Cyprinidae is a

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

freshwater fish Freshwater fish are those that spend some or all of their lives in fresh water, such as rivers and lakes, with a salinity of less than 1.05%. These environments differ from marine conditions in many ways, especially the difference in levels of s ...

commonly called the carp or minnow family. It includes the

carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...

s, the true

minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are ...

s, and relatives like the

barb Barb or the BARBs or ''variation'' may refer to: People * Barb (name), a list of people and fictional characters with the given name or surname * Barb, a term used by fans of Nicki Minaj to refer to themselves * The Barbs, a band Places * Barb, ...

s and

barbel Barbel may refer to: *Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles *Barbel (fish), a common name for certain species of fish **''Barbus barbus'', a species of cyprinid ...

s. Cyprinidae is the largest and most diverse fish family and the largest vertebrate animal family in general with about 3,000

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

, of which only 1,270 remain extant, divided into about 370

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...

. Cyprinids range from about 12 mm in size to the

giant barb The giant barb, Siamese giant carp, or simply Siamese carp (''Catlocarpio siamensis'') ( km, ត្រីគល់រាំង, ; th, กระโห้, , , or , , ; Vietnamese: ''cá Hô''), is the largest species of cyprinid in the world. Th ...

(''Catlocarpio siamensis''). By genus and species count, the family makes up more than two-thirds of the

ostariophysi Ostariophysi is the second-largest superorder of fish. Members of this superorder are called ostariophysians. This diverse group contains 10,758 species, about 28% of known fish species in the world and 68% of freshwater species, and are present ...

an order

Cypriniformes Cypriniformes is an order of ray-finned fish, including the carps, minnows, loaches, and relatives. Cypriniformes is an Order within the Superorder Ostariophysi consisting of "Carp-like" Ostariophysins. This order contains 11-12 familie ...

. The family name is derived from the Greek word ( 'carp').

Biology and ecology

Cyprinids are stomachless fish with toothless jaws. Even so, food can be effectively chewed by the

gill raker Gill rakers in fish are bony or cartilaginous processes that project from the branchial arch (gill arch) and are involved with suspension feeding tiny prey. They are not to be confused with the gill filaments that compose the fleshy part of th ...

s of the specialized last gill bow. These

pharyngeal teeth Pharyngeal teeth are teeth in the pharyngeal arch of the throat of cyprinids, suckers, and a number of other fish species otherwise lacking teeth.

allow the fish to make chewing motions against a chewing plate formed by a bony process of the

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...

. The pharyngeal teeth are unique to each species and are used by scientists to identify species. Strong pharyngeal teeth allow fish such as the

common carp The Eurasian carp or European carp (''Cyprinus carpio''), widely known as the common carp, is a widespread freshwater fish of eutrophic waters in lakes and large rivers in Europe and Asia.Fishbase''Cyprinus carpio'' Linnaeus, 1758/ref>Arkive The ...

and ide to eat hard baits such as

snails A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Ga ...

and

bivalves Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, biv ...

. Hearing is a well-developed sense in the cyprinids since they have the

Weberian organ The Weberian apparatus is an anatomical structure that connects the swim bladder to the auditory system in fishes belonging to the superorder Ostariophysi. When it is fully developed in adult fish, the elements of the apparatus are sometimes coll ...

, three specialized vertebral processes that transfer motion of the

gas bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth wi ...

to the inner ear. The vertebral processes of the Weberian organ also permit a cyprinid to detect changes in motion of the gas bladder due to atmospheric conditions or depth changes. The cyprinids are considered

physostome Physostomes are fishes that have a pneumatic duct connecting the gas bladder to the alimentary canal. This allows the gas bladder to be filled or emptied via the mouth. This not only allows the fish to fill their bladder by gulping air, but also t ...

s because the

pneumatic duct The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth wi ...

is retained in adult stages and the fish are able to gulp air to fill the gas bladder, or they can dispose of excess gas to the gut. Giant Barb

Cyprinids are native to North America,

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, and

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...

. The largest known cyprinid is the

(''Catlocarpio siamensis''), which may grow up to in length and in weight. Other very large species that can surpass are the

golden mahseer ''Tor putitora'', the Putitor mahseer, Himalayan mahseer, or golden mahseer, is an endangered species of cyprinid fish that is found in rapid streams, riverine pools, and lakes in the Himalayan region. Its native range is within the basins of th ...

(''Tor putitora'') and mangar (''Luciobarbus esocinus''). The largest North American species is the

Colorado pikeminnow The Colorado pikeminnow (''Ptychocheilus lucius'', formerly squawfish) is the largest cyprinid fish of North America and one of the largest in the world, with reports of individuals up to 6 ft (1.8 m) long and weighing over 100 p ...

(''Ptychocheilus lucius''), which can reach up to in length. Conversely, many species are smaller than . The smallest known fish is '' Paedocypris progenetica'', reaching at the longest. All fish in this family are egg-layers and most do not guard their eggs; however, a few species build nests and/or guard the eggs. The bitterlings of subfamily

Acheilognathinae The bitterling-like cyprinids form the cyprinid subfamily Acheilognathinae. This subfamily contains four genera, although the Khanka spiny bitterling is often placed in '' Acheilognathus'', and at least 71 described species to date. Over half of ...

are notable for depositing their eggs in bivalve

molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...

, where the young develop until able to fend for themselves. Cyprinids contain the first and only known example of androgenesis in a vertebrate, in the

Squalius alburnoides ''Squalius alburnoides'' is a species of fish in the family Cyprinidae. It is found in Portugal and Spain. Its natural habitats are rivers and intermittent rivers. It may be threatened by habitat loss. This species is a highly peculiar fish in ...

allopolyploid Polyploidy is a condition in which the cells of an organism have more than one pair of (homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ...

complex. Most cyprinids feed mainly on

invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

and

vegetation Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic charact ...

, probably due to the lack of teeth and stomach; however, some species, like the asp, are predators that specialize in fish. Many species, such as the ide and the

common rudd The common rudd (''Scardinius erythrophthalmus'') is a bentho-pelagic freshwater fish, widely spread in Europe and middle Asia, around the basins of the North, Baltic, Black, Caspian and Aral seas. Identification Morphologically, this specie ...

, prey on small fish when individuals become large enough. Even small species, such as the moderlieschen, are opportunistic predators that will eat larvae of the

common frog The common frog or grass frog (''Rana temporaria''), also known as the European common frog, European common brown frog, European grass frog, European Holarctic true frog, European pond frog or European brown frog, is a semi-aquatic amphibian ...

in artificial circumstances. Some cyprinids, such as the

grass carp The grass carp (''Ctenopharyngodon idella'') is a species of large herbivorous freshwater fish in the family Cyprinidae, native to the Pacific Far East, with a native range stretching from northern Vietnam to the Amur River on the Sino-Russi ...

, are specialized herbivores; others, such as the

common nase The common nase (''Chondrostoma nasus'') is a European potamodromous cyprinid fish. It is often simply called the nase, but that can refer to any species of its genus '' Chondrostoma''. Another name is sneep. Distribution The nase is found natur ...

, eat algae and

biofilm A biofilm comprises any syntrophic consortium of microorganisms in which cells stick to each other and often also to a surface. These adherent cells become embedded within a slimy extracellular matrix that is composed of extracellular ...

s, while others, such as the

black carp The black carp (''Mylopharyngodon piceus'') or Chinese black roach is a species of cyprinid freshwater fish and the sole extant species of the genus '' Mylopharyngodon''. It is native to lakes and rivers in East Asia, ranging from the Amur Basin ...

, specialize in snails, and some, such as the

silver carp The silver carp (''Hypophthalmichthys molitrix'') is a species of freshwater cyprinid fish, a variety of Asian carp native to China and eastern Siberia, from the Amur River drainage in the north to the Xi Jiang River drainage in the south. Al ...

, are specialized

filter feeders Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...

. For this reason, cyprinids are often introduced as a management tool to control various factors in the aquatic environment, such as aquatic vegetation and diseases transmitted by snails. Unlike most fish species, cyprinids generally increase in abundance in

eutrophic Eutrophication is the process by which an entire body of water, or parts of it, becomes progressively enriched with minerals and nutrients, particularly nitrogen and phosphorus. It has also been defined as "nutrient-induced increase in phytoplank ...

lakes. Here, they contribute towards positive feedback as they are efficient at eating the

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

that would otherwise graze on the algae, reducing its abundance.

Relationship with humans

Food

Cyprinids are highly important food fish; they are fished and

farmed Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled peop ...

across

. In

land-locked A landlocked country is a country that does not have territory connected to an ocean or whose coastlines lie on endorheic basins. There are currently 44 landlocked countries and 4 landlocked de facto states. Kazakhstan is the world's largest ...

countries in particular, cyprinids are often the major species of fish eaten because they make the largest part of

biomass Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms biom ...

in most water types except for fast-flowing rivers. In Eastern Europe, they are often prepared with traditional methods such as drying and salting. The prevalence of inexpensive frozen fish products made this less important now than it was in earlier times. Nonetheless, in certain places, they remain popular for food, as well as

recreational fishing Recreational fishing, also called sport fishing or game fishing, is fishing for leisure, exercise or competition. It can be contrasted with commercial fishing, which is professional fishing for profit; or subsistence fishing, which is fishing ...

, for ornamental use, and have been deliberately stocked in ponds and lakes for centuries for this reason.

Sport

Cyprinids are popular for angling especially for

match fishing In Britain and Ireland, coarse fishing (, ) refers to angling for rough fish, which are fish species traditionally considered undesirable as a food or game fish. Freshwater game fish are all salmonids — most particularly salmon, trout and cha ...

(due to their dominance in biomass and numbers) and fishing for common carp because of its size and strength.

As pest control

Several cyprinids have been introduced to waters outside their natural ranges to provide food, sport, or biological control for some

pest Pest or The Pest may refer to: Science and medicine * Pest (organism), an animal or plant deemed to be detrimental to humans or human concerns ** Weed, a plant considered undesirable * Infectious disease, an illness resulting from an infection ** ...

species. The common carp (''Cyprinus carpio'') and the grass carp (''Ctenopharyngodon idella'') are the most important of these, for example in

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, a ...

As a pest species

Carp in particular can stir up

sediment Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sa ...

, reducing the clarity of the water and making plant growth difficult. In America and Australia, such as the Asian carp in the

Mississippi Basin The Mississippi River is the second-longest river and chief river of the second-largest drainage system in North America, second only to the Hudson Bay drainage system. From its traditional source of Lake Itasca in northern Minnesota, it f ...

, they have become

invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...

that compete with native fishes or disrupt the environment. ''Cyprinus carpio'' is a major pest species in Australia impacting freshwater environments, amenity, and the agricultural economy, devastating

biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity' ...

by decimating native fish populations where they first became established as a major pest in the wild in the 1960s. In the major river system of eastern Australia, the Murray-Darling Basin, they constitute 80-90 per cent of fish biomass. In 2016 the federal government announced A$15.2 million to fund the National Carp Control Plan to investigate using Cyprinid herpesvirus 3 (carp virus) as a biological control agent while minimising impacts on industry and environment should a carp virus release go ahead. Despite initial, favourable assessment, in 2020 this plan was found to be unlikely to work due to the high fecundity of the fish.

Aquarium fish

Numerous cyprinids have become important in the aquarium and fishpond hobbies, most famously the

goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have b ...

, which was bred in China from the

Prussian carp The Prussian carp, silver Prussian carp or Gibel carp (''Carassius gibelio''), is a member of the family Cyprinidae, which includes many other fish, such as the common carp, goldfish, and the smaller minnows. It is a medium-sized cyprinid, and d ...

(''Carassius (auratus) gibelio''). First imported into Europe around 1728, it was much fancied by Chinese nobility as early as 1150AD and after it arrived there in 1502, also in Japan. In the latter country, from the 18th century onwards, the common carp was bred into the ornamental variety known as

koi or more specifically , are colored varieties of the Amur carp ('' Cyprinus rubrofuscus'') that are kept for decorative purposes in outdoor koi ponds or water gardens. Koi is an informal name for the colored variants of ''C. rubrofuscus'' k ...

