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Bryantolepis
''Bryantolepis'' is an extinct genus of arthrodire placoderm from the Early Devonian of Wyoming, Idaho, and Utah. It is known from two species, B. brachycephala, and B. williamsi. Description The genus is known from multiple parts of the skull roof, the suborbital, endocranium, and trunk shield. Taxonomy ''Bryantolepis'' is regarded as part of the subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ... Actinolepinae, or within Actinolepidoidei. References {{Taxonbar, from=Q116235524 Arthrodires Arthrodire genera ...
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Arthrodira
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus'' measured just ...
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Arthrodires
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus'' measured just ...
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Early Devonian
The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first Ammonoidea, ammonoids appeared, descending from Bactritida, bactritoid Nautiloidea, nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatite, Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era. References

{{Geological history Early Devonian, Geological epochs Devonian geochronology, *01 ...
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Extinction
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Placodermi
Placodermi (from Greek πλάξ 'plate' and δέρμα 'skin', literally 'Plate (animal anatomy), plate-skinned') is a Class (biology), class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scale (zoology), scaled or naked, depending on the species. Placoderms were among the first jawed fish; their Fish jaw, jaws likely evolved from the first of their gill arches. Placoderms are thought to be paraphyly, paraphyletic, consisting of several distinct Outgroup (cladistics), outgroups or sister taxon, sister taxa to all living jawed vertebrates, which originated among their ranks. In contrast, one 2016 analysis concluded that placodermi are likely monophyletic, though these analyses have been further dismissed with more transitional taxa between placoderms and modern gnathosthomes, solidifying their paraphyletic status. Plac ...
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Wyoming
Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the south. With a population of 576,851 in the 2020 United States census, Wyoming is the List of U.S. states and territories by population, least populous state despite being the List of U.S. states and territories by area, 10th largest by area, with the List of U.S. states by population density, second-lowest population density after Alaska. The state capital and List of municipalities in Wyoming, most populous city is Cheyenne, Wyoming, Cheyenne, which had an estimated population of 63,957 in 2018. Wyoming's western half is covered mostly by the ranges and rangelands of the Rocky Mountains, while the eastern half of the state is high-elevation prairie called the High Plains (United States), High Plains. It is drier ...
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Idaho
Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyoming to the east, Nevada and Utah to the south, and Washington and Oregon to the west. The state's capital and largest city is Boise. With an area of , Idaho is the 14th largest state by land area, but with a population of approximately 1.8 million, it ranks as the 13th least populous and the 7th least densely populated of the 50 U.S. states. For thousands of years, and prior to European colonization, Idaho has been inhabited by native peoples. In the early 19th century, Idaho was considered part of the Oregon Country, an area of dispute between the U.S. and the British Empire. It officially became U.S. territory with the signing of the Oregon Treaty of 1846, but a separate Idaho Territory was not organized until 1863, instead ...
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Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its west by Nevada. Utah also touches a corner of New Mexico in the southeast. Of the fifty U.S. states, Utah is the 13th-largest by area; with a population over three million, it is the 30th-most-populous and 11th-least-densely populated. Urban development is mostly concentrated in two areas: the Wasatch Front in the north-central part of the state, which is home to roughly two-thirds of the population and includes the capital city, Salt Lake City; and Washington County in the southwest, with more than 180,000 residents. Most of the western half of Utah lies in the Great Basin. Utah has been inhabited for thousands of years by various indigenous groups such as the ancient Puebloans, Navajo and Ute. The Spanish were the first Europe ...
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Skull Roof
The skull roof, or the roofing bones of the skull, are a set of bones covering the brain, eyes and nostrils in bony fishes and all land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparative anatomy the term is used on the full dermatocranium. Romer, A.S. & T.S. Parsons. 1977. ''The Vertebrate Body.'' 5th ed. Saunders, Philadelphia. (6th ed. 1985) In general anatomy, the roofing bones may refer specifically to the bones that form above and alongside the brain and neurocranium (i.e., excluding the marginal upper jaw bones such as the maxilla and premaxilla), and in human anatomy, the skull roof often refers specifically to the skullcap. Origin Early armoured fish did not have a skull in the common understanding of the word, but had an endocranium that was partially open above, topped by dermal bones forming armour. The dermal bones gradually evolved into a fixed unit overlaying the endocranium like a heavy "lid", protec ...
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Endocranium
The endocranium in comparative anatomy is a part of the skull base in vertebrates and it represents the basal, inner part of the cranium. The term is also applied to the outer layer of the dura mater in human anatomy. Structure Structurally, the endocranium consists of a boxlike shape, open at the top. The posterior margin exhibit the '' foramen magnum'', an opening for the spinal cord. The floor of the endocranium has several paired openings for the cranial nerves, and the anterior margin holds a spongy construction, allowing for the external nasal nerves to pass through. Romer, A.S. & T.S. Parsons. 1977. ''The Vertebrate Body.'' 5th ed. Saunders, Philadelphia. (6th ed. 1985) All bones of the structure derive from the cranial neural crest during fetal development. Endocranial elements in humans In humans and other mammals, the endocranium forms during fetal development as a cartilaginous neurocranium, that ossifies from several centers. Several of these bones merge, and in t ...
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Subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoological names with "-inae". See also * International Code of Nomenclature for algae, fungi, and plants * International Code of Zoological Nomenclature * Rank (botany) * Rank (zoology) In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family, order, class, phylum, kingdom, domain. While ... Sources {{biology-stub ...
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Actinolepidae
Actinolepidae is an extinct family of placoderm fishes which lived during the Early Devonian period. They are considered to be among the most primitive of the arthrodires, and are widely accepted to be phylogenetically basal to the group. Description The bodies of Actinolepids are wide and flat, suggesting that most members of this family were benthic fish. Their jaws were comparatively underdeveloped in comparison to the more robust-jawed arthrodires that would come after them, such as ''Dunkleosteus ''Dunkleosteus'' is an extinct genus of large armored, jawed fishes that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: ...'' and '' Coccosteus'', indicating that it likely subsisted primarily on smaller, softer-bodied animals such as mollusks or worms instead of larger, tougher prey animals. References External links * Devonian placoderms ...
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