Grasshopper
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Grasshoppers are a group of
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s belonging to the
suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
Caelifera The Caelifera are a suborder of orthopteran insects. They include the grasshoppers and grasshopper-like insects, as well as other superfamilies classified with them: the ground-hoppers ( Tetrigoidea) and pygmy mole crickets ( Tridactyloidea). ...
. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the
early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.9 Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which ...
around 250 million years ago. Grasshoppers are typically ground-dwelling insects with powerful hind legs which allow them to escape from threats by leaping vigorously. Their front legs are shorter and used for grasping food. As hemimetabolous insects, they do not undergo complete metamorphosis; they hatch from an egg into a
nymph A nymph (; ; sometimes spelled nymphe) is a minor female nature deity in ancient Greek folklore. Distinct from other Greek goddesses, nymphs are generally regarded as personifications of nature; they are typically tied to a specific place, land ...
or "hopper" which undergoes five
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at ...
s, becoming more similar to the adult insect at each developmental stage. The grasshopper hears through the tympanal organ which can be found in the first segment of the abdomen attached to the thorax; while its sense of vision is in the compound eyes, a change in light intensity is perceived in the simple eyes (ocelli). At high population densities and under certain environmental conditions, some grasshopper species can change colour and behavior and form swarms. Under these circumstances, they are known as
locust Locusts (derived from the Latin ''locusta'', locust or lobster) are various species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstances they b ...
s. Grasshoppers are plant-eaters, with a few species at times becoming serious pests of cereals, vegetables and pasture, especially when they swarm in the millions as locusts and destroy crops over wide areas. They protect themselves from
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s by
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
; when detected, many species attempt to startle the predator with a brilliantly coloured wing flash while jumping and (if adult) launching themselves into the air, usually flying for only a short distance. Other species such as the
rainbow grasshopper ''Dactylotum bicolor'', also known as the rainbow grasshopper, painted grasshopper, or the barber pole grasshopper, is a species of grasshopper in the family Acrididae. It is native to the United States, Canada and northern Mexico and exhibits ...
have
warning coloration Aposematism is the Advertising in biology, advertising by an animal, whether terrestrial or marine, to potential predation, predators that it is not worth attacking or eating. This unprofitability may consist of any defenses which make the pr ...
which deters predators. Grasshoppers are affected by parasites and various diseases, and many predatory creatures feed on both nymphs and adults. The eggs are subject to attack by
parasitoid In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable str ...
s and predators. Grasshoppers are diurnal insects, meaning they are most active during the day time. Grasshoppers have had a long relationship with humans. Swarms of locusts can have devastating effects and cause famine, having done so since Biblical times. Even in smaller numbers, the insects can be serious pests. They are used as food in countries such as Mexico and Indonesia. They feature in art, symbolism and literature. The study of grasshopper species is called
acridology Orthopterology is the scientific study of the order Orthoptera, which includes grasshoppers, crickets, locusts and some other insects. Someone that studies in this field is an orthopterist. The term is derived from the Ancient Greek words (''orth ...
.