– or more accurately , as simply means "common carp" in

Japanese Japanese may refer to: * Something from or related to Japan, an island country in East Asia * Japanese language, spoken mainly in Japan * Japanese people, the ethnic group that identifies with Japan through ancestry or culture ** Japanese diaspor ...

. Other popular aquarium cyprinids include

danionin The danionins are a group of small, minnow-type fish belonging to the family Cyprinidae. Members of this group are mostly in the genera ''Danio'', '' Devario'', and ''Rasbora''. They are primarily native to the fresh waters of South and Southeast ...

s, rasborines, and true barbs. Larger species are bred by the thousands in outdoor ponds, particularly in

Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...

, and trade in these aquarium fishes is of considerable commercial importance. The small rasborines and danionines are perhaps only rivalled by

characid Characidae, the characids or characins is a family of freshwater subtropical and tropical fish, belonging to the order Characiformes. The name "characins" is the historical one, but scientists today tend to prefer "characids" to reflect their st ...

s and poecilid livebearers in their popularity for community aquaria. One particular species of these small and undemanding danionines is the

zebrafish The zebrafish (''Danio rerio'') is a freshwater fish belonging to the minnow family (Cyprinidae) of the order Cypriniformes. Native to South Asia, it is a popular aquarium fish, frequently sold under the trade name zebra danio (and thus often c ...

(''Danio rerio''). It has become the standard

model species A model organism (often shortened to model) is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workin ...

for studying developmental genetics of

vertebrate Vertebrates () comprise all animal taxon, taxa within the subphylum Vertebrata () (chordates with vertebral column, backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the ...

s, in particular fish.

Threatened families

Habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

and other causes have reduced the wild stocks of several cyprinids to dangerously low levels; some are already entirely extinct. In particular, the cyprinids of the subfamily

Leuciscinae Leuciscinae is a subfamily of the freshwater fish family Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnows. As the name suggests, most members of the ...

from southwestern North America have been hit hard by

pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the ...

and unsustainable water use in the early to mid-20th century; most globally extinct cypriniform species are in fact leuciscinid cyprinids from the southwestern United States and northern Mexico.

Systematics

The massive diversity of cyprinids has so far made it difficult to resolve their

phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ...

in sufficient detail to make assignment to

subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

more than tentative in many cases. Some distinct lineages obviously exist – for example, the

Cultrinae The Cultrinae are one of at least 13 subfamilies of cyprinid fish. It contains ten genera. Recognized genera * ''Anabarilius'' (21 species) * '' Ancherythroculter'' (5 species) * '' Chanodichthys'' (5 species) * '' Culter'' (4 species) * '' Cul ...

and Leuciscinae, regardless of their exact delimitation, are rather close relatives and stand apart from

Cyprininae The Cyprininae are one of at least 11 subfamilies of cyprinid fish. It contains three genera in its strictest definition but many more are included depending on which authority is defining it, especially if the Labeobarbinae is not considered to ...

but the overall

systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic t ...

and

taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

of the Cyprinidae remain a subject of considerable debate. A large number of

are ''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...

'', too equivocal in their traits and/or too little-studied to permit assignment to a particular subfamily with any certainty. Part of the solution seems that the delicate rasborines are the core group, consisting of minor lineages that have not shifted far from their evolutionary niche, or have coevolved for millions of years. These are among the most basal lineages of living cyprinids. Other "rasborines" are apparently distributed across the diverse lineages of the family. The validity and circumscription of proposed subfamilies like the

Labeoninae Labeoninae is a doubtfully distinct subfamily of ray-finned fishes in the family Cyprinidae of order Cypriniformes. They inhabit fresh water and the largest species richness is in the region around southern China, but there are also species el ...

Squaliobarbinae Squaliobarbinae is a small subfamily of the carp and minnow family, Cyprinidae, which consists of three monotypic genera which have their natural distributions in eastern Asia. Two species, the grass carp (''Ctenopharyngodon idella'') and the ...

also remain doubtful, although the latter do appear to correspond to a distinct lineage. The sometimes-seen grouping of the large-headed carps ( Hypophthalmichthyinae) with ''

Xenocypris ''Xenocypris'' is a genus of cyprinid fish found in eastern Asia. There are currently seven species in this genus, one, ''X. yushensis'', being known only from fossils, and one, ''X. yunnanensis'', which is critically endangered or recently exti ...

'', though, seems quite in error. More likely, the latter are part of the Cultrinae. The entirely

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

"Barbinae" and the disputed Labeoninae might be better treated as part of the Cyprininae, forming a close-knit group whose internal relationships are still little known. The small African "

s" do not belong in ''

Barbus ''Barbus'' is a genus of ray-finned fish in the family Cyprinidae. The type species of ''Barbus'' is the common barbel, first described as ''Cyprinus barbus'' and now named ''Barbus barbus''. ''Barbus'' is the namesake genus of the subfamily B ...

'' ''

sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...

'' – indeed, they are as distant from the typical

s and the

typical carp ''Cyprinus'' is the genus of typical carps in family Cyprinidae. Most species in the genus are of East Asia origin with only the common carp (''C. carpio'') in Western Asia and Europe; this invasive species has also been introduced to many othe ...

s (''Cyprinus'') as these are from ''

Garra ''Garra'' is a genus of fish in the family Cyprinidae. These fish are one example of the " log suckers", sucker-mouthed barbs and other cyprinids commonly kept in aquaria to keep down algae. The doctor fish of Anatolia and the Middle East be ...

'' (which is placed in the Labeoninae by most who accept the latter as distinct) and thus might form another as yet unnamed subfamily. However, as noted above, how various minor lineages tie into this has not yet been resolved; therefore, such a radical move, though reasonable, is probably premature. The

tench The tench or doctor fish (''Tinca tinca'') is a fresh- and brackish-water fish of the order Cypriniformes found throughout Eurasia from Western Europe including the British Isles east into Asia as far as the Ob and Yenisei Rivers. It is als ...

(''Tinca tinca''), a significant food species farmed in western Eurasia in large numbers, is unusual. It is most often grouped with the Leuciscinae, but even when these were rather loosely circumscribed, it always stood apart. A

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...

analysis of

DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...

data of the S7

ribosomal protein A ribosomal protein (r-protein or rProtein) is any of the proteins that, in conjunction with rRNA, make up the ribosomal subunits involved in the cellular process of translation. ''E. coli'', other bacteria and Archaea have a 30S small subunit ...

intron An intron is any nucleotide sequence within a gene that is not expressed or operative in the final RNA product. The word ''intron'' is derived from the term ''intragenic region'', i.e. a region inside a gene."The notion of the cistron .e., gene ...

1 supports the view that it is distinct enough to constitute a

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

subfamily. It also suggests it may be closer to the small

East Asia East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea ...

n ''

Aphyocypris ''Aphyocypris'' is a genus of cyprinid fishes consisting of eight species, all of which are restricted to East Asia. Species There are currently ten recognized species in this genus: * ''Aphyocypris amnis'' T. Y. Liao, S. O. Kullander & H. ...

'', '' Hemigrammocypris'', and '' Yaoshanicus''. They would have diverged roughly at the same time from cyprinids of east-central Asia, perhaps as a result of the Alpide orogeny that vastly changed the

topography Topography is the study of the forms and features of land surfaces. The topography of an area may refer to the land forms and features themselves, or a description or depiction in maps. Topography is a field of geoscience and planetary s ...

of that region in the late

Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...

, when their divergence presumably occurred. A DNA-based analysis of these fish places the Rasborinae as the basal lineage with the Cyprininae as a sister clade to the Leuciscinae. The subfamilies

Gobioninae Gobioninae is a monophyletic subfamily of Eurasian cyprinid fishes. A species-rich subfamily, it is divided into five tribes: Gobionini, Pseudogobionini, Hemibarbini, Coreiini, and Sarcocheilichthyini. To adapt to different masticatory operatio ...

, and

are monophyletic.

Subfamilies and genera

Pseudogobio esocinus(Hamamatsu,Shizuoka,Japan)

The 5th Edition of

Fishes of the World ''Fishes of the World'' by the American Ichthyology, ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classificat ...

sets out the following subfamilies:

Subfamily
Acheilognathinae The bitterling-like cyprinids form the cyprinid subfamily Acheilognathinae. This subfamily contains four genera, although the Khanka spiny bitterling is often placed in '' Acheilognathus'', and at least 71 described species to date. Over half of ...

* ''

Acanthorhodeus The Khanka spiny bitterling (''Acanthorhodeus chankaensis'') is a temperate freshwater fish belonging to the Acheilognathinae subfamily of the family Cyprinidae. It originates in the inland rivers in Asia, and is found in China, Korea, and Russi ...

'' (Khanka spiny bitterling) * ''

Acheilognathus ''Acheilognathus'' is a genus of cyprinid fish native to Asia. The name is derived from the Greek ''a'', meaning "without", the Greek ''cheilos'', meaning "lip", and the Greek ''gnathos'', meaning "jaw". Morphology Fishes of this genus have l ...

'' (bitterlings) * ''

Rhodeus ''Rhodeus'' is a genus of cyprinid fish, consisting of 23 species called bitterlings. The scientific name is derived from the Greek word ', meaning "rose". Most species in the genus are restricted to Asia, but two species are found in Europe (' ...

'' (bitterlings) * ''

Tanakia ''Tanakia'' is a genus of cyprinid fish, consisting of five species that occurs in Eastern Asia. The type species is the '' Tanakia limbata''. Genus is named for Shigeho Tanaka (1878-1974), “accomplished” ichthyologist of the Imperial Univer ...

'' (bitterlings)

Subfamily Alburninae

* ''

Alburnoides ''Alburnoides'' is a genus of cyprinid fishes native to Europe and Asia. Many species are known as riffle minnows or spirlins. Species There are currently 30 recognized species in this genus: * '' Alburnoides bipunctatus'' (Bloch, 1782) * '' Alb ...

'' * ''

Alburnus ''Alburnus'' is a genus of fish in the family Cyprinidae, the carps and minnows. They are known commonly as bleaks. A group of species in the genus is known as shemayas. The genus occurs in the western Palearctic realm, and the center of diversit ...

'' (bleaks) * ''

Aspiolucius ''Aspiolucius esocinus'' or the pike asp is a species of cyprinid fish native to rivers, such as Amu Darya and Syr Darya, and occasionally lakes in Tajikistan, Kyrgyzstan, Turkmenistan, and Uzbekistan, where it was a common fish until 1980s. Afte ...

'' (pike asp) * ''

Metzia ''Metzia'' is a genus of cyprinid fish that is found in eastern Asia. The genus is named in honor of the American ichthyologist Charles William Metz of Stanford University. Species There are currently 8 recognized species in this genus: * '' M ...

Subfamily Barbinae

* '' Acrossocheilus'' * '' Anchicyclocheilus'' * '' Aulopyge'' (Dalmatian barbelgudgeon) * '' Balantiocheilos'' * ''

'' (typical barbels and barbs) * '' Carasobarbus'' * '' Clypeobarbus'' * ''

Dawkinsia ''Dawkinsia'' is a genus of cyprinid fishes from freshwater in South India and Sri Lanka. It was split off from genus ''Puntius'' in 2012. Etymology ''Dawkinsia'' is named after the evolutionary biologist Richard Dawkins in recognition of his ...

'' * ''

Desmopuntius ''Desmopuntius'' is a genus of small freshwater cyprinids native to Southeast Asia. They were formerly included in ''Puntius''. Etymology The name ''Desmopuntius'' is derived from the Greek language δεσμψτης ("desmotes") meaning 'pris ...

'' * ''

Diptychus ''Diptychus'' is a genus of cyprinid freshwater fish, consisting of two species found in Himalaya and the Tibetan Plateau of China, India, Nepal and Pakistan, ranging west to the Tien Shan Mountains and Central Asia.Li, G.; Y. Tang; R. Zhang; an ...

'' * ''

Enteromius ''Enteromius'' is a genus of small to medium-sized cyprinid fish native to tropical Africa. Most species were placed in the genus '' Barbus''. Species Species in this genus are: References External links {{Taxonbar, from=Q23072870 C ...

