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Buxales
The Buxales are a small order of eudicot flowering plants, recognized by the APG IV system of 2016. The order includes the family Buxaceae; the families Didymelaceae and Haptanthaceae may also be recognized or may be included in the Buxaceae. Many members of the order are evergreen shrubs or trees, although some are herbaceous perennials. They have separate "male" (staminate) and "female" (carpellate) flowers, mostly on the same plant (i.e. they are mostly monoecious). Some species are of economic importance either for the wood they produce or as ornamental plants. Description The Buxales have relatively few obvious shared features distinguishing them from related groups of plants (i.e. few obvious synapomorphies). One is the presence of a particular type of alkaloid or pseudoalkaloid, pregnane steroids. They have unisexual flowers (i.e. separate staminate or "male" flowers and carpellate or "female" flowers), most being monoecious. The flowers are small, less than 7 mm acr ...
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APG III System
The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a further revision, the APG IV system. Along with the publication outlining the new system, there were two accompanying publications in the same issue of the Botanical Journal of the Linnean Society: * The first, by Chase & Reveal, was a formal phylogenetic classification of all land plants (embryophytes), compatible with the APG III classification. As the APG have chosen to eschew ranks above order, this paper was meant to fit the system into the existing Linnaean hierarchy for those that prefer such a classification. The result was that all land plants were placed in the class Equisetopsida, which was then divided into 16 subclasses and a multitude of superorders. * The second, by Haston ''et al.'', was a linear sequence of families followi ...
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Didymeles
''Didymeles'' is a genus of flowering plants. It is variously treated as the only genus of the family Didymelaceae — or in the family Buxaceae, as in the APG IV system. The genus ''Didymeles'' is restricted to Madagascar and the Comoros. In Madagascar, the two species are found in humid montane habitats, up to 1,500 m, in two disconnected areas in the north-east and south-east of the island. Fossils assignable to this genus have been found in New Zealand, suggesting a much wider distribution in the past. The conservation status of ''D. integrifolia'' has been assessed as of "least concern". Description Species are dioecious (i.e. "male" and "female" flowers occur on separate plants). Individual flowers have a very simple structure, without obvious petals or sepals. Female flowers have been interpreted in different ways, either as having two carpels or as occurring close together in pairs, each with a single carpel. ''Didymeles'' species are evergreen trees. The simp ...
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Eudicot
The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the later evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots. Numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Some common and familiar eudicots include sunflower, dandelion, forget-me-not, cabbage, apple, buttercup, maple, and macadamia. Most leafy trees of midlatitudes also belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, and ''Ginkgo biloba'', which is not an angiosperm. Description The close relationships among flowering plants with tricolpate po ...
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Eudicots
The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the later evolutionary divergence of tricolpate dicots from earlier, less specialized, dicots. Numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Some common and familiar eudicots include sunflower, dandelion, forget-me-not, cabbage, apple, buttercup, maple, and macadamia. Most leafy trees of midlatitudes also belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, and ''Ginkgo biloba'', which is not an angiosperm. Description The close relationships among flowering plants with tricolpat ...
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Core Eudicots
The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicots by previous authors. The botanical terms were introduced in 1991 by evolutionary botanist James A. Doyle and paleobotanist Carol L. Hotton to emphasize the later evolutionary divergence of tricolpate dicotyledon, dicots from earlier, less specialized, dicots. Numerous familiar plants are eudicots, including many common food plants, trees, and ornamentals. Some common and familiar eudicots include sunflower, dandelion, forget-me-not, cabbage, apple, buttercup, maple, and macadamia. Most leafy trees of midlatitudes also belong to eudicots, with notable exceptions being magnolias and tulip trees which belong to magnoliids, and ''Ginkgo biloba'', which is not an angiosperm. Description The close relationships among flowering plants with poll ...
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Didymelaceae
''Didymeles'' is a genus of flowering plants. It is variously treated as the only genus of the family Didymelaceae — or in the family Buxaceae, as in the APG IV system. The genus ''Didymeles'' is restricted to Madagascar and the Comoros. In Madagascar, the two species are found in humid montane habitats, up to 1,500 m, in two disconnected areas in the north-east and south-east of the island. Fossils assignable to this genus have been found in New Zealand, suggesting a much wider distribution in the past. The conservation status of ''D. integrifolia'' has been assessed as of "least concern". Description Species are dioecious (i.e. "male" and "female" flowers occur on separate plants). Individual flowers have a very simple structure, without obvious petals or sepals. Female flowers have been interpreted in different ways, either as having two carpels or as occurring close together in pairs, each with a single carpel. ''Didymeles'' species are evergreen trees. The simp ...
