Lexovisaurus
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''Lexovisaurus'' is a genus of
stegosaur Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Euro ...
from mid-to-Late
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, 165.7-164.7  mya. Fossils of limb bones and armor fragments have been found in middle to late Jurassic-aged strata of
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...
.


Discovery and taxonomy

In the early 1880s collector
Alfred Nicholson Leeds Alfred Nicholson Leeds (9 March 184725 August 1917) was an English amateur palaeontologist. Biography Leeds was born at Eyebury, Peterborough, the youngest of the eight children of Edward Thurlow Leeds (180251) and Eliza Mary Leeds (née Nichols ...
acquired a skeleton of a dinosaur excavated at a small brick pit at the hamlet of Tanholt, close to Eye,
Cambridgeshire Cambridgeshire (abbreviated Cambs.) is a county in the East of England, bordering Lincolnshire to the north, Norfolk to the north-east, Suffolk to the east, Essex and Hertfordshire to the south, and Bedfordshire and Northamptonshire to t ...
. In September 1885 the remains were shown to paleontologist Henry Woodward whose notes form the first documentation on the subject. Later it was mistakenly assumed the find had been made at the industrial brick pits at
Fletton Fletton is an area of the city of Peterborough, in the Peterborough district, in the ceremonial county of Cambridgeshire, England, south of the River Nene. Notable for its large brickworks, the area has given its name to "Fletton bricks", Admin ...
, the usual source of Leeds' specimens.Leslie F. Noè, Jeff J. Liston and Sandra D. Chapman, 2010, "‘Old bones, dry subject’: the dinosaurs and pterosaur collected by Alfred Nicholson Leeds of Peterborough, England", ''Geological Society, London, Special Publications'' 343: 49-77 In 1887 the fossil was described by
John Whitaker Hulke John Whitaker Hulke FRCS FRS FGS (6 November 1830 – 19 February 1895) was a British surgeon, geologist and fossil collector. He was the son of a physician in Deal, who became a Huxleyite despite being deeply religious. Hulke became Huxley's ...
and named as a new species of the stegosaurian ''
Omosaurus ''Omosaurus'' is a dubious genus of extinct crurotarsan reptile, possibly a phytosaur, from the Late Triassic (Carnian) of North Carolina. Only scant remains are known, which makes ''Omosaurus'' hard to classify. The type, and only specie ...
'': ''Omosaurus durobrivensis''. The specific name referred to the old Roman town of Durobrivae. On 30 May 1892 the specimen was bought by the
British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum ...
. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, BMNH R1989, was found in the
Peterborough Member Peterborough () is a cathedral city in Cambridgeshire, east of England. It is the largest part of the City of Peterborough unitary authority district (which covers a larger area than Peterborough itself). It was part of Northamptonshire until ...
of the Oxford Clay Formation, more especially the '' Kosmoceras jason''
biozone In biostratigraphy, biostratigraphic units or biozones are intervals of geological strata that are defined on the basis of their characteristic fossil taxa, as opposed to a lithostratigraphic unit which is defined by the lithological properties ...
dating from the middle
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
. Hulke mistakenly assumed a provenance from the younger
Kimmeridge Clay Formation The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for Nort ...
. It consists of a sacrum of five vertebrae, and two ilia. Other bones were referred to the species, among them two plates thought to be part of the dermal armour. However, on 22 August 1888
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among ...
visited Leeds' collection at Eyebury and recognised these elements as belonging to a giant fish, in 1889 by
Arthur Smith Woodward Sir Arthur Smith Woodward, FRS (23 May 1864 – 2 September 1944) was an English palaeontologist, known as a world expert in fossil fish. He also described the Piltdown Man fossils, which were later determined to be fraudulent. He is not relate ...
named ''
Leedsichthys ''Leedsichthys'' is an extinct genus of pachycormid fish that lived in the oceans of the Middle to Late Jurassic.Liston, JJ (2004). An overview of the pachycormiform ''Leedsichthys''. In: Arratia G and Tintori A (eds) Mesozoic Fishes 3 - Syste ...
''. The plates are in fact part of the latter's skull roof. In 1915 ''Omosaurus durobrivensis'' was renamed ''Dacentrurus durobrivensis'', as the name ''Omosaurus'' had been preoccupied, which had already been indicated by Marsh in the 1870s. In 1957 the French palaeontologist