Phylogeny

Grasshoppers belong to the suborder Caelifera. Although "grasshopper" has been used as a common name for the suborder in general, modern sources restrict it to the more "evolved"
families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...
. They may be placed in the infraorder
Acrididea Acrididea including the Acridomorpha is an infraorder of insects that describe the grasshoppers (thus also locusts) and ground-hoppers. It contains a large majority of species in the suborder Caelifera and the taxon Acridomorpha may also be us ...
and have been referred to as "short-horned grasshoppers" in older textsImms A.D., rev. Richards O.W. & Davies R.G. (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 886 pp. to distinguish them from the also-obsolete term "long-horned grasshoppers" (now bush-crickets or katydids) with their much longer antennae. The
phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...
of the Caelifera, based on mitochondrial
ribosomal RNA Ribosomal ribonucleic acid (rRNA) is a type of non-coding RNA which is the primary component of ribosomes, essential to all cells. rRNA is a ribozyme which carries out protein synthesis in ribosomes. Ribosomal RNA is transcribed from ribosomal ...
of thirty-two taxa in six out of seven superfamilies, is shown as a
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
. The
Ensifera Ensifera is a Order (biology), suborder of insects that includes the various types of crickets and their allies, including cricket (insect), true crickets, Rhaphidophoridae, camel crickets, Tettigoniidae, bush crickets or katydids, Prophalangops ...
(crickets, ''etc.''), Caelifera and all the superfamilies of grasshoppers except " Pamphagoidea" appear to be
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
. In evolutionary terms, the split between the Caelifera and the Ensifera is no more recent than the Permo-Triassic boundary; the earliest insects that are certainly Caeliferans are in the extinct families Locustopseidae and Locustavidae from the early Triassic, roughly 250 million years ago. The group diversified during the Triassic and have remained important plant-eaters from that time to now. The first modern families such as the Eumastacidae, Tetrigidae and Tridactylidae appeared in the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
, though some insects that might belong to the last two of these groups are found in the
early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic ...
. Morphological classification is difficult because many taxa have converged towards a common habitat type; recent taxonomists have concentrated on the internal genitalia, especially those of the male. This information is not available from fossil specimens, and the palaeontological taxonomy is founded principally on the venation of the hindwings. The Caelifera includes some 2,400 valid genera and about 11,000 known species. Many undescribed species probably exist, especially in tropical wet forests. The Caelifera have a predominantly tropical distribution with fewer species known from temperate zones, but most of the superfamilies have representatives worldwide. They are almost exclusively herbivorous and are probably the oldest living group of chewing herbivorous insects. The most diverse superfamily is the Acridoidea, with around 8,000 species. The two main families in this are the Acrididae (grasshoppers and locusts) with a worldwide distribution, and the Romaleidae (lubber grasshoppers), found chiefly in the New World. The Ommexechidae and Tristiridae are South American, and the Lentulidae, Lithidiidae and Pamphagidae are mainly African. The Pauliniids are nocturnal and can swim or skate on water, and the Lentulids are wingless. Pneumoridae are native to Africa, particularly southern Africa, and are distinguished by the inflated abdomens of the males.


Characteristics

Grasshoppers have the typical insect body plan of head,
thorax The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main di ...
, and
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
. The head is held vertically at an angle to the body, with the mouth at the bottom. The head bears a large pair of
compound eye A compound eye is a Eye, visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidium, ommatidia, which are tiny independent photoreception units that consist of a cornea, lens (anatomy), lens, and p ...
s which give all-round vision, three simple eyes which can detect light and dark, and a pair of thread-like antennae that are sensitive to touch and smell. The downward-directed mouthparts are modified for chewing and there are two sensory palps in front of the jaws.Pfadt, 1994. pp. 1–8 The thorax and abdomen are segmented and have a rigid
cuticle A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structu ...
made up of overlapping plates composed of
chitin Chitin (carbon, C8hydrogen, H13oxygen, O5nitrogen, N)n ( ) is a long-chain polymer of N-Acetylglucosamine, ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is the second most abundant polysaccharide in nature (behind only cell ...
. The three fused thoracic segments bear three pairs of legs and two pairs of wings. The forewings, known as tegmina, are narrow and leathery while the hindwings are large and membranous, the veins providing strength. The legs are terminated by claws for gripping. The hind leg is particularly powerful. The legs of these species are so powerful that they can jump quite a long distance. they also use this to flee from danger. The femur is robust and has several ridges where different surfaces join and the inner ridges bear stridulatory pegs in some species. The posterior edge of the tibia bears a double row of spines and there are a pair of articulated spurs near its lower end. The interior of the thorax houses the muscles that control the wings and legs. The abdomen has eleven segments, the first of which is fused to the thorax and contains the
tympanal organ A tympanal organ (or tympanic organ) is a hearing organ in insects, consisting of a tympanal membrane ( tympanum) stretched across a frame backed by an air sac and associated sensory neurons. Sounds vibrate the membrane, and the vibrations are s ...
and hearing system. Segments two to eight are ring-shaped and joined by flexible membranes. Segments nine to eleven are reduced in size; segment nine bears a pair of cerci and segments ten and eleven have the reproductive organs. Female grasshoppers are normally larger than males, with short ovipositors. The name of the suborder "Caelifera" comes from the Latin and means ''chisel-bearing'', referring to the shape of the ovipositor. The grasshopper's auditory organs are located on its abdomen, rather than on its head. These organs consist of a pair of membranes, each positioned on either side of the first abdominal segment and tucked under the wings. Known as tympanal organs, these simple eardrums vibrate in response to sound waves, enabling the grasshopper to hear the songs of other grasshoppers. Those species that make easily heard noises usually do so by rubbing a row of pegs on the hind legs against the edges of the forewings (stridulation). These sounds are produced mainly by males to attract females, though in some species the females also stridulate. Grasshoppers may be confused with crickets, but they differ in many aspects; these include the number of segments in their antennae and the structure of the ovipositor, as well as the location of the tympanal organ and the methods by which sound is produced. Ensiferans have antennae that can be much longer than the body and have at least 20–24 segments, while caeliferans have fewer segments in their shorter, stouter antennae.