'' * ''

Haludaria ''Haludaria'' is a genus of cyprinids native to freshwater habitats in the Western Ghats of India. Originally the genus was named ''Dravidia'' Pethiyagoda, Meegaskumbura & Maduwage, 2012 which is preoccupied by the dipteran genus ''Dravidia'' L ...

'' * '' Hsianwenia'' * '' Kalimantania'' * '' Luciobarbus'' * '' Mesopotamichthys'' * '' Neobarynotus'' * ''

Oliotius The checker barb (''Oliotius oligolepis'')Kottelat, M. (2013)The Fishes of the Inland Waters of Southeast Asia: A Catalogue and Core Bibliography of the Fishes Known to Occur in Freshwaters, Mangroves and Estuaries. ''The Raffles Bulletin of Zool ...

'' * '' Oreichthys'' * ''

Pethia ''Pethia'' is a genus of small freshwater fish in the family Cyprinidae native to South Asia, East Asia (only Pethia stoliczkana recorded) and Mainland Southeast Asia. Some species are commonly seen in the aquarium trade. The name ''Pethia'' is ...

'' * ''

Pseudobarbus ''Pseudobarbus'' is a ray-finned fish genus in the family Cyprinidae. The type species is Burchell's redfin (''P. burchelli''). The scientific name is derived from the Ancient Greek ''pseudes'' ("false") and the Latin word ''barbus'' ("bear ...

'' (redfins) * ''

Puntigrus ''Puntigrus'' is a genus of cyprinids native to Southeast Asia. Etymology The name ''Puntigrus'' is derived from the first syllable of the cyprinid genus ''Puntius'' and "tigrus" (to allude to the Latin ''"tigris"'' meaning "tiger"). Species ...

'' * ''

Puntius ''Puntius'' is a genus of small freshwater fish in the family Cyprinidae native to South Asia and Mainland Southeast Asia, as well as Taiwan. Many species formerly placed in ''Puntius'' have been moved to other genera such as '' Barbodes'', '' ...

'' (spotted barbs) * ''

Sahyadria ''Sahyadria'' is a genus of cyprinid fishes native to India where they are endemic to rivers and streams in the Western Ghats. They were formerly included in ''Puntius''. The largest ''Sahyadria'' measure total length Fish measurement is the ...

'' * ''

Schizopyge ''Schizopyge'' is a genus of cyprinid freshwater fish found in Pakistan and the northwestern part of India. ''Schizopyge'' is closely related to ''Schizothorax ''Schizothorax'' is a genus of cyprinid fish found in southern and western China, ...

'' (snowtrouts) * ''

Schizothorax ''Schizothorax'' is a genus of cyprinid fish found in southern and western China, through northern South Asia ( Himalaya) and Central Asia, to Iran, with a single species, ''S. prophylax'', in Turkey.Yang, J.; J.X. Yang; and X.Y. Chen (2012). A ...

'' (snowtrouts) * ''

Sinocyclocheilus ''Sinocyclocheilus'' is a genus of freshwater fish in the family Cyprinidae endemic to China, where only found in Guangxi, Guizhou and Yunnan.Ma, L.; and Y.-H. Zhao (2012). Cavefish of China. Pp. 107—125 in: White, W.B.; and D.C. Cuvier, edit ...

'' (golden-line fish) * '' Striuntius'' * '' Systomus''

Subfamily
Cultrinae The Cultrinae are one of at least 13 subfamilies of cyprinid fish. It contains ten genera. Recognized genera * ''Anabarilius'' (21 species) * '' Ancherythroculter'' (5 species) * '' Chanodichthys'' (5 species) * '' Culter'' (4 species) * '' Cul ...

* ''

Anabarilius ''Anabarilius'' is a genus of ray-finned fish in the family Cyprinidae, most of them only occurring in China. Many of the species have very restricted geographic range and have been negatively impacted by introduced species, fishing, and habitat ...

'' * '' Chanodichthys'' * '' Culter'' * '' Cultrichthys'' * '' Hainania'' * '' Hemiculter'' (sharpbellies) * '' Ischikauia'' * ''

Megalobrama ''Megalobrama'' is a genus of cyprinid fish, consisting of six species native to fresh water in China and eastern Russia. The name is derived from the Greek word ''megalos'', meaning "great", and the Old French word ''breme'', a type of fres ...

'' * '' Parabramis'' (white Amur bream) * '' Paralaubuca'' * '' Pseudohemiculter'' * ''

Pseudolaubuca ''Pseudolaubuca'' is a genus of cyprinid fish from eastern Asia. There are currently four species in this genus. Species * '' Pseudolaubuca engraulis'' ( Nichols, 1925) * '' Pseudolaubuca hotaya'' Đ. Y. Mai, 1978 * '' Pseudolaubuca jouyi'' ( ...

'' * ''

Sinibrama ''Sinibrama'' is a small genus of cyprinid fishes, the five currently recognized species distributed in south China, Taiwan, Laos, and Vietnam. These are silvery fishes with deep, laterally compressed bodies, large eyes, and terminal mouths. Th ...

'' * ''

Toxabramis ''Toxabramis'' is a genus of cyprinid fish found in eastern Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares th ...

Subfamily
Cyprininae The Cyprininae are one of at least 11 subfamilies of cyprinid fish. It contains three genera in its strictest definition but many more are included depending on which authority is defining it, especially if the Labeobarbinae is not considered to ...

* '' Arabibarbus'' * ''

Barboides ''Barboides'' is a genus of very small ray-finned fish in the family Cyprinidae from freshwater habitats in West and Middle Africa. Species *''Barboides britzi'' Conway & Moritz, 2006 *''Barboides gracilis ''Barboides gracilis'' is a species ...

'' * '' Barbonymus'' (tinfoil barbs) * '' Capoeta'' (khramulyas) * ''

Carassioides ''Carassioides'' is a genus of cyprinid fish that occurs in East and Southeast Asia. There are currently four recognized species. Species *'' Carassioides acuminatus'' ( J. Richardson, 1846) *''Carassioides argentea'' V. H. Nguyễn, 2001 * ...

'' * ''

Carassius ''Carassius ''is a genus in the ray-finned fish family Cyprinidae. Most species in this genus are commonly known as crucian carps, though this term often specifically refers to '' C. carassius''. The most well known is the goldfish (''C. auratu ...

'' (Crucian carps and

) * ''

Cyprinus ''Cyprinus'' is the genus of typical carps in family Cyprinidae. Most species in the genus are of East Asia origin with only the common carp (''C. carpio'') in Western Asia and Europe; this invasive species has also been introduced to many othe ...

'' (typical carps) * ''

Eechathalakenda The Channa barb (''Eechathalakenda ophicephalus'') (formerly ''Puntius ophicephalus''), is a species of cyprinid fish endemism, endemic to India where it occurs in hill streams in forested areas. This species can reach a length of fish measureme ...

'' * '' Gymnocypris''

Subfamily Danioninae

* '' Amblypharyngodon'' (carplets) * ''

Aspidoparia ''Aspidoparia'' is a small genus of cyprinid fishes that are found in South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghan ...

'' * '' Barilius'' * ''

Betadevario ''Betadevario ramachandrani'' is a species of cyprinid fish that is found only in the upper Seetha River drainage in Karnataka, India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-la ...

'' * '' Boraras'' (rasboras) * '' Cabdio'' * ''

Chelaethiops ''Chelaethiops'' is a genus of cyprinid fish found in lakes and rivers of Africa. While commonly called "sardines", they are not related to the true sardines of the family Clupeidae. There are currently five recognized species in this genus. ...

'' * '' Chela'' * ''

Danio ''Danio'' is a genus of small freshwater fish in the family Cyprinidae found in South and Southeast Asia, commonly kept in aquaria. They are generally characterised by a pattern of horizontal stripes, rows of spots or vertical bars. Some species ...

'' (danios) * ''

Danionella ''Danionella'' is a genus of danionin fish found in freshwater habitats in Myanmar and West Bengal, India. It includes some of the smallest fishes. Distribution Four out of Five Described species of ''Danionella'' are found in Myanmar. ''D. tran ...

'' * ''

Devario ''Devario'' is a genus of fish in the family Cyprinidae native to the rivers and streams of South and Southeast Asia. These fishes have short barbels and many species having vertical or horizontal stripes. These species consume various small, aqu ...

'' * ''

Esomus ''Esomus'', or flying barbs, are freshwater fish native to South and Mainland Southeast Asia. They are closely related to the genus ''Danio'' and are distinctive for their extremely long barbels. Species * '' Esomus ahli'' Hora & Mukerji, 19 ...

'' (flying barbs) * ''

Engraulicypris ''Engraulicypris'' is a genus of fish in the family Cyprinidae endemic to Africa. In a study of mitochondrial genealogy, the species formerly included in ''Mesobola'' are not phylogenetically separated from ''Engraulicypris'' and therefore shoul ...

'' * '' Fangfangia'' * '' Horadandia'' * ''

Inlecypris ''Inlecypris jayarami'' is a danionin cyprinid from Lake Inle, Myanmar. Fishbase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfi ...

'' * ''

Laubuka ''Laubuka'' is a genus of cyprinid fish found in South and Southeast Asia. Species There are currently 14 recognized species in this genus: * ''Laubuka brahmaputraensis'' Kulabtong, Suksri & Nonpayom, 2012 Kulabtong, S., Suksri, S. & Nonpayom ...

'' * ''

Leptocypris ''Leptocypris'' is a genus of cyprinid fish found in Africa. There are currently nine recognized species in this genus. Species * '' Leptocypris crossensis'' Howes & Teugels, 1989 * '' Leptocypris guineensis'' (Daget, 1962) * '' Leptocypris ko ...

'' * '' Luciosoma'' * '' Malayochela'' * '' Microdevario'' * ''

Microrasbora ''Microrasbora'' is a genus of small fishes. The generic name means "small Rasbora", however these are more closely related to the danios than rasboras. They inhabit freshwater in Myanmar and Yunnan, China. Taxonomy ''Microrasbora'', until rece ...

'' * ''

Nematabramis ''Nematabramis'' is a genus of cyprinid freshwater fish that occur in Borneo and the Philippines. There are currently five described species in this genus. Species * '' Nematabramis alestes'' ( Seale & B. A. Bean, 1907) * '' Nematabramis borne ...

'' * ''

Neobola ''Neobola'' is a genus of cyprinid fishes found mostly in East Africa. There are currently five species in this genus. Species * '' Neobola bottegoi'' Vinciguerra, 1895 * '' Neobola fluviatilis'' ( Whitehead, 1962) (Athi Sardine) *'' Neobola ...

'' * '' Opsaridium'' * ''

Opsarius Species * ''Opsarius barna'' (Francis Buchanan-Hamilton, F. Hamilton, 1822) (''Barilius barna'') * ''Opsarius barnoides'' Decio Vinciguerra, 1890 * ''Opsarius bernatziki'' Koumans, 1937 * ''Opsarius cocsa'' (Francis Buchanan-Hamilton, F. Hamilton, 1 ...

'' * '' Paedocypris'' * ''

Pectenocypris ''Pectenocypris'' is a genus of small fish in the family Cyprinidae endemic to freshwater habitats in Borneo and Sumatra in Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between th ...

'' * '' Raiamas'' * ''

Rasbora ''Rasbora'' is a genus of fish in the family Cyprinidae. They are native to freshwater habitats in South and specimen that reputedly originated from Africa (Cameroon), but this locality is considered doubtful. They are small, up to long, altho ...

'' * ''

Rasboroides ''Rasboroides'' is a genus of small cyprinid fishes endemic to Sri Lanka. They are found in small, slow-flowing and shaded streams in the southwestern part of the island. They are essentially restricted to lowlands, although one introduced po ...

'' * ''

Rasbosoma ''Rasbosoma'' is a genus of cyprinid fish found in the Mekong Basin The Mekong or Mekong River is a trans-boundary river in East Asia and Southeast Asia. It is the world's twelfth longest river and the third longest in Asia. Its estima ...

