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Borophagina
Borophagina is a subtribe of the Borophaginae, a group of extinct canids. They inhabited much of North America from the Early Miocene to the Zanclean stage of the Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Mya, and existed for approximately . Like some other borophagines, they were short-faced, heavy-jawed canids although the group included both omnivorous and hypercarnivorous species.


References

{{Taxonbar, from=Q4946287 Zanclean extinctio ...
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Borophaginae
The extinct Borophaginae form one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Hesperocyoninae and extant Caninae. Borophaginae, called "bone-crushing dogs", were endemic to North America during the Oligocene to Pliocene and lived roughly 34—2.5 million years ago, existing for about . Origin The Borophaginae descended from the subfamily Hesperocyoninae. The earliest and most primitive borophagine is the genus ''Archaeocyon'', which is a small fox-sized animal mostly found in the fossil beds in western North America. The borophagines soon diversified into several major groups. They evolved to become considerably larger than their predecessors, and filled a wide range of niches in late Cenozoic North America, from small omnivores to powerful, bear-sized carnivores, such as ''Epicyon''. Species There are 66 identified borophagine species, including 18 new ones that range from the Orellan to Blancan ages. A phylogenetic anal ...
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Canidae
Canidae (; from Latin, ''canis'', "dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within the canid family, which are the extinct Borophaginae and Hesperocyoninae, and the extant Caninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other extant and extinct species. Canids are found on all continents except Antarctica, having arrived independently or accompanied human beings over extended periods of time. Canids vary in size from the gray wolf to the fennec fox. The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails. They are mostly social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a group bree ...
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Epicyon
''Epicyon'' ("more than a dog") is a large, extinct, canid genus of the subfamily Borophaginae ("bone-crushing dogs"), native to North America. ''Epicyon'' existed for about from the Hemingfordian age of the Early Miocene, to the Hemphillian of the Late Miocene. ''Epicyon'' is the largest known canid of all time, with the type species reaching 2.4 m (7.9 ft) in length, 90 cm (35 in) in shoulder height and approximately 100–125 kg (220–276 lb) in body mass. The largest known humerus specimen belonged to an individual weighing up to . Description ''Epicyon'' had a massive head and powerful jaws that were well adapted for bone-crushing, with enlarged fourth premolars like some hyenas, giving its skull a lion-like shape rather than having a skull similar in shape to that of a wolf; the adaptation would have allowed ''Epicyon'' to scavenge as well as hunt, giving it access to the nutritious marrow other contemporary carnivores couldn't access. ''Epic ...
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Borophagus
''Borophagus'' ("gluttonous eater") is an extinct genus of the subfamily Borophaginae, a group of canids endemic to North America from the Middle Miocene epoch through the Early Pleistocene epoch 12—1.8 Mya. Evolution ''Borophagus'', like other borophagines, are loosely known as "bone-crushing" or " hyena-like" dogs. Though not the most massive borophagine by size or weight, it had a more highly evolved capacity to crunch bone than earlier, larger genera such as ''Epicyon'', which seems to be an evolutionary trend of the group (Turner, 2004). During the Pliocene epoch, ''Borophagus'' began being displaced by other ''Canid'' species such as ''Canis edwardii'' and later by ''Aenocyon dirus''. Early species of ''Borophagus'' were placed in the genus ''Osteoborus'' until recently, but the genera are now considered synonyms. Description Typical features of this genus are a bulging forehead and powerful jaws; ''Borophagus'' has been considered to be probably a scavenger by paleonto ...
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Paratomarctus
''Paratomarctus'' is an extinct monospecific genus of the Borophaginae subfamily of canids native to North America. It lived from the Middle to Late Miocene, 16.3—5.3 mya, existing for approximately . It was about the size of a coyote, and was probably a generalised predator, without the specialised adaptations of most later borophagines. ''Paratomarctus'' was one of the last of the Borophaginae and shared its habitat with other canids, including ''Borophagus'', ''Epicyon'', ''Carpocyon'', ''Aelurodon'', and the true canine, ''Canis lepophagus''. Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ... have been uncovered throughout most of the western United States. References zipcodezoo.comwww.sciencemag.org Borophagines Miocene carnivorans Miocene mammals of North A ...