Biology


Diet and digestion

Most grasshoppers are
polyphagous Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffixes -vore, -vory, or -vorous from Latin ''vorare'', meaning "to devour", or -phage, -phagy, or -phagous from Greek φαγε ...
, eating vegetation from multiple plant sources, but some are
omnivorous An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...
and also eat animal tissue and animal faeces. In general their preference is for grasses, including many
cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
s grown as crops. The digestive system is typical of insects, with Malpighian tubules discharging into the midgut. Carbohydrates are digested mainly in the crop, while proteins are digested in the ceca of the midgut. Saliva is abundant but largely free of enzymes, helping to move food and Malpighian secretions along the gut. Some grasshoppers possess
cellulase Cellulase (; systematic name 4-β-D-glucan 4-glucanohydrolase) is any of several enzymes produced chiefly by fungi, bacteria, and protozoans that catalyze cellulolysis, the decomposition of cellulose and of some related polysaccharides: : Endo ...
, which by softening plant cell walls makes plant cell contents accessible to other digestive enzymes. Grasshoppers can also be cannibalistic when swarming.


Sensory organs

Grasshoppers have a typical insect nervous system, and have an extensive set of external sense organs. On the side of the head are a pair of large compound eyes which give a broad field of vision and can detect movement, shape, colour and distance. There are also three simple eyes (
ocelli A simple eye or ocellus (sometimes called a pigment pit) is a form of eye or an optical arrangement which has a single lens without the sort of elaborate retina that occurs in most vertebrates. These eyes are called "simple" to distinguish the ...
) on the forehead which can detect light intensity, a pair of antennae containing olfactory (smell) and touch receptors, and mouthparts containing gustatory (taste) receptors. At the front end of the abdomen there is a pair of tympanal organs for sound reception. There are numerous fine hairs (
setae In biology, setae (; seta ; ) are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Depending partly on their form and function, protostome setae may be called macrotrichia, chaetae ...
) covering the whole body that act as mechanoreceptors (touch and wind sensors), and these are most dense on the antennae, the palps (part of the mouth), and on the cerci at the tip of the abdomen.Chapman, 2013. pp. 745–755 There are special receptors ( campaniform sensillae) embedded in the cuticle of the legs that sense pressure and cuticle distortion. There are internal "chordotonal" sense organs specialized to detect position and movement about the joints of the exoskeleton. The receptors convey information to the central nervous system through sensory neurons, and most of these have their cell bodies located in the periphery near the receptor site itself.


Circulation and respiration

Like other insects, grasshoppers have an
open circulatory system In vertebrates, the circulatory system is a system of organs that includes the heart, blood vessels, and blood which is circulated throughout the body. It includes the cardiovascular system, or vascular system, that consists of the heart a ...
and their body cavities are filled with
haemolymph Hemolymph, or haemolymph, is a fluid, similar to the blood in invertebrates, that circulates in the inside of the arthropod's body, remaining in direct contact with the animal's tissues. It is composed of a fluid plasma in which hemolymph ce ...
. A heart-like structure in the upper part of the abdomen pumps the fluid to the head from where it percolates past the tissues and organs on its way back to the abdomen. This system circulates nutrients throughout the body and carries
metabolic waste Metabolic wastes or excrements are substances left over from metabolic processes (such as cellular respiration) which cannot be used by the organism (they are surplus or toxic), and must therefore be excreted. This includes nitrogen compounds ...
s to be excreted into the gut. Other functions of the haemolymph include wound healing, heat transfer and the provision of hydrostatic pressure, but the circulatory system is not involved in gaseous exchange. Respiration is performed using
trachea The trachea (: tracheae or tracheas), also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all animals' lungs. The trachea extends from ...
e, air-filled tubes, which open at the surfaces of the thorax and abdomen through pairs of valved spiracles. Larger insects may need to actively ventilate their bodies by opening some spiracles while others remain closed, using abdominal muscles to expand and contract the body and pump air through the system.