'' (dwarf scissortail rasbora) * '' Rastrineobola'' (silver cyprinid) * '' Salmostoma'' (razorbelly minnows) * '' Securicula'' * ''

Sundadanio ''Sundadanio'' is a genus of cyprinid fishes from freshwater habitats, typically peat swamps and blackwater streams, in Borneo and Sumatra (as well as nearby smaller islands) in southeast Asia. At up to in standard length they are very small, ...

'' * ''

Trigonopoma ''Trigonopoma'' is a genus of cyprinid fish found in Southeast Asia. There are two described species in this genus. Species * '' Trigonopoma gracile'' ( Kottelat, 1991) * ''Trigonopoma pauciperforatum ''Trigonopoma'' is a genus of cyprinid ...

'' * ''

Trigonostigma ''Trigonostigma'' is a genus of small cyprinid fish found in Southeast Asia. These social, colorful freshwater fish are often kept in aquariums. Species There are currently four recognized species in this genus: * ''Trigonostigma espei'' ( Mei ...

Subfamily
Gobioninae Gobioninae is a monophyletic subfamily of Eurasian cyprinid fishes. A species-rich subfamily, it is divided into five tribes: Gobionini, Pseudogobionini, Hemibarbini, Coreiini, and Sarcocheilichthyini. To adapt to different masticatory operatio ...

* ''

Abbottina ''Abbottina'' is a genus of ray-finned fish in the family Cyprinidae, the carps and minnows. They are native to eastern Asia (China, Korea, Japan, and Vietnam).Froese, R. and D. Pauly. (Eds.''Abbottina'' species list.FishBase. 2011. The genus w ...

'' (false gudgeons) * ''

Belligobio ''Belligobio'' is a small genus of cyprinid fishes containing two recognized species endemic to China. The type species of ''Belligobio'' is ''Belligobio eristigma'', a synonym of ''Hemibarbus mylodon'', and this taxa is not currently classified ...

'' * '' Biwia'' * ''

Coreius ''Coreius'' is a genus of cyprinid fish belonging to the subfamily Gobioninae. They are known as gudgeons. This genus is endemic to freshwater habitats in China. It currently contains four recognized species. Species * '' Coreius cetopsis'' ( ...

'' * ''

Coreoleuciscus ''Coreoleuciscus'' is a genus of fish from the cyprinid family endemic to rivers A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into ...

'' * '' Gnathopogon'' * ''

Gobio ''Gobio'' is a genus of typical gudgeons, ray-finned fish in the family Cyprinidae many of which are endemics of south-eastern Europe. Members of the genus are usually small fish, rarely longer than 10 cm. Many species previously in ''Gobio ...

'' (typical gudgeons) * '' Gobiobotia'' * '' Gobiocypris'' * ''

Hemibarbus ''Hemibarbus'' is a genus of cyprinid fish found in eastern Asia. There are currently 12 recognized species in this genus. Species * '' Hemibarbus brevipennus'' P. Q. Yue, 1995 * '' Hemibarbus labeo'' ( Pallas, 1776) (Barbel steed) * ''Hemib ...

'' (steeds) * ''

Huigobio ''Huigobio'' is a genus of cyprinid fish that is found in Eastern Asia East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongoli ...

'' * '' Ladislavia'' * ''

Mesogobio ''Mesogobio'' is a genus of cyprinid fish that is found in Eastern Asia. There are currently two described species. Species There are currently two recognized species in this genus: * '' Mesogobio lachneri'' Bănărescu & Nalbant Nalbant is ...

'' * ''

Microphysogobio ''Microphysogobio'' is a genus of freshwater fish in the family Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barb ...

'' * '' Paracanthobrama'' * ''

Paraleucogobio ''Paraleucogobio notacanthus'' is a species of cyprinid fish endemism, endemic to China. It is the only member of the genus ''Paraleucogobio''. References

* Gobioninae Monotypic fish genera Cyprinid fish of Asia Freshwater fish of Chin ...

'' * '' Parasqualidus'' * '' Platysmacheilus'' * '' Pseudogobio'' * ''

Pseudopungtungia ''Pseudopungtungia'' is a genus of cyprinid fish found in Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic ...

'' * '' Pseudorasbora'' * '' Pungtungia'' * '' Rhinogobio'' * ''

Romanogobio ''Romanogobio'' is a genus of cyprinid fish found in Europe and Asia. Species Currently, 18 described species are in this genus, including the extinct ''R. antipai'': * ''Romanogobio albipinnatus'' (Lukasch, 1933) (white-finned gudgeon) * ''Rom ...

'' * ''

Sarcocheilichthys ''Sarcocheilichthys'' is a genus of cyprinid fishes found in eastern Asia. There are currently 12 species in the genus. Species * '' Sarcocheilichthys biwaensis'' K. Hosoya, 1982 * ''Sarcocheilichthys caobangensis'' V. H. Nguyễn & V. B. V ...

'' * ''

Saurogobio ''Saurogobio'' is a genus of cyprinid fish found in eastern Asia. Species There are currently 7 recognized species in this genus: * '' Saurogobio dabryi'' Bleeker, 1871 (Chinese lizard gudgeon) * '' Saurogobio dumerili'' Bleeker, 1871 * ''Saur ...

'' * '' Squalidus'' * ''

Xenophysogobio ''Xenophysogobio'' is a genus of cyprinid fish endemic to China. There are currently two described species in this genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as ...

Subfamily
Labeoninae Labeoninae is a doubtfully distinct subfamily of ray-finned fishes in the family Cyprinidae of order Cypriniformes. They inhabit fresh water and the largest species richness is in the region around southern China, but there are also species el ...

* '' Bangana'' * '' Brevibora'' * ''

Cirrhinus ''Cirrhinus'' is a genus of fish in the family Cyprinidae, the carps and minnows. Members of this genus are native to freshwater in South Asia, Indochina and southern China.Froese, R. and D. Pauly. (Eds.''Cirrhinus'' species list. FishBase. 2011 ...

'' (mud carps) * ''

Cophecheilus ''Cophecheilus'' is a genus of fish in the family Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinida ...

'' * ''

Crossocheilus ''Crossocheilus'', also known as the fringe barbs, flying foxes, or "algae eaters", is a genus of fish in the family Cyprinidae. It is distributed in China, India, Indonesia, Malaysia and Thailand in Asia. These fish occur in several types of ha ...

'' * '' Discocheilus'' * ''

Discogobio ''Discogobio'' is a genus of cyprinid fish that are found in eastern Asia. So far the 16 species have only been identified from China (11 species) and Vietnam (5 species) wherein each species is endemic to the country in which it is found. Spec ...

'' * '' Discolabeo'' * ''

'' * '' Henicorhynchus'' * ''

Horalabiosa ''Horalabiosa''Eschmeyer, W.N. (ed.) (2004) Catalog of fishes. Updated database version of January 2004., Catalog databases as made available to FishBase in January 2004. is a disputed genus of cyprinid fish found only in India. There are curre ...

'' * '' Hongshuia'' * ''

Labeo ''Labeo'' is a genus of carps in the family Cyprinidae. They are found in freshwater habitats in the tropics and subtropics of Africa and Asia. It contains the typical labeos in the subfamily Labeoninae, which may not be a valid group, howeve ...

'' (labeos) * ''

Labeobarbus ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular fo ...

'' (yellowfish) * ''

Labiobarbus ''Labiobarbus'' is a genus of cyprinid fish found in Southeast Asia. It currently contains nine described species. Species A total of 9 species have been assigned: * '' Labiobarbus cyanopareja'' ( Heckel, 1843) * '' Labiobarbus fasciatus'' ( Bl ...

'' * '' Longanalus'' * '' Osteochilichthys'' * '' Osteochilus'' * '' Paraqianlabeo'' * ''

Parasinilabeo ''Parasinilabeo'' is a genus of cyprinid fish endemic to China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ...

'' * ''

Placocheilus ''Placocheilus caudofasciatus'' is a species of cyprinid native to Laos, Vietnam and China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population e ...

'' * '' Protolabeo'' * '' Pseudocrossocheilus'' * '' Pseudogyrinocheilus'' * ''

Ptychidio ''Ptychidio'' is a genus of cyprinid fish endemic to China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly a ...

'' * '' Qianlabeo'' * ''

Rectoris ''Rectoris'' is a genus of cyprinid fish found in China and Vietnam. Species There are currently five recognized species in this genus: * ''Rectoris longibarbus ''Rectoris longibarbus'' is a species of cyprinid of the genus '' Rectoris''. ...

'' * '' Semilabeo'' * '' Sinigarra'' * '' Sinilabeo'' * '' Sinocrossocheilus'' * '' Stenorynchoacrum'' * ''

Tariqilabeo ''Tariqilabeo'' is a genus of fish in the family Cyprinidae native to Asia.Kottelat, M. (2016): On ''Gonorynchus'', ''Gonorhynchus'', ''Gonorinchus'', ''Gonorhinchus'' and ''Gonorrhynchus'', and some other names of labeonine fishes (Teleostei: G ...

'' * ''

Vinagarra ''Vinagarra'' is a genus of ray-finned fish in the family Cyprinidae endemic to Vietnam. Species There are currently 4 recognized species in this genus: * ''Vinagarra elongata'' V. H. Nguyễn & T. A. Bùi, 2010 * ''Vinagarra findolabium'' ( F. ...

'' * '' Vinalabeo''

Subfamily
Leptobarbinae ''Leptobarbus'' is a genus of cyprinid fish that are native to freshwater habitats in Southeast Asia. They are important food fish. It is the only genus in the subfamily Leptobarbinae. Leptobarbus hoevenii or "sultan fish" migrate the fresh water ...

* ''

Leptobarbus ''Leptobarbus'' is a genus of cyprinid fish that are native to freshwater habitats in Southeast Asia. They are important food fish. It is the only genus in the subfamily Leptobarbinae. Leptobarbus hoevenii or "sultan fish" migrate the fresh water ...

Subfamily
Leuciscinae Leuciscinae is a subfamily of the freshwater fish family Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnows. As the name suggests, most members of the ...

* ''

Abramis The common bream, freshwater bream, bream, bronze bream, carp bream or sweaty bream (''Abramis brama''), is a European species of freshwater fish in the family Cyprinidae. It is now considered to be the only species in the genus ''Abramis''. ...

'' (common bream) * ''

Acanthobrama ''Acanthobrama'' is a genus of ray-finned fish in the family Cyprinidae found mostly in the Near East. Species * ''Acanthobrama centisquama'' Heckel, 1843 * ''Acanthobrama hadiyahensis'' Coad, Alkahem & Behnke, 1983 * †''Acanthobrama hu ...

'' (bleaks) * ''

Achondrostoma ''Achondrostoma'' is a genus of freshwater fish in the family Cyprinidae, the carps and minnows. It is endemic to the Iberian Peninsula.Robalo, J. I., et al. (2007)Re-examination and phylogeny of the genus ''Chondrostoma'' based on mitochondrial ...

'' * ''

Anaecypris ''Anaecypris hispanica'', the Spanish minnowcarp, is a small species of ray-finned fish species in the family Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true mi ...

'' * '' Acrocheilus'' (chiselmouth) * '' Agosia'' (longfin dace) * ''

Algansea ''Algansea'' is a genus of ray-finned fish in the family Leuciscidae, distributed in the Lerma–Chapala– Grande de Santiago, Pátzcuaro, Armería, Ameca, Ayutla and Tuxpan basins in west-central Mexico (mostly Jalisco, but also surroundin ...

'' (Mexican chubs) * '' Aztecula'' (Aztec chub) * '' Ballerus'' (breams) * ''

Blicca ''Blicca bjoerkna'', alternatively called the white bream or the silver bream, is a European species of freshwater fish in the family Cyprinidae. Distribution ''Blicca bjoerkna'' is distributed across most of Europe and in adjacent Western Asia ...

'' (silver bream) * '' Campostoma'' (stonerollers) * '' Chondrostoma'' (typical nases) * ''

Chrosomus ''Chrosomus'' is a genus of small cyprinid fish found in freshwater habitats in the eastern half of the United States and Canada. There are currently seven recognized species in this genus. They have sometimes been included in ''Phoxinus''. They ...