Robert Hoffstetter Robert Julien Hoffstetter (11 June 1908 in Fargniers – 29 December 1999 in Gennevilliers) was a French taxonomist and herpetologist who was influential in categorizing reptiles Reptiles, as most commonly defined are the animals in the clas ...
created a separate genus for the species: ''Lexovisaurus''. The generic name is derived from the
Lexovii The Lexovii (Gaulish: *''Lexsouioi'', 'the leaning, lame'), were a Gallic tribe dwelling immediately west of the mouth of the Seine, around present-day Lisieux, during the Iron Age and the Roman period. Name They are mentioned as ''Lexovii'' (v ...
, a Gallic tribe in ancient times inhabiting the region of
Normandy Normandy (; french: link=no, Normandie ; nrf, Normaundie, Nouormandie ; from Old French , plural of ''Normant'', originally from the word for "northman" in several Scandinavian languages) is a geographical and cultural region in Northwestern ...
, where several stegosaurian specimens had been discovered which by Hoffstetter were referred to ''Lexovisaurus''. While the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
remains ''Omosaurus durobrivensis'', the ''
combinatio nova ''Combinatio nova'', abbreviated ''comb. nov.'' (sometimes ''n. comb.''), is Latin for "new combination". It is used in taxonomic biology literature when a new name is introduced based on a pre-existing name. The term should not to be confused w ...
'' is ''Lexovisaurus durobrivensis''. Hofstetter also referred a much more complete stegosaurian skeleton in 1901 discovered by Leeds in the Fletton brick pit, specimen BMNH R3167 that in 1911 had been named ''Stegosaurus priscus'', placed as a stegosaurine.Hoffstetter, R., 1957, "Quelques observations sur les Stégosaurinés", ''Bulletin du Muséum National d’Histoire Naturelle, Paris, 2nde série'' 29: 537-547 Subsequently, in 1964,
Oskar Kuhn Oskar Kuhn (7 March 1908, Munich – 1990) was a German palaeontologist. Life and career Kuhn was educated in Dinkelsbühl and Bamberg and then studied natural science, specialising in geology and paleontology, at the University of Munich, f ...
referred the ''nomen nudum'' "Omosaurus leedsi" Seeley vide Huene 1901 to ''Lexovisaurus'' as ''Lexovisaurus leedsi''. In 1983,
Peter Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosa ...
renamed ''Omosaurus vetustus'' Huene 1910 into ''Lexovisaurus vetustus''. However, in 2008 Susannah Maidment and colleagues concluded that the holotype of ''Lexovisaurus'', BMNH R1989, was undiagnostic, so they split off BMNH R3167 and the French finds naming them as a separate new genus: '' Loricatosaurus''. This made ''Lexovisaurus'' a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
'', while ''O. vetustus'' was found to be undiagnostic and declared a ''nomen dubium''. Other workers though, combining the English material collected by Leeds due to its shared provenance, have considered ''Lexovisaurus'' a valid
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
. In the meantime, ''Omosaurus vetustus'' has been renamed ''
Eoplophysis This list of informally named dinosaurs is a listing of dinosaurs (excluding Aves; birds and their extinct relatives) that have never been given formally published scientific names. This list only includes names that were not properly published ...
'', although this genus is not regarded as valid. The ''nomen nudum'' "Omosaurus leedsi" (mistakenly considered a ''nomen dubium'' by Maidment et al. 2008) has since been referred to ''Loricatosaurus''.


Description

''Lexovisaurus'' was a medium-sized stegosaur, reaching in length and in body mass. Little information about the holotype is available apart from it having a general stegosaurian build and a pelvis width of . Part of the material described by Hulke was a left femur, specimen BMNH R1991, with a length of . The Fletton and Normandy material show a combination of narrow flat plates on the back and round pointed spines that ran along the tail. A large spine was found that by Hoffstetter was placed on the shoulder, by Galton on the hip and by Maidment on the tail.


See also

*
Timeline of stegosaur research This timeline of stegosaur research is a chronological listing of events in the history of paleontology focused on the stegosaurs, the iconic plate-backed, spike-tailed herbivorous eurypod dinosaurs that predominated during the Jurassic period. ...


References


Further reading

* Benton, Michael. 1992. ''Dinosaur and other prehistoric animal Fact Finder'' {{Taxonbar, from=Q133863 Stegosaurs Middle Jurassic dinosaurs of Europe Jurassic England Fossils of England Oxford Clay Jurassic France Fossils of France Fossil taxa described in 1957 Taxa named by Robert Hoffstetter Controversial dinosaur taxa Ornithischian genera