Jumping

Grasshoppers jump by extending their large back legs and pushing against the substrate (the ground, a twig, a blade of grass or whatever else they are standing on); the reaction force propels them into the air. A large grasshopper, such as a locust, can jump about a metre (20 body lengths) without using its wings; the acceleration peaks at about 20 g. They jump for several reasons; to escape from a predator, to launch themselves into flight, or simply to move from place to place. For the escape jump in particular there is strong selective pressure to maximize take-off velocity, since this determines the range. This means that the legs must thrust against the ground with both high force and a high velocity of movement. A fundamental property of
muscle Muscle is a soft tissue, one of the four basic types of animal tissue. There are three types of muscle tissue in vertebrates: skeletal muscle, cardiac muscle, and smooth muscle. Muscle tissue gives skeletal muscles the ability to muscle contra ...
is that it cannot contract with high force and high velocity at the same time. Grasshoppers overcome this by using a
catapult A catapult is a ballistics, ballistic device used to launch a projectile at a great distance without the aid of gunpowder or other propellants – particularly various types of ancient and medieval siege engines. A catapult uses the sudden rel ...
mechanism to amplify the mechanical power produced by their muscles. The jump is a three-stage process. First, the grasshopper fully flexes the lower part of the leg (tibia) against the upper part (femur) by activating the flexor tibiae muscle (the back legs of the grasshopper in the top photograph are in this preparatory position). Second, there is a period of co-contraction in which force builds up in the large, pennate extensor tibiae muscle, but the tibia is kept flexed by the simultaneous contraction of the flexor tibiae muscle. The extensor muscle is much stronger than the flexor muscle, but the latter is aided by specialisations in the joint that give it a large effective mechanical advantage over the former when the tibia is fully flexed. Co-contraction can last for up to half a second, and during this period the extensor muscle shortens and stores elastic strain energy by distorting stiff cuticular structures in the leg. The extensor muscle contraction is quite slow (almost isometric), which allows it to develop high force (up to 14 N in the desert locust), but because it is slow only low power is needed. The third stage of the jump is the trigger relaxation of the flexor muscle, which releases the tibia from the flexed position. The subsequent rapid tibial extension is driven mainly by the relaxation of the elastic structures, rather than by further shortening of the extensor muscle. In this way the stiff cuticle acts like the elastic of a
catapult A catapult is a ballistics, ballistic device used to launch a projectile at a great distance without the aid of gunpowder or other propellants – particularly various types of ancient and medieval siege engines. A catapult uses the sudden rel ...
, or the bow of a bow-and-arrow. Energy is put into the store at low power by slow but strong muscle contraction, and retrieved from the store at high power by rapid relaxation of the mechanical elastic structures.


Stridulation

Male grasshoppers spend much of the day stridulating, singing more actively under optimal conditions and being more subdued when conditions are adverse; females also stridulate, but their efforts are insignificant when compared to the males. Late-stage male nymphs can sometimes be seen making stridulatory movements, although they lack the equipment to make sounds, demonstrating the importance of this behavioural trait. The songs are a means of communication; the male stridulation seems to express reproductive maturity, the desire for social cohesion and individual well-being. Social cohesion becomes necessary among grasshoppers because of their ability to jump or fly large distances, and the song can serve to limit dispersal and guide others to favourable habitat. The generalised song can vary in phraseology and intensity, and is modified in the presence of a rival male, and changes again to a courtship song when a female is nearby. In male grasshoppers of the family Pneumoridae, the enlarged abdomen amplifies stridulation.