'' (typical daces) * ''

Clinostomus ''Clinostomus'' is a genus of cyprinid fish that occur in eastern North America. There are two species in this genus. Species * ''Clinostomus elongatus'' ( Kirtland, 1840) (Redside dace) * ''Clinostomus funduloides The rosyside dace (''Clinost ...

'' (redside daces) * '' Codoma'' (ornate shiner) * ''

'' (Korean splendid dace) * '' Couesius'' (lake chub) * ''

Cyprinella ''Cyprinella'' is a genus of fish in the family Cyprinidae, the carps and minnows. They are known as the satinfin shiners. They are native to North America, and some are among the most common freshwater fish species on the eastern side of the con ...

'' (satinfin shiners) * '' Delminichthys'' * ''

Dionda ''Dionda'' is the genus of desert minnows, small fish belonging to the family Cyprinidae. They are native to fresh waters in the United States and Mexico. Their range is centered in the Rio Grande basin, but they also occur in associated systems ...

'' (desert minnows) * '' Eremichthys'' (desert dace) * ''

Ericymba ''Ericymba'' is a genus of fish in the family Cyprinidae. They are native to North America.Yang, L., Sado, T., Hirt, M.V., Pasco-Viel, E., Arunachalam, M., Li, J., Wang, X., Freyhof, J., Saitoh, K., Simons, A.M., Miya, M., He, S. & Mayden, R.L. ...

'' (longjaw minnows) * ''

Erimystax ''Erimystax'' is a genus of ray-finned fish in the family Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. C ...

'' (slender chubs) *

† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...

Evarra ''Evarra'' was a genus of small ray-finned fish in the family Cyprinidae. All species in the genus were restricted to waters in the Valley of Mexico and are now extinct due to habitat loss (their habitat dried out with remaining waters in the re ...

'' (Mexican daces) * ''

Exoglossum ''Exoglossum'' is a genus of mound-building freshwater fish of the minnow family (Cyprinidae) containing two species, commonly known as cutlip minnows, although the individual species, particularly '' Exoglossum maxillingua'', are also locally k ...

'' (cutlips minnows) * '' Gila'' (western chubs) * ''

Hemitremia The flame chub (''Hemitremia flammea'') is a species of freshwater fish in the family Cyprinidae found only in the United States. Its range broadly follows the Tennessee River from above Knoxville, Tennessee, to the mouth of the Duck River. Hist ...

'' (flame chub) * ''

Hesperoleucus The California roach (''Hesperoleucus symmetricus'') is a cyprinid fish native to western North America and abundant in the intermittent streams throughout central California. Once considered the sole member of its genus, it has recently been s ...

'' (California roach) * ''

Hybognathus ''Hybognathus'' is a genus of ray-finned fish in the family Cyprinidae. Its members are collectively known as the silvery minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the f ...

'' (silvery minnows) * ''

Hybopsis ''Hybopsis'' is a genus of cyprinid fish endemic to the United States. There are currently six described species in this genus. Species * ''Hybopsis amblops'' (Rafinesque, 1820) (Bigeye chub) * ''Hybopsis amnis'' ( C. L. Hubbs & Greene, 1951) (P ...

'' (bigeye chubs) * ''

Iberochondrostoma ''Iberochondrostoma'' is a genus of cyprinid fish endemic to the Iberian Peninsula. Species There are currently five recognized species in this genus: * ''Iberochondrostoma almacai'' ( M. M. Coelho, Mesquita & Collares-Pereira, 2005) * ''Iberoc ...

'' * '' Iberocypris'' * '' Iotichthys'' (least chub) * '' Kottelatia'' * '' Ladigesocypris'' * ''

Lavinia In Roman mythology, Lavinia ( ; ) is the daughter of Latinus and Amata, and the last wife of Aeneas. Creation It has been proposed that the character was in part intended to represent Servilia Isaurica, Emperor Augustus's first fiancée. Stor ...

'' (hitch) * ''

Lepidomeda ''Lepidomeda'' is a genus of cyprinid fish, commonly known as the spinedaces, found in western North America. Of the four known species, one is extinct and two are threatened. They appear to be fairly close to the leatherside chub and the spike ...

'' (spinedaces) * '' Leucalburnus'' * '' Leucaspius'' (moderlieschen) * '' Leuciscus'' (Eurasian daces) * ''

Leucos ''Leucos'' is a genus of fishes in the family Cyprinidae, from Southern Europe. They are carp close to the genus ''Rutilus'', and were only recently taxonomically distinguished from that genus.Bianco, P.G., Ketmaier, V. (2014)A revision of the ...

'' * ''

Luxilus ''Luxilus'' is a genus of cyprinid fish found in North America. They are commonly known as highscale shiners. There are currently nine species in the genus. Species * '' Luxilus albeolus'' ( D. S. Jordan, 1889) (white shiner) * '' Luxilus car ...

'' (highscale shiners) * '' Lythrurus'' (finescale shiners) * '' Macrhybopsis'' (blacktail chubs) * '' Margariscus'' (pearl daces) * ''

Meda Meda may refer to: Places * Meda de Mouros, a parish in Tábua Municipality, Portugal * Medas, a parish in Gondomar Municipality, Portugal * Meda-Ela, Sri Lanka * Međa (Leskovac), village in the municipality of Leskovac, Serbia * Meda, Lombard ...

'' (pikedace) * '' Moapa'' (moapa dace) * ''

Mylocheilus ''Mylocheilus caurinus'', the peamouth, peamouth chub, redmouth sucker or northwestern dace, is a species of freshwater ray-finned fish from the family Cyprinidae, the carps and minnows, that is found in western North America. It is the only sp ...

'' (peamouth) * '' Mylopharodon'' (hardhead) * '' Nocomis'' (hornyhead chubs) * '' Notemigonus'' (golden shiner) * ''

Notropis ''Notropis'' is a genus of freshwater fish in the family Cyprinidae. They are known commonly as eastern shiners.''Not ...

'' (eastern shiners) * ''

Opsopoeodus The pugnose minnow (''Opsopoeodus emiliae'') is a species of cyprinid fish found in the eastern North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered ...

'' (pugnose minnow) * ''

Oregonichthys ''Oregonichthys'' is a genus of ray-finned fish in the family Cyprinidae. Collectively known as Oregon chubs, that term can also refer to ''O. crameri'' in particular. Species * ''Oregonichthys crameri'' ( Snyder, 1908) (Oregon chub) * ''Oreg ...

'' (Oregon chubs) * '' Orthodon'' (Sacramento blackfish) * '' Pachychilon'' * ''

Parachondrostoma ''Parachondrostoma'' is a genus of cyprinid fish found mostly in Spain with one species entering France. There are currently four species in this genus. Species There are currently four recognized species in this genus: * '' Parachondrostoma ...

'' * '' Pararhinichthys'' (cheat minnow) * ''

Pelasgus In Greek mythology, Pelasgus ( grc, Πελασγός, ''Pelasgós'' means "ancient") was the eponymous ancestor of the Pelasgians, the mythical inhabitants of Greece who established the worship of the Dodonaean Zeus, Hephaestus, the Cabeiri, and o ...

'' * ''

Pelecus ''Pelecus cultratus'', commonly known as the ziege, sichel, sabre carp or sabrefish, is a cyprinid fish species from Eastern Europe and adjacent Asian regions, the only one in its genus, inhabiting the lower reaches of rivers and brackish waters ...

'' (sabre carp) * ''

Petroleuciscus ''Petroleuciscus'' is a genus of four species of ray-finned fish in the family Cyprinidae. It was usually included in ''Leuciscus'' until recently. This genus unites the Ponto-Caspian chubs and daces. Recent research has indicated that ''Petroleu ...

'' (Ponto-Caspian chubs and daces) * ''

Phenacobius ''Phenacobius'', the suckermouth minnows, is a genus of cyprinid fishes endemic to the United States. Historically the suckermouth minnow was not found as far eastward as Ohio, now they seem to be a stable species living throughout the Midwest an ...

'' (suckermouth minnows) * '' Phoxinellus'' * ''

Phoxinus ''Phoxinus'' is a genus of freshwater fish in the family (biology), family Leuciscidae of order (biology), order Cypriniformes, and the only members of the subfamily Phoxininae, or Eurasian minnows. The type species is ''Phoxinus phoxinus''. The ...

'' (Eurasian minnows and daces) * '' Pimephales'' (bluntnose minnows) * ''

Plagopterus The woundfin (''Plagopterus argentissimus'') is a species of minnow endemic to the Virgin River of the southwestern United States. It is the only species in the monotypic genus ''Plagopterus''. Description The woundfin is a small slender, silv ...

'' (woundfin) * ''

Platygobio The flathead chub (''Platygobio gracilis'') is a species of fish in the carp family, Cyprinidae. It is the only member of the monotypic genus ''Platygobio''.Froese, R. and D. Pauly, Eds. (2011)''Platygobio''.FishBase. It is native to North Ameri ...

'' (flathead chub) * '' Pogonichthys'' (splittails) * '' Protochondrostoma'' (South European nase) * ''

Pseudochondrostoma ''Pseudochondrostoma'' is a genus of cyprinid fish that occurs in the Iberian Peninsula. Species There are currently three recognized species in this genus: * ''Pseudochondrostoma duriense'' ( M. M. Coelho, 1985) * ''Pseudochondrostoma polylepis ...

'' * ''

Pseudophoxinus ''Pseudophoxinus'' is a genus of fish in the family Cyprinidae found in Western Asia. Species There are currently 29 recognized species in this genus: * '' Pseudophoxinus alii'' Küçük, 2007 (Pamphylian spring minnow) * '' Pseudophoxinus ...

'' * ''

Pteronotropis ''Pteronotropis'' is a genus of cyprinid fish endemic to the United States. Characteristics The genus is characterized by a broad dark blue stripe along the side. The genus has large anal and dorsal fins, as well as a compressed body. Species ...

'' (flagfin shiners) * ''

Ptychocheilus Pikeminnows, formerly squawfish, are cyprinid fish of the genus ''Ptychocheilus'' consisting of four species native to western North America. Voracious predators, they are considered an "undesirable" species in many waters, largely due to the spe ...

'' (pikeminnows) * '' Relictus'' (relict dace) * ''

Rhinichthys ''Rhinichthys'', known as the riffle daces, is a genus of freshwater fish in the carp family (Cyprinidae) of the order Cypriniformes. The type species is '' Rhinichthys atratulus'', the blacknose dace. ''Rhinichthys'' species range throughout No ...

'' (riffle daces, loach minnows) (including ''Tiaroga'') * '' Rhynchocypris'' (Eurasian minnows) * ''

Richardsonius ''Richardsonius'' is a genus of cyprinid fish consisting of two species native to western North America. Commonly known as redside shiners, that term is also used to refer to ''Richardsonius balteatus'' specifically. The genus is named after n ...

'' (redside shiners) * ''

Rutilus ''Rutilus'' is a genus of fish in the family Cyprinidae found in Eurasia. This genus is a widely distributed lineage of cyprinids and ranges from West Europe to East Siberia. Species In FishBase (2022), nine species are included in the genus: * ...

'' (roaches) * '' Sarmarutilus'' * ''

Scardinius ''Scardinius'' is a genus of ray-finned fish in the family Cyprinidae commonly called rudds. Locally, the name "rudd" without any further qualifiers is also used for individual species, particularly the common rudd (''S. erythrophthalmus''). The ...

'' (rudds) * ''

Semotilus ''Semotilus'' is the genus of creek chubs, ray-finned fish in the family Cyprinidae. The term "creek chub" is sometimes used for individual species, particularly the common creek chub, ''S. atromaculatus''. The creek chub species of minnows can ...

'' (creek chubs) * ''

Siphateles ''Siphateles'' is a genus of fish belonging to the family Cyprinidae, native to the Western United States. They were formerly placed in the genus '' Gila''. Species The species include: *''Siphateles alvordensis'' C. L. Hubbs & R. R. Miller, 1 ...

'' * '' Snyderichthys'' (spinedaces) * ''

Squalius ''Squalius'' is a genus of fish in the family Cyprinidae found in Europe and Asia. Hybridization is not rare in the Cyprinidae, including this genus. '' S. alburnoides'' is known to be of ancient hybrid origin, with the paternal lineage deriving ...