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Carpocyon
''Carpocyon'' is an extinct genus of the Borophaginae subfamily of canids native to North America. It lived from the Middle to the Late Miocene, 13.6 to 5.3 Ma Mya, existing for approximately . The four species in the genus varied in size, with the largest (''C. webbi'') being about the size of a wolf; all had relatively small teeth, suggesting a diet that was more omnivorous than that of other contemporary borophagines. Species *''Carpocyon compressus'' (syn. ''Cynodesmus cuspidatus''), fossils have been found in Nevada, Nebraska, Colorado, and Kansas. *''Carpocyon limosus'', fossils have been found in Nebraska, Oklahoma, and Florida. *''Carpocyon robustus'', fossils have been found in Arizona, California, Colorado, South Dakota, and Texas. *''Carpocyon webbi'', fossils have been found in Nebraska and New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , ...
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Protepicyon
''Protepicyon'' is an extinct monospecific genus of the Borophaginae subfamily of canids native to North America. It lived during the Barstovian stage of the Middle Miocene 16.0—13.6 mya. One of the top predators of its time, it was the probable ancestor of the better known ''Epicyon ''Epicyon'' ("more than a dog") is a large, extinct, canid genus of the subfamily Borophaginae ("bone-crushing dogs"), native to North America. ''Epicyon'' existed for about from the Hemingfordian age of the Early Miocene, to the Hemphillian of ...'', and is known from remains in California and New Mexico. References zipcodezoo.comserials.cib.unibo.it
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Subtribe
Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclude ...). The early use of this word is from 19th century. An example of subtribe is Hyptidinae that contains approximately 400 accepted species distributed in 19 genera. References Botanical nomenclature Plant taxonomy Zoological nomenclature {{Botany-stub ...
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North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Caribbean Sea, and to the west and south by the Pacific Ocean. Because it is on the North American Plate, North American Tectonic Plate, Greenland is included as a part of North America geographically. North America covers an area of about , about 16.5% of Earth's land area and about 4.8% of its total surface. North America is the third-largest continent by area, following Asia and Africa, and the list of continents and continental subregions by population, fourth by population after Asia, Africa, and Europe. In 2013, its population was estimated at nearly 579 million people in List of sovereign states and dependent territories in North America, 23 independent states, or about 7.5% of the world's population. In Americas (terminology)#Human ge ...
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Early Miocene
The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene Epoch made up of two stages: the Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 Ma to 15.97 ± 0.05 Ma (million years ago). It was preceded by the Oligocene epoch. As the climate started to get cooler, the landscape started to change. New mammals evolved to replace the extinct animals of the Oligocene epoch. The first members of the hyena and weasel family started to evolve to replace the extinct ''Hyaenodon'', entelodonts and bear-dogs. The chalicotheres survived the Oligocene epoch. A new genus of entelodont called ''Daeodon'' evolved in order to adapt to the new habitats and hunt the new prey animals of the Early Miocene epoch; it quickly became the top predator of North America. But it became extinct due to competition from '' Amphicyon'', a newcomer from Eurasia. ''Amphicyon'' bested ''Daeodon'' because the bear-dog Amphicyonidae is an extinct family of terrestr ...
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Zanclean
The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, and followed by the Piacenzian Age. The Zanclean can be correlated with regionally used stages, such as the Opoitian of New Zealand, and the Tabianian or Dacian of Central Europe. It also corresponds to the late Hemphillian to mid-Blancan North American Land Mammal Ages. In California, the Zanclean roughly corresponds to the middle part of the Delmontian stage. Definition The Zanclean Stage was introduced by Giuseppe Seguenza in 1868. It is named after ''Zancle'', the pre-Roman name for the Italian city of Messina on Sicily. The base of the Zanclean (and the Pliocene Series) lies with the top of magnetic chronozone Cr3 (about 100,000 years before the Thvera normal subchronozone C3n.4n). The base is also close to the extinction level of the calc ...
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Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Neogene Period in the . The Pliocene follows the Epoch and is followed by the Epoch. Prior to the 2009 ...
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