Life cycle

In most grasshopper species, conflicts between males over females rarely escalate beyond ritualistic displays. Some exceptions include the chameleon grasshopper ('' Kosciuscola tristis''), where males may fight on top of ovipositing females; engaging in leg grappling, biting, kicking and mounting. Female grasshoppers of the species '' Chorthippus biguttulus'' appear to be able to integrate information from male calling songs.Clemens J, Krämer S, Ronacher B. Asymmetrical integration of sensory information during mating decisions in grasshoppers. Proc Natl Acad Sci U S A. 2014 Nov 18;111(46):16562-7. doi: 10.1073/pnas.1412741111. Epub 2014 Nov 3. PMID: 25368152; PMCID: PMC4246278. An unattractive song subunit far outweighs an attractive song subunit, and this asymmetrical integration is consistent with theories of
sexual selection Sexual selection is a mechanism of evolution in which members of one sex mate choice, choose mates of the other sex to mating, mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex ...
because it helps females avoid potentially costly interaction with unsuitable mating partners if the song belongs to another species or indicates a low-quality male. The newly emerged female grasshopper has a preoviposition period of a week or two while she increases in weight and her eggs mature. After mating, the female of most species digs a hole with her
ovipositor The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typica ...
and lays a batch of eggs in a pod in the ground near food plants, generally in the summer. After laying the eggs, she covers the hole with soil and litter. Some, like the semi-aquatic '' Cornops aquaticum'', deposit the pod directly into plant tissue. The eggs in the pod are glued together with a froth in some species. After a few weeks of development, the eggs of most species in temperate climates go into
diapause In animal dormancy, diapause is the delay in development in response to regular and recurring periods of adverse environmental conditions.Tauber, M.J., Tauber, C.A., Masaki, S. (1986) ''Seasonal Adaptations of Insects''. Oxford University Press It ...
, and pass the winter in this state. Diapause is broken by a sufficiently low ground temperature, with development resuming as soon as the ground warms above a certain threshold temperature. The embryos in a pod generally all hatch out within a few minutes of each other. They soon shed their membranes and their exoskeletons harden. These first
instar An instar (, from the Latin '' īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the exoskeleton in order to ...
nymphs can then jump away from predators. Pfadt, 1994. pp. 11–16
Diagrams
Grasshoppers undergo incomplete metamorphosis: they repeatedly moult, each instar becoming larger and more like an adult, with the wing-buds increasing in size at each stage. The number of instars varies between species but is often six. After the final moult, the wings are inflated and become fully functional. The migratory grasshopper, '' Melanoplus sanguinipes'', spends about 25 to 30 days as a nymph, depending on sex and temperature, and lives for about 51 days as an adult.


Swarming

Locusts are the swarming phase of certain species of short-horned grasshoppers in the family Acrididae. Swarming behaviour is a response to overcrowding. Increased tactile stimulation of the hind legs causes an increase in levels of
serotonin Serotonin (), also known as 5-hydroxytryptamine (5-HT), is a monoamine neurotransmitter with a wide range of functions in both the central nervous system (CNS) and also peripheral tissues. It is involved in mood, cognition, reward, learning, ...
. This causes the grasshopper to change colour, feed more and breed faster. The transformation of a solitary individual into a swarming one is induced by several contacts per minute over a short period. Following this transformation, under suitable conditions dense nomadic bands of flightless nymphs known as "hoppers" can occur, producing
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
s which attract the insects to each other. With several generations in a year, the locust population can build up from localised groups into vast accumulations of flying insects known as plagues, devouring all the vegetation they encounter. The largest recorded locust swarm was one formed by the now-extinct Rocky Mountain locust in 1875; the swarm was long and wide, and one estimate puts the number of locusts involved at 3.5 trillion.ffol An adult
desert locust The desert locust (''Schistocerca gregaria'') is a species of locust, a periodically swarming, short-horned grasshopper in the family Acrididae. They are found primarily in the deserts and dry areas of northern and eastern Africa, Arabia, and ...
can eat about of plant material each day, so the billions of insects in a large swarm can be very destructive, stripping all the foliage from plants in an affected area and consuming stems, flowers, fruits, seeds and bark.