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Buxaceae
The Buxaceae are a small family of six genera and about 123 known species of flowering plants. They are shrubs and small trees, with a cosmopolitan distribution. A seventh genus, sometimes accepted in the past (''Notobuxus''), has been shown by genetic studies to be included within ''Buxus'' (Balthazar ''et al.'', 2000). The family is recognised by most taxonomists, and it is commonly known as the box family. However, its placement and circumscription has varied; some taxonomists treated '' Styloceras'' in its own family Stylocerataceae, ''Didymeles'' in its own family Didymelaceae, ''Haptanthus'' in Haptanthaceae (now all included in Buxaceae)), and formerly '' Simmondsia'' was included, which is not related and now usually placed in its own family Simmondsiaceae. The APG II system of 2003 recognises the family, but in a new circumscription in that it includes the genus ''Didymeles'' (two species of evergreen trees from Madagascar). However, APG II does allow the option of segr ...
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Haptanthaceae
''Haptanthus'' is a monotypic genus containing the sole species ''Haptanthus hazlettii'', a shrub or small tree known only from the locality of Matarras in the Arizona Municipality in Honduras. Its flowers are unique among the flowering plants. A single "female" (carpellate) flower has two branches on either side which carry "male" (staminate) flowers. The flowers are very simple, lacking obvious sepals or petals. The family placement of the genus has been uncertain, but based on molecular phylogenetic research, it is included in the family Buxaceae . Very few individuals have ever been found and its habitat is threatened by logging. Description ''Haptanthus hazlettii'' is a shrub or tree. It has opposite leaves spaced at 5.5–6.0 cm apart, usually arranged in two ranks (distichous). The leaves are simple with untoothed (entire) margins. There are no stipules. The leaf stalk ( petiole) is short, 7–8 mm long, the leaf blade (lamina) 10–13.5 cm long by 4.1â ...
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APG IV System
The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published in 2016, seven years after its predecessor the APG III system was published in 2009, and 18 years after the first APG system was published in 1998. In 2009, a linear arrangement of the system was published separately; the APG IV paper includes such an arrangement, cross-referenced to the 2009 one. Compared to the APG III system, the APG IV system recognizes five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, making a total of 64 angiosperm orders and 416 families. In general, the authors describe their philosophy as "conservative", based on making changes from APG III only where "a well-supported need" has been demonstrated. This has sometimes resulted in placements that a ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils ar ...
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Seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosperm plants. Seeds are the product of the ripened ovule, after the embryo sac is fertilized by sperm from pollen, forming a zygote. The embryo within a seed develops from the zygote, and grows within the mother plant to a certain size before growth is halted. The seed coat arises from the integuments of the ovule. Seeds have been an important development in the reproduction and success of vegetable gymnosperm and angiosperm plants, relative to more primitive plants such as ferns, mosses and liverworts, which do not have seeds and use water-dependent means to propagate themselves. Seed plants now dominate biological niches on land, from forests to grasslands both in hot and cold climates. The term "seed" also has a general me ...
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Angiosperm Phylogeny Website
The Angiosperm Phylogeny Website (or APweb) is a website dedicated to research on angiosperm phylogeny and taxonomy. The site is hosted by the Missouri Botanical Garden website and maintained by researchers, Peter F. Stevens and Hilary M. Davis. Peter F. Stevens is a member of the Angiosperm Phylogeny Group The Angiosperm Phylogeny Group (APG) is an informal international group of systematic botanists who collaborate to establish a consensus on the taxonomy of flowering plants (angiosperms) that reflects new knowledge about plant relationships disco ... (APG). The taxonomy presented is broadly based on the work of the APG, with modifications to incorporate new results. References External links * , hosted by thMissouri Botanical Garden Plant taxonomy American science websites Angiosperm taxonomy Angiosperm Phylogeny Group Online taxonomy databases Missouri Botanical Garden {{botany-stub ...
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