'' (European chubs) *

'' Stypodon'' (stumptooth minnow) * ''

Tampichthys ''Tampichthys'' is a genus of cyprinid fish endemic to east–central Mexico. They are entirely restricted to the Pánuco River basin, except ''T. ipni'' which also occurs in some other Mexican rivers that drain into the Gulf of Mexico. These ar ...

'' * ''

Telestes ''Telestes'' is a genus of cyprinid fish. It was formerly usually included in '' Leuciscus''. Species There are currently 14 recognized species in this genus:Bogutskaya, N.G., Zupancic, P., Bogut, I. & Naseka, A.M. (2012): Two new freshwater fi ...

'' * '' Tribolodon'' * ''

Tropidophoxinellus ''Tropidophoxinellus'' is a genus of cyprinid fish found in Europe. There are currently two described species. Species * '' Tropidophoxinellus hellenicus'' ( Stephanidis, 1971) * '' Tropidophoxinellus spartiaticus'' ( Schmidt-Ries, 1943) Ref ...

'' * ''

Vimba ''Vimba'' is a genus of cyprinid fish that is found in Europe and western Asia. There are currently four described species. Species * '' Vimba elongata'' (Valenciennes, 1844) (Bavarian vimba) * '' Vimba melanops'' ( Heckel, 1837) (Macedonian v ...

'' (Vimbas) * ''

Yuriria Yuriria, Guanajuato, Mexico,(), is one of 46 municipalities in the Mexican state of Guanajuato. Geography Yuriria is located in the "Bajío", or lowlands, of Guanajuato. It is bordered to the east by Salvatierra and Santiago Maravatío, to t ...

Subfamily
Tincinae Tincinae is a subfamily of freshwater ray-finned fish from the family Cyprinidae, it consists of the tench of Eurasia and the east Asian clod minnows. Tinca tinca is a freshwater tincinae fish that is found in the Danube basin Genera There ar ...

* ''

Tanichthys ''Tanichthys'' is a genus of freshwater fish in the carp family (Cyprinidae) of order Cypriniformes. They are native to rivers and streams in China and Vietnam. Until recently, the type species, ''T. albonubes,'' was the only one known. In 2001, ...

'' * '' Tinca'' Trigonostigma somphongsi

Subfamily
Squaliobarbinae Squaliobarbinae is a small subfamily of the carp and minnow family, Cyprinidae, which consists of three monotypic genera which have their natural distributions in eastern Asia. Two species, the grass carp (''Ctenopharyngodon idella'') and the ...

* '' Ctenopharyngodon'' (grass carp) * '' Mylopharyngodon'' (black carp) * ''

Squaliobarbus The barbel chub (''Squaliobarbus curriculus'') is a species of cyprinid fish found in China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population e ...

Subfamily
Xenocyprinae Xenocyprinae, is a contentious subfamily of the family Cyprinidae, the carp and minnow family, originally from eastern Asia. Genera The 5th edition of ''Fishes of the World'' assigns the follow general to this subfamily: *'' Aristichthys'' Oshi ...

* ''

Distoechodon ''Distoechodon'' is a genus of cyprinid fishes that occur in East Asia, one species endemic to mainland China and the other in both mainland China and Taiwan. Species *''Distoechodon macrophthalmus'' Y. H. Zhao, F. F. Kullander, S. O. Kullan ...

'' * '' Hypophthalmichthys'' (bighead carps) * '' Plagiognathops'' * ''

Pseudobrama ''Pseudobrama simoni'' is a species of cyprinid fish endemic to China, and the only species in its genus. It is a pseudobream species living in fresh and brackish water in the Yangtze The Yangtze or Yangzi ( or ; ) is the longest r ...

'' * ''

''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''

* '' Aaptosyax'' (giant salmon carp) * '' Acanthalburnus'' (bleaks) * '' Acanthogobio'' * '' Acapoeta'' * '' Albulichthys'' * '' Amblyrhynchichthys'' * '' Ancherythroculter'' * ''

'' * '' Araiocypris'' * '' Aspiorhynchus'' * '' Atrilinea'' * '' Barbichthys'' * ''

Barbodes ''Barbodes'' is a genus of small to medium-sized cyprinid fish native to tropical Asia. The majority of the species are from Southeast Asia. Many species are threatened and some from the Philippines (Lake Lanao) are already extinct. A survey car ...

'' * '' Barbopsis'' (Somalian blind barb) * '' Caecobarbus'' (Congo blind barb) * ''

Caecocypris ''Caecocypris basimi'', the Haditha cavefish, is a species of cyprinid fish endemic to Iraq, only occurring in aquifers near Haditha. This cavefish is the only member of its genus. The species is classed as Critically endangered, possibly extin ...

'' * ''

Candidia ''Candidia'' is a genus of small cyprinid freshwater fish endemic to Taiwan. Species There are currently two recognized species in this genus. * '' Candidia barbata'' (Regan The family name Regan, along with its cognates O'Regan, O Regan, R ...

'' * ''

Capoetobrama ''Capoetobrama kuschakewitschi'' is a cyprinid fish that is found in Central Asia, in a region shared by Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan and Uzbekistan. It is the only member of its genus. There are two recognized subspecie ...

'' * '' Catlocarpio'' * ''

Chagunius ''Chagunius'' is a genus of cyprinid fish containing three species which occur in South and Southeast Asia. Species There are currently three recognized species in this genus: * ''Chagunius baileyi ''Chagunius baileyi'' is a species of fres ...

'' * ''

Chuanchia ''Chuanchia labiosa'' is a species of cyprinid fish that is only found in the upper reaches of the Yellow River basin in the Qinghai–Tibet Plateau of China, where it mostly inhabits slow-flowing cold waters at altitudes above . It is the only ...

'' * ''

Coptostomabarbus ''Coptostomabarbus'' is a small genus of cyprinid fish containing only two Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million squar ...

'' * '' Cosmochilus'' * ''

Cyclocheilichthys ''Cyclocheilichthys'' is a genus of ray-finned fish in the family Cyprinidae containing approximately eight valid species. They are native to freshwater habitats in Southeast Asia and China. Two additional species are alternately placed in the g ...

'' * '' Cyprinion'' * '' Diplocheilichthys'' * ''

Discherodontus ''Discherodontus'' is a genus of small cyprinid fishes found in rivers and streams in Mainland Southeast Asia and Yunnan in China. Species There are currently six recognized species in this genus:Kottelat, M. (2013)The Fishes of the Inland Wate ...

'' * ''

Eirmotus ''Eirmotus'' is a genus of cyprinid fish that is found in Southeast Asia. There are currently four recognized species in this genus. Species * '' Eirmotus furvus'' H. H. Tan & Kottelat, 2008 * '' Eirmotus insignis'' H. H. Tan & Kottelat, ...

'' * '' Elopichthys'' * ''

Epalzeorhynchos ''Epalzeorhynchos'' is a small ray-finned fish genus of the family Cyprinidae. Its members are – like some other cyprinids – known as " freshwater sharks" or simply "sharks". They are, however, freshwater members of the Osteichthyes l ...

'' * '' Folifer'' * '' Gibelion'' (catla) (some authorities consider this species to belong in the genus ''Catla'') * '' Gymnodanio'' * ''

Gymnodiptychus ''Gymnodiptychus'' is a genus of cyprinid fish found in freshwater habitats in the highlands of China, Kyrgyzstan and Tajikistan.Kustareva, L.A., and A.M. Naseka (2015). Fish diversity in Kyrgyzstan: Species composition, fisheries and management ...

'' * '' Hampala'' * ''

Hemiculterella ''Hemiculterella'' is a genus of cyprinid fish that occurs in eastern Asia. There are currently three species in this genus. Species * '' Hemiculterella macrolepis'' Y. R. Chen, 1989 * '' Hemiculterella sauvagei'' Warpachowski, 1887 * '' He ...

'' * '' Hemigrammocypris'' (close to ''Aphyocypris''?) * '' Herzensteinia'' * ''

Hypselobarbus ''Hypselobarbus'' is a genus of fish in the family Cyprinidae endemic to India. Species There are currently 26 recognized species in this genus: * ''Hypselobarbus basavarajai'' Arunachalam, Chinnaraja & Mayden, 2016 Arunachalam, M., Chinnaraj ...

'' * '' Hypsibarbus'' * '' Laocypris'' * '' Lepidopygopsis'' * '' Linichthys'' * '' Lobocheilos'' * '' Longiculter'' * '' Luciobrama'' * ''

Luciocyprinus ''Luciocyprinus'' is a genus of cyprinid fish found in China and Southeast Asia. There are currently two described species in this genus. Species * ''Luciocyprinus langsoni ''Luciocyprinus langsoni'' is a species of cyprinid in the genus ' ...

'' * ''

Macrochirichthys ''Macrochirichthys macrochirus'', the long pectoral-fin minnow or giant sword minnow, is a species of cyprinid fish found in rivers and lakes in Southeast Asia (Vietnam and Thailand to the Greater Sundas) where it is used as a food fish. It is t ...

'' (long pectoral-fin minnow) * ''

Megarasbora ''Megarasbora elanga'' ( bn, এলং), commonly known as the Bengala barb is a fish of the carp family Cyprinidae, found commonly in rivers and freshwater lakes in and around Bangladesh. It reaches a maximum length of . It is a valued food fish ...

'' * '' Mekongina'' * ''

'' * ''

Mystacoleucus ''Mystacoleucus'' is a genus of cyprinid fishes that are found in Southeast Asia. There are currently eight described species in this genus. Species * '' Mystacoleucus argenteus'' ( F. Day, 1888) * '' Mystacoleucus atridorsalis'' Fowler, 1937 ...

'' * '' Naziritor'' (Zhobi mahseers) * ''

Neolissochilus ''Neolissochilus'' is a genus of fish in the family Cyprinidae native to freshwater habitats in Asia that are often grouped with the mahseers. The largest reach up to in length, but most species are much smaller. Species There are currently 28 ...

'' (''mahseers'') * ''

Nipponocypris ''Nipponocypris'' is a genus of cyprinid fish containing three extant species, one endemic to Japan, one to South Korea while the third occurs in Japan, Korea and China. A fourth, extinct species is known from Middle Pleistocene-aged freshwater ...

'' * '' Ochetobius'' * '' Onychostoma'' * '' Opsariichthys'' * ''

Oreoleuciscus ''Oreoleuciscus'' is a genus of medium to large cyprinid freshwater fish which is only found in Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered b ...

'' * ''

Osteobrama ''Osteobrama'' is a genus of cyprinid fish found in southern Asia consisting of eight species. The name is derived from the Greek word ''osteon'', meaning "bone", and the Old French word ''breme'', a type of freshwater fish. Species *''Ost ...

'' * ''

Oxygaster ''Oxygaster'' is a genus of cyprinid fishes found in Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geosche ...

'' * ''

Oxygymnocypris ''Oxygymnocypris stewartii'' is a species of cyprinid fish endemic to Tibet and occurs in the Yarlung Tsangpo River (=upper Brahmaputra) and its tributaries at altitudes above in the Qinghai-Tibet Plateau. It is the only species in its genus. ...

'' * '' Parachela'' * '' Paracrossochilus'' * '' Parapsilorhynchus'' * '' Parasikukia'' * '' Paraspinibarbus'' * '' Parator'' * ''

Parazacco ''Parazacco'' is a genus of small freshwater fish in the family Cyprinidae, native to China and Vietnam. Species According to FishBase, there are currently two recognized species in this genus, but the Catalog of Fishes considers the first as a ...

'' * '' Percocypris'' * '' Phreatichthys'' (Somalian cavefish) * '' Placogobio'' * '' Platypharodon'' * '' Pogobrama'' * '' Poropuntius'' * '' Probarbus'' * '' Procypris'' * '' Prolabeo'' * '' Prolabeops'' * '' Pseudaspius'' * ''

Ptychobarbus ''Ptychobarbus'' is a genus of cyprinid fish that is found in rivers, streams and lakes in the Himalayas and Tibetan Plateau of China, India, Nepal and Pakistan, extending into the highlands of Afghanistan. They reach up to in weight and about ...