Predators, parasites, and pathogens

Grasshoppers have a wide range of
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s at different stages of their lives; eggs are eaten by bee-flies,
ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it ...
s and blister beetles; hoppers and adults are taken by other insects such as ants, robber flies and sphecid wasps, by
spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
s, and by many birds and small mammals including dogs and cats.Capinera, 2008. pp. 1709–1710 The eggs and nymphs are under attack by
parasitoid In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable str ...
s including blow flies, flesh flies, and tachinid flies. External parasites of adults and nymphs include mites. Female grasshoppers parasitised by mites produce fewer eggs and thus have fewer offspring than unaffected individuals. The grasshopper nematode (''Mermis nigrescens'') is a long slender worm that infects grasshoppers, living in the insects'
hemocoel In vertebrates, the circulatory system is a organ system, system of organs that includes the heart, blood vessels, and blood which is circulated throughout the body. It includes the cardiovascular system, or vascular system, that consists of ...
. Adult worms lay eggs on plants and the host becomes infected when the foliage is eaten. '' Spinochordodes tellinii'' and '' Paragordius tricuspidatus'' are parasitic worms that infect grasshoppers and alter the behaviour of their hosts. When the worms are sufficiently developed, the grasshopper is persuaded to leap into a nearby body of water where it drowns, thus enabling the parasite to continue with the next stage of its life cycle, which takes place in water. Grasshoppers are affected by diseases caused by
bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
,
virus A virus is a submicroscopic infectious agent that replicates only inside the living Cell (biology), cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Viruses are ...
es,
fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
and
protozoa Protozoa (: protozoan or protozoon; alternative plural: protozoans) are a polyphyletic group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic debris. Historically ...
. The bacteria ''
Serratia marcescens ''Serratia marcescens'' () is a species of bacillus (shape), rod-shaped, Gram-negative bacteria in the family Yersiniaceae. It is a facultative anaerobe and an opportunistic pathogen in humans. It was discovered in 1819 by Bartolomeo Bizio in Pa ...
'' and ''
Pseudomonas aeruginosa ''Pseudomonas aeruginosa'' is a common Bacterial capsule, encapsulated, Gram-negative bacteria, Gram-negative, Aerobic organism, aerobic–facultative anaerobe, facultatively anaerobic, Bacillus (shape), rod-shaped bacteria, bacterium that can c ...
'' have both been implicated in causing disease in grasshoppers, as has the entomopathogenic fungus '' Beauveria bassiana''. This widespread fungus has been used to control various pest insects around the world, but although it infects grasshoppers, the infection is not usually lethal because basking in the sun has the result of raising the insects' temperature above a threshold tolerated by the fungus.Capinera, 2008. pp. 1229–1230 The fungal
pathogen In biology, a pathogen (, "suffering", "passion" and , "producer of"), in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a Germ theory of d ...
'' Entomophaga grylli'' is able to influence the behaviour of its grasshopper host, causing it to climb to the top of a plant and cling to the stem as it dies. This ensures wide dispersal of the fungal spores liberated from the corpse. The fungal pathogen '' Metarhizium acridum'' is found in Africa, Australia and Brazil where it has caused
epizootic In epizoology, an epizootic (or epizoötic, from Greek: ''epi-'' "upon" + ''zoon'' "animal") is a disease event in a nonhuman animal population analogous to an epidemic in humans. An epizootic disease (or ) may occur in a specific locale (an ...
s in grasshoppers. It is being investigated for possible use as a microbial insecticide for locust control. The
microsporidia Microsporidia are a group of spore-forming unicellular parasites. These spores contain an extrusion apparatus that has a coiled polar tube ending in an anchoring disc at the apical part of the spore.Franzen, C. (2005). How do Microsporidia inva ...
n fungus '' Nosema locustae'', once considered to be a protozoan, can be lethal to grasshoppers. It has to be consumed by mouth and is the basis for a bait-based commercial microbial pesticide. Various other microsporidians and protozoans are found in the gut.


Anti-predator defences

Grasshoppers exemplify a range of
anti-predator adaptation Anti-predator adaptations are mechanisms developed through evolution that assist Predation, prey organisms in their constant struggle against predators. Throughout the animal kingdom, adaptations have evolved for every stage of this struggle, na ...
s, enabling them to avoid detection, to escape if detected, and in some cases to avoid being eaten if captured. Grasshoppers are often
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
d to avoid detection by predators that hunt by sight; some species can change their coloration to suit their surroundings. Several species such as the hooded leaf grasshopper '' Phyllochoreia ramakrishnai'' (Eumastacoidea) are detailed mimics of leaves. Stick grasshoppers (Proscopiidae) mimic wooden sticks in form and coloration. Grasshoppers often have