae within the Boraginales order, by the Boraginales Working Group. Species include: * '' Wellste ...
'' * ''
Wigandia ''Wigandia'' is a genus of flowering plants within the waterleaf subfamily, Hydrophylloideae. They are found mainly in Central America and South America, though one or two species are found as far north as the United States. Some are grown as o ...
''


References


External links


Distribution MapMissouri Botanical Garden

BoraginaceaeMabberley's Plant-Book

BoraginaceaePlant NamesIPNI

Home page of James L. Reveal and C. Rose Broome

Boraginaceae (Search Exact)Name SearchTropicosMissouri Botanical Garden


Topwalks.net: Walking Routes in Spain.
Boraginaceae.
Integrated Taxonomic Information System (ITIS).


Further reading

* Diane, N., et al. 2002
A systematic analysis of ''Heliotropium'', ''Tournefortia'', and allied taxa of the Heliotropiaceae (Boraginales) based on ITS1 sequences and morphological data.
''American Journal of Botany'' 89(2), 287-95. * Gottschling, M., et al. (2001). Secondary structure of the ITS1 transcript and its application in a reconstruction of the phylogeny of Boraginales. ''Plant Biology'' 3, 629-36. {{Authority control Asterid families