'' * '' Puntioplites'' * '' Rasborichthys'' * '' Rohtee'' (Vatani rohtee) * '' Rohteichthys'' * '' Sanagia'' * ''

Sawbwa Chao-Pha (; Tai Ahom: 𑜋𑜧𑜨 𑜇𑜡, th, เจ้าฟ้า}, shn, ၸဝ်ႈၾႃႉ, translit=Jao3 Fa5 Jao3 Fa5, my, စော်ဘွား ''Sawbwa,'' ) was a royal title used by the hereditary rulers of the Tai peoples of ...

'' (Sawbwa barb) * '' Scaphiodonichthys'' * '' Scaphognathops'' * ''

'' (rudds) * ''

Schismatorhynchos ''Schismatorhynchos'' is an Asian genus of cyprinid freshwater fish found in India, Borneo and Sumatra. There are four species in this genus, although some authorities raise ''Nukta'' to full generic status as a monotypic genus. Species * ' ...

'' * '' Schizocypris'' (snowtrouts) * ''

Schizopygopsis ''Schizopygopsis'' is a genus of cyprinid fish. Most species are endemic to river basins in the Himalayas and Qinghai–Tibet Plateau of China, but ''S. stoliczkai'' extends into the highlands of Afghanistan, Iran, northern India, Kyrgyzstan, Pa ...

'' (snowtrouts) * '' Semiplotus'' * '' Sikukia'' * '' Spinibarbus'' * '' Thryssocypris'' * ''

Thynnichthys ''Thynnichthys'' is a genus of cyprinid fish found in southern Asia from India to Borneo. There are currently four species in this genus. Species * '' Thynnichthys polylepis'' Bleeker, 1860 * '' Thynnichthys sandkhol'' (Sykes, 1839) (Sandkhol c ...

'' * '' Tor'' (''

mahseer Mahseer is the common name used for the genera '' Tor'', '' Neolissochilus'', '' Naziritor'' and '' Parator'' in the family Cyprinidae (carps). The name is, however, more often restricted to members of the genus ''Tor''.Sen TK, Jayaram KC, 1982 ...

s'') * '' Troglocyclocheilus'' * ''

'' * '' Typhlobarbus'' * '' Typhlogarra'' (Iraq blind barb) * '' Xenobarbus'' * '' Xenocyprioides'' * '' Zacco'' With such a large and diverse family the taxonomy and phylogenies are always being worked on so alternative classifications are being created as new information is discovered, for example:

Phylogeny

Subfamily Probarbinae * '' Catlocarpio'' * '' Probarbus'' Subfamily

* Tribe Parapsilorhynchini ** '' Diplocheilichthys'' ** '' Neorohita'' ** '' Parapsilorhynchus'' ** '' Longanalus'' ** '' Protolabeo'' ** '' Sinilabeo'' * Tribe

Labeonini Labeoninae is a doubtfully distinct subfamily of ray-finned fishes in the family Cyprinidae of order Cypriniformes. They inhabit fresh water and the largest species richness is in the region around southern China, but there are also species e ...

** '' Bangana'' ** ''

'' (mud carps) ** '' Decourus'' ** '' Gymnostomus'' ** '' Incisilabeo'' ** ''

'' (labeos) ** '' Speolabeo'' ** ''

'' * Tribe Garrini ** ''

'' ** '' Paracrossocheilus'' ** ''

'' ** '' Osteochilus'' clade *** '' Barbichthys'' *** ''

'' *** ''

'' *** '' Henicorhynchus'' *** ''

'' *** '' Lobocheilos'' *** '' Osteochilus'' *** ''

'' ** '' Semilabeo'' clade *** ''

Ageneiogarra ''Garra'' is a genus of fish in the family Cyprinidae. These fish are one example of the " log suckers", sucker-mouthed barbs and other cyprinids commonly kept in aquaria to keep down algae. The doctor fish of Anatolia and the Middle East be ...

'' *** '' Altigena'' *** ''

'' *** ''

'' *** '' Hongshuia'' *** '' Linichthys'' *** '' Mekongina'' *** '' Paraqianlabeo'' *** ''

'' *** ''

'' *** '' Prolixicheilus'' *** '' Pseudocrossocheilus'' *** '' Pseudogyrinocheilus'' *** ''

'' *** '' Qianlabeo'' *** ''

'' *** '' Semilabeo'' *** '' Sinigarra'' *** '' Sinocrossocheilus'' *** '' Stenorynchoacrum'' Subfamily Torinae * '' Acapoeta'' * '' Arabibarbus'' * '' Barbopsis'' (Somalian blind barb) * '' Carasobarbus'' * ''

'' * ''

'' (yellowfish) * '' Lepidopygopsis'' * '' Mesopotamichthys'' * '' Naziritor'' (Zhobi mahseers) * ''

'' (

s) * '' Osteochilichthys'' * ''

Pterocapoeta ''Pterocapoeta maroccana'' is a species of cyprinid fish. It is found only in Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Se ...

'' * '' Sanagia'' * '' Tor'' (

s) * ''

'' Subfamily Smiliogastrinae * ''

'' * ''

'' * '' Caecobarbus'' (Congo blind barb) * ''

'' * '' Clypeobarbus'' * ''

'' * ''

'' * '' Hampala'' * ''

'' * '' Oreichthys'' * ''

'' * ''

'' * '' Prolabeo'' * '' Prolabeops'' * ''

'' (redfins) * ''

'' * ''

'' (spotted barbs) * '' Rohtee'' (Vatani rohtee) * ''

'' * '' Striuntius'' * '' Systomus'' * '' Xenobarbus'' Subfamily

Barbinae">ncl. Barbinae * Tribe Cyprinini ** '' Aaptosyax'' (giant salmon carp) ** ''

'' ** ''

'' (Crucian carps and

) ** ''

'' (typical carps) ** ''

'' ** '' Paraspinibarbus'' ** '' Parator'' ** '' Procypris'' ** ''

Pseudosinocyclocheilus ''Sinocyclocheilus jinxiensis'' is a species of cavefish in the family (biology), family Cyprinidae endemism, endemic to Xiaolong Spring in Jinxi County, Guangxi, China. In 2016, it was proposed that this species should be moved to its own genus ( ...

'' ** '' Sinibarbus'' ** ''

'' (golden-line fish) ** '' Typhlobarbus'' * Tribe Rohteichthyini ** '' Albulichthys'' ** '' Amblyrhynchichthys'' ** '' Anematichthys'' ** '' Balantiocheilos'' ** '' Barbonymus'' (tinfoil barbs) ** '' Cosmochilus'' ** ''

'' ** '' Cyclocheilos'' ** ''

'' ** ''

'' ** '' Hypsibarbus'' ** '' Kalimantania'' ** '' Laocypris'' ** ''

'' ** '' Parasikukia'' ** '' Poropuntius'' ** '' Puntioplites'' ** '' Rohteichthys'' ** ''

'' (Sawbwa barb) ** '' Scaphognathops'' ** '' Sikukia'' ** '' Troglocyclocheilus'' * Tribe Acrossocheilini ** '' Acrossocheilus'' ** '' Folifer'' ** '' Onychostoma'' * Tribe Spinibarbini ** '' Spinibarbus'' ** '' Spinibarbichthys'' * Tribe Schizothoracini ** '' Aspiorhynchus'' ** '' Percocypris'' ** ''

'' (snowtrouts) ** ''

'' (snowtrouts) * Tribe Schizopygopsini ** ''

'' ** ''

'' ** '' Gymnocypris'' ** ''

'' ** '' Oreinus'' ** ''

'' ** '' Platypharodon'' ** ''

'' ** ''

'' (snowtrouts) * Tribe Barbini ** '' Aulopyge'' (Dalmatian barbelgudgeon) ** ''

'' (typical barbels and barbs) ** '' Hsianwenia'' ** ''

'' ** '' Capoeta'' (khramulyas) ** '' Cyprinion'' ** '' Kantaka'' ** '' Luciobarbus'' ** '' Scaphiodonichthys'' ** '' Schizocypris'' (snowtrouts) ** '' Semiplotus'' Subfamily Danioninae * Tribe Paedocypridini ** '' Paedocypris'' * Tribe Sundadanionini ** '' Fangfangia'' ** ''

'' * Tribe Rasborini ** '' Amblypharyngodon'' (carplets) ** '' Boraras'' (rasboras) ** '' Brevibora'' ** '' Horadandia'' ** '' Kottelatia'' ** ''

'' ** ''

'' (dwarf scissortail rasbora) ** ''

'' ** ''

'' * Tribe Danionini ** ''

'' ** ''

Brachydanio The danionins are a group of small, minnow-type fish belonging to the family Cyprinidae. Members of this group are mostly in the genera ''Danio'', '' Devario'', and '' Rasbora''. They are primarily native to the fresh waters of South and Southe ...

'' ** '' Celestichthys'' ** '' Chela'' ** ''

'' (danios) ** ''

'' ** ''

'' ** '' Microdevario'' ** ''

'' * Tribe Chedrini ** '' Barilius'' ** '' Bengala'' ** '' Cabdio'' [''

''] ** ''

'' ** ''

'' (flying barbs) ** ''

'' ** '' Luciosoma'' ** '' Malayochela'' ** ''

'' ** ''

'' ** '' Opsaridium'' ** ''

'' ** '' Raiamas'' ** '' Rastrineobola'' (silver cyprinid) ** '' Salmostoma'' (razorbelly minnows) ** '' Securicula'' ** '' Thryssocypris'' Subfamily

Leptobarbinae ''Leptobarbus'' is a genus of cyprinid fish that are native to freshwater habitats in Southeast Asia. They are important food fish. It is the only genus in the subfamily Leptobarbinae. Leptobarbus hoevenii or "sultan fish" migrate the fresh water ...

* ''

Subfamily Xenocypridinae [incl.

] * Tribe Squaliobarbini ** ''

'' * Tribe Opsariichthyini ** ''

'' ** ''

'' ** '' Opsariichthys'' ** ''

'' ** '' Xenocyprioides'' * Tribe Oxygastrini ** ''

'' ** '' Araiocypris'' ** '' Gymnodanio'' ** '' Hemigrammocypris'' ** ''

'' (long pectoral-fin minnow) ** ''

'' ** ''

'' ** '' Parachela'' ** '' Paralaubuca'' ** '' Rasborichthys'' * Tribe Hypophthalmichthyini ** '' Atrilinea'' ** '' Ctenopharyngodon'' (grass carp) ** '' Elopichthys'' ** '' Hypophthalmichthys'' (bighead carps) ** '' Luciobrama'' ** '' Mylopharyngodon'' (black carp) ** '' Ochetobius'' * Tribe Xenocypridini ** Subtribe Xenocypridina *** ''

'' *** '' Plagiognathops'' *** ''

'' *** ''

'' ** Subtribe Cultrina *** ''

'' *** '' Chanodichthys'' *** '' Culter'' *** '' Ischikauia'' *** '' Longiculter'' *** ''

'' *** '' Parabramis'' (white Amur bream) *** '' Pogobrama'' *** ''

'' ** '' Hemiculter'' clade *** '' Hainania'' *** '' Hemiculter'' (sharpbellies) *** '' Pseudohemiculter'' *** ''

'' *** ''

'' Subfamily

Tincinae Tincinae is a subfamily of freshwater ray-finned fish from the family Cyprinidae, it consists of the tench of Eurasia and the east Asian clod minnows. Tinca tinca is a freshwater tincinae fish that is found in the Danube basin Genera There ar ...

* '' Tinca'' Subfamily

( bitterlings) * ?''

'' (Khanka spiny bitterling) * ''

'' * ''

Paratanakia ''Tanakia himantegus'' is a species of fishes in the family Cyprinidae found in China and Taiwan. Subspecies *''T. h. chii'' (''Paratanakia himantegus chii'') (Miao, 1934):China mainland and Taiwan *''T. h. himantegus'' (''Paratanakia himantegu ...

'' * ''

Pseudorhodeus The Tokyo bitterling (''Tanakia tanago'') is a temperate freshwater fish of the carp family (Cyprinidae). Taxonomically, it belongs to the subfamily Acheilognathinae. The species was first described as ''Rhodeus tanago'' by Shigeho Tanaka in 190 ...

'' * ''

'' Subfamily

* ''

''-'' Squalidus'' clade ** ''

'' ** ''

'' (steeds) ** '' Squalidus'' * Tribe Gobionini ** Subtribe Gobiobotiina *** '' Gobiobotia'' *** ''

'' ** Subtribe Gobionina *** ''

'' (typical gudgeons) *** ''

'' *** ''

'' *** '' Acanthogobio'' ** Subtribe Armatogobionina *** ''

'' (false gudgeons) *** '' Biwia'' *** ?''

'' *** ''

'' *** '' Platysmacheilus'' *** '' Pseudogobio'' *** ''

'' * Tribe Sarcocheilichthyini ** ''

'' ** ''

'' (Korean splendid dace) ** '' Gnathopogon'' ** '' Gobiocypris'' ** '' Ladislavia'' ** '' Paracanthobrama'' ** ''

Paraleucogobio ''Paraleucogobio notacanthus'' is a species of cyprinid fish endemism, endemic to China. It is the only member of the genus ''Paraleucogobio''. References

* Gobioninae Monotypic fish genera Cyprinid fish of Asia Freshwater fish of Chin ...

'' ** ?'' Parasqualidus'' ** ''

'' ** '' Pseudorasbora'' ** '' Pungtungia'' ** '' Rhinogobio'' ** ''

'' Subfamily Tanichthyinae * ''

'' Subfamily

Alburninae">ncl. Alburninae * Tribe Phoxinini ** ''

'' ** ''

'' (Eurasian minnows and daces) ** '' Pseudaspius'' * Tribe Laviniini ** Subtribe Chrosomina *** ''

'' (typical daces) ** Subtribe Laviniina *** '' Eremichthys'' (desert dace) *** '' Gila'' (western chubs) *** ''

'' (California roach) *** '' Klamathella'' *** ''

'' (hitch) *** '' Mylopharodon'' (hardhead) *** '' Orthodon'' (Sacramento blackfish) *** ''

'' (pikeminnows) *** '' Relictus'' (relict dace) *** ''

'' * Tribe Leuciscini ** '' Pachychilon'' clade *** '' Pachychilon'' ** ''

'' clade *** ''

'' ** Primitive Leuciscine clade *** '' Delminichthys'' *** '' Leucalburnus'' *** '' Notemigonus'' (golden shiner) *** ''

'' ** Subtribe Leuciscina *** ''

'' (pike asp) *** '' Leuciscus'' (Eurasian daces) *** ''

'' (sabre carp) ** Subtribe Abramina *** ''

'' (common bream) *** ''

'' (bleaks) *** ''

'' *** '' Mirogrex'' *** ''

'' (Vimbas) ** Subtribe Chondrostomina *** ''

'' *** ''

'' (bleaks) *** ''

'' *** '' Chondrostoma'' (typical nases) *** ''

'' *** '' Leucaspius'' (moderlieschen) *** ''

'' *** ''

'' (Ponto-Caspian chubs and daces) *** '' Phoxinellus'' *** '' Protochondrostoma'' (South European nase) *** ''

'' *** ''

'' (roaches) *** '' Sarmarutilus'' *** ''

'' (rudds) *** ''

'' (European chubs) *** ''

'' *** ''

'' * Tribe Plagiopterini ** '' Couesius'' (lake chub) ** ''

'' (flame chub) ** ''

'' (spinedaces) ** '' Margariscus'' (pearl daces) ** ''

'' (pikedace) ** ''

'' (woundfin) ** '' Rhynchocypris'' (Eurasian minnows) ** ''

'' (creek chubs) **

'' Stypodon'' (stumptooth minnow) * Tribe Pogonichthyini ** Subtribe Pogonichthyina *** ''

'' (redside daces) *** '' Iotichthys'' (least chub) *** ''

'' (peamouth) *** '' Pogonichthys'' (splittails) *** ''

'' (redside shiners) ** Subtribe Exoglossina *** ''

'' (cutlips minnows) *** ''

'' (Oregon chubs) *** '' Pararhinichthys'' (cheat minnow) *** ''

'' (riffle daces, loach minnows) *** '' Tiaroga'' ** Subtribe Campostomina *** '' Campostoma'' (stonerollers) *** '' Nocomis'' (hornyhead chubs) ** Subtribe Hybognathina *** '' Agosia'' (longfin dace) *** '' Alburnops'' *** ''

'' (Mexican chubs) *** ?'' Aztecula'' (Aztec chub) *** ?'' Ballerus'' (breams) *** ?''

'' (silver bream) *** '' Codoma'' (ornate shiner) *** ''

'' (satinfin shiners) *** ''

'' (desert minnows) *** ?''

'' (longjaw minnows) *** ''

Erimonax Spotfin chub (''Cyprinella monacha'') is a ray-finned fish in the family Cyprinidae that is endemic to the Tennessee River watershed. Its other common names include turquoise shiner and chromium shiner. Appearance and anatomy Spotfin chub have an ...

'' *** ''

'' (slender chubs) ***

'' (Mexican daces) *** '' Graodus'' *** '' Hudsonius'' *** ''

'' (silvery minnows) *** ''

'' (bigeye chubs) *** ?'' Iberocypris'' *** ?'' Ladigesocypris'' *** ''

'' (highscale shiners) *** '' Lythrurus'' (finescale shiners) *** '' Macrhybopsis'' (blacktail chubs) *** '' Miniellus'' *** ?'' Moapa'' (moapa dace) *** ''

Notropis ''Notropis'' is a genus of freshwater fish in the family Cyprinidae. They are known commonly as eastern shiners.''Not ...

'' (eastern shiners) *** ''

'' (pugnose minnow) *** ''

'' (suckermouth minnows) *** '' Pimephales'' (bluntnose minnows) *** ''

'' (flathead chub) *** ''

'' (flagfin shiners) *** ?'' Snyderichthys'' (spinedaces) *** ''

'' *** ?'' Tribolodon'' *** ?''

''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''

* '' Acanthalburnus'' (bleaks) * '' Acrocheilus'' (chiselmouth) * '' Ancherythroculter'' * '' Anchicyclocheilus'' * '' Gibelion'' (catla) (some authorities consider this species to belong in the genus ''Catla'') * '' Cultrichthys'' * '' Discocheilus'' * '' Discolabeo'' * ''

'' * '' Herzensteinia'' * ''

'' * ''

'' * '' Neobarynotus'' * '' Paracrossochilus'' * '' Phreatichthys'' (Somalian cavefish) * '' Placogobio'' * ''

'' (rudds) * ''

'' * '' Typhlogarra'' (Iraq blind barb) * '' Zacco''

References

External links

{{Authority control Ray-finned fish families Fish of North America Fish of Asia Fish of Europe Fish of Africa Extant Eocene first appearances

Biology and ecology

Relationship with humans

Food

Sport

As pest control

As a pest species

Aquarium fish

Threatened families

Systematics

Subfamilies and genera

Subfamily Acheilognathinae The bitterling-like cyprinids form the cyprinid subfamily Acheilognathinae. This subfamily contains four genera, although the Khanka spiny bitterling is often placed in '' Acheilognathus'', and at least 71 described species to date. Over half of ...

Subfamily Alburninae

Subfamily Barbinae

Subfamily Cultrinae The Cultrinae are one of at least 13 subfamilies of cyprinid fish. It contains ten genera. Recognized genera * ''Anabarilius'' (21 species) * '' Ancherythroculter'' (5 species) * '' Chanodichthys'' (5 species) * '' Culter'' (4 species) * '' Cul ...

Subfamily Cyprininae The Cyprininae are one of at least 11 subfamilies of cyprinid fish. It contains three genera in its strictest definition but many more are included depending on which authority is defining it, especially if the Labeobarbinae is not considered to ...

Subfamily Danioninae

Subfamily Gobioninae Gobioninae is a monophyletic subfamily of Eurasian cyprinid fishes. A species-rich subfamily, it is divided into five tribes: Gobionini, Pseudogobionini, Hemibarbini, Coreiini, and Sarcocheilichthyini. To adapt to different masticatory operatio ...

Subfamily Labeoninae Labeoninae is a doubtfully distinct subfamily of ray-finned fishes in the family Cyprinidae of order Cypriniformes. They inhabit fresh water and the largest species richness is in the region around southern China, but there are also species el ...

Subfamily Leptobarbinae ''Leptobarbus'' is a genus of cyprinid fish that are native to freshwater habitats in Southeast Asia. They are important food fish. It is the only genus in the subfamily Leptobarbinae. Leptobarbus hoevenii or "sultan fish" migrate the fresh water ...

Subfamily Leuciscinae Leuciscinae is a subfamily of the freshwater fish family Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnows. As the name suggests, most members of the ...

Subfamily Tincinae Tincinae is a subfamily of freshwater ray-finned fish from the family Cyprinidae, it consists of the tench of Eurasia and the east Asian clod minnows. Tinca tinca is a freshwater tincinae fish that is found in the Danube basin Genera There ar ...

Subfamily Squaliobarbinae Squaliobarbinae is a small subfamily of the carp and minnow family, Cyprinidae, which consists of three monotypic genera which have their natural distributions in eastern Asia. Two species, the grass carp (''Ctenopharyngodon idella'') and the ...

Subfamily Xenocyprinae Xenocyprinae, is a contentious subfamily of the family Cyprinidae, the carp and minnow family, originally from eastern Asia. Genera The 5th edition of ''Fishes of the World'' assigns the follow general to this subfamily: *'' Aristichthys'' Oshi ...

''Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...''

Phylogeny

''Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...''

See also

References

External links

Subfamily
Acheilognathinae The bitterling-like cyprinids form the cyprinid subfamily Acheilognathinae. This subfamily contains four genera, although the Khanka spiny bitterling is often placed in '' Acheilognathus'', and at least 71 described species to date. Over half of ...

Subfamily
Cultrinae The Cultrinae are one of at least 13 subfamilies of cyprinid fish. It contains ten genera. Recognized genera * ''Anabarilius'' (21 species) * '' Ancherythroculter'' (5 species) * '' Chanodichthys'' (5 species) * '' Culter'' (4 species) * '' Cul ...

Subfamily
Cyprininae The Cyprininae are one of at least 11 subfamilies of cyprinid fish. It contains three genera in its strictest definition but many more are included depending on which authority is defining it, especially if the Labeobarbinae is not considered to ...

Subfamily
Gobioninae Gobioninae is a monophyletic subfamily of Eurasian cyprinid fishes. A species-rich subfamily, it is divided into five tribes: Gobionini, Pseudogobionini, Hemibarbini, Coreiini, and Sarcocheilichthyini. To adapt to different masticatory operatio ...

Subfamily
Labeoninae Labeoninae is a doubtfully distinct subfamily of ray-finned fishes in the family Cyprinidae of order Cypriniformes. They inhabit fresh water and the largest species richness is in the region around southern China, but there are also species el ...

Subfamily
Leptobarbinae ''Leptobarbus'' is a genus of cyprinid fish that are native to freshwater habitats in Southeast Asia. They are important food fish. It is the only genus in the subfamily Leptobarbinae. Leptobarbus hoevenii or "sultan fish" migrate the fresh water ...

Subfamily
Leuciscinae Leuciscinae is a subfamily of the freshwater fish family Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnows. As the name suggests, most members of the ...

Subfamily
Tincinae Tincinae is a subfamily of freshwater ray-finned fish from the family Cyprinidae, it consists of the tench of Eurasia and the east Asian clod minnows. Tinca tinca is a freshwater tincinae fish that is found in the Danube basin Genera There ar ...

Subfamily
Squaliobarbinae Squaliobarbinae is a small subfamily of the carp and minnow family, Cyprinidae, which consists of three monotypic genera which have their natural distributions in eastern Asia. Two species, the grass carp (''Ctenopharyngodon idella'') and the ...

Subfamily
Xenocyprinae Xenocyprinae, is a contentious subfamily of the family Cyprinidae, the carp and minnow family, originally from eastern Asia. Genera The 5th edition of ''Fishes of the World'' assigns the follow general to this subfamily: *'' Aristichthys'' Oshi ...

''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''

''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''