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Paratomarctus
''Paratomarctus'' is an extinct monospecific genus of the Borophaginae subfamily of canids native to North America. It lived from the Middle to Late Miocene, 16.3—5.3 mya, existing for approximately . It was about the size of a coyote, and was probably a generalised predator, without the specialised adaptations of most later borophagines. ''Paratomarctus'' was one of the last of the Borophaginae and shared its habitat with other canids, including ''Borophagus'', ''Epicyon'', ''Carpocyon'', ''Aelurodon'', and the true canine, ''Canis lepophagus''. Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ... have been uncovered throughout most of the western United States. References zipcodezoo.comwww.sciencemag.org Borophagines Miocene carnivorans Miocene mammals of North A ...
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Borophaginae
The extinct Borophaginae form one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Hesperocyoninae and extant Caninae. Borophaginae, called "bone-crushing dogs", were endemic to North America during the Oligocene to Pliocene and lived roughly 34—2.5 million years ago, existing for about . Origin The Borophaginae descended from the subfamily Hesperocyoninae. The earliest and most primitive borophagine is the genus ''Archaeocyon'', which is a small fox-sized animal mostly found in the fossil beds in western North America. The borophagines soon diversified into several major groups. They evolved to become considerably larger than their predecessors, and filled a wide range of niches in late Cenozoic North America, from small omnivores to powerful, bear-sized carnivores, such as ''Epicyon''. Species There are 66 identified borophagine species, including 18 new ones that range from the Orellan to Blancan ages. A phylogenetic anal ...
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Epicyon
''Epicyon'' ("more than a dog") is a large, extinct, canid genus of the subfamily Borophaginae ("bone-crushing dogs"), native to North America. ''Epicyon'' existed for about from the Hemingfordian age of the Early Miocene, to the Hemphillian of the Late Miocene. ''Epicyon'' is the largest known canid of all time, with the type species reaching 2.4 m (7.9 ft) in length, 90 cm (35 in) in shoulder height and approximately 100–125 kg (220–276 lb) in body mass. The largest known humerus specimen belonged to an individual weighing up to . Description ''Epicyon'' had a massive head and powerful jaws that were well adapted for bone-crushing, with enlarged fourth premolars like some hyenas, giving its skull a lion-like shape rather than having a skull similar in shape to that of a wolf; the adaptation would have allowed ''Epicyon'' to scavenge as well as hunt, giving it access to the nutritious marrow other contemporary carnivores couldn't access. ''Epic ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Miocene Carnivorans
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the late ...
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Fossils
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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Canis Lepophagus
''Canis lepophagus'' (Latin: : 'dog'; : 'hare' or 'rabbit'; suffix : '-eating'; hence hare-eating dog) is an extinct species of canid which was endemic to much of North America during the Early Pliocene. It is notable because its lineage is proposed to have led to both wolves and coyotes. Evolution Kurten proposed that the Blancan ''C. lepophagus'' derived from smaller Miocene ''Canis'' species in North America. It then became widespread across Eurasia where it was either identical to, or closely related with, '' C. arnensis'' of Europe. Wang proposed a linear progression from ''Eucyon davisi'' to ''C. lepophagus'' to the coyote. Johnston describes ''C. lepophagus'' as having a more slender skull and skeleton than in the modern coyote. Nowak found that the early populations had small, delicate and narrowly proportioned skulls that resemble small coyotes and appear to be ancestral to ''C. latrans''. Johnson noted that some specimens found in Cita Canyon, ...
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Caninae
The Caninae, known as canines, are one of three subfamily, subfamilies found within the Canidae, canid family. The other two canid subfamilies are the extinct Borophaginae and Hesperocyoninae. The Caninae includes all living canids and their most recent fossil relatives. Their fossils were first found in North America and dated to the Oligocene era, then spreading to Asia at the end of the Miocene era, some 7 million to 8 million years ago. Taxonomy and lineage The genus ''Leptocyon'' (Greek: ''leptos'' slender + ''cyon'' dog) includes 11 species and was the first primitive canine. They were small and weighed around 2 kg. They first appeared in Sioux County, Nebraska in the Orellan era 34-32 million years ago, which was the beginning of the Oligocene. This was the same time as the appearance of the Borophaginae with whom they share features, indicating that these were two sister groups. Borophaginae skull and dentition were designed for a powerful killi ...
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Aelurodon
''Aelurodon'' is an extinct canid genus of the subfamily Borophaginae which lived from the Barstovian land mammal age () of the middle Miocene to the late Miocene epoch ().Aelurodon, Age Range and Collections, PaleoBiology Database
yu ''Aelurodon'' existed for approximately .


Description

''Aelurodon'' are a part of a clade of loosely known as "bone-crushing" or "hyena-like" dogs, that apparently descended from the earlier genera ''



Carpocyon
''Carpocyon'' is an extinct genus of the Borophaginae subfamily of canids native to North America. It lived from the Middle to the Late Miocene, 13.6 to 5.3 Ma Mya, existing for approximately . The four species in the genus varied in size, with the largest (''C. webbi'') being about the size of a wolf; all had relatively small teeth, suggesting a diet that was more omnivorous than that of other contemporary borophagines. Species *''Carpocyon compressus'' (syn. ''Cynodesmus cuspidatus''), fossils have been found in Nevada, Nebraska, Colorado, and Kansas. *''Carpocyon limosus'', fossils have been found in Nebraska, Oklahoma, and Florida. *''Carpocyon robustus'', fossils have been found in Arizona, California, Colorado, South Dakota, and Texas. *''Carpocyon webbi'', fossils have been found in Nebraska and New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , ...
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Borophagus
''Borophagus'' ("gluttonous eater") is an extinct genus of the subfamily Borophaginae, a group of canids endemic to North America from the Middle Miocene epoch through the Early Pleistocene epoch 12—1.8 Mya. Evolution ''Borophagus'', like other borophagines, are loosely known as "bone-crushing" or " hyena-like" dogs. Though not the most massive borophagine by size or weight, it had a more highly evolved capacity to crunch bone than earlier, larger genera such as ''Epicyon'', which seems to be an evolutionary trend of the group (Turner, 2004). During the Pliocene epoch, ''Borophagus'' began being displaced by other ''Canid'' species such as ''Canis edwardii'' and later by ''Aenocyon dirus''. Early species of ''Borophagus'' were placed in the genus ''Osteoborus'' until recently, but the genera are now considered synonyms. Description Typical features of this genus are a bulging forehead and powerful jaws; ''Borophagus'' has been considered to be probably a scavenger by paleonto ...
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Monotypic Taxon
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described. In contrast, an oligotypic taxon contains more than one but only a very few subordinate taxa. Examples Just as the term ''monotypic'' is used to describe a taxon including only one subdivision, the contained taxon can also be referred to as monotypic within the higher-level taxon, e.g. a genus monotypic within a family. Some examples of monotypic groups are: Plants * In the order Amborellales, there is only one family, Amborellaceae and there is only one genus, '' Amborella'', and in this genus there is only one species, namely ''Amborella trichopoda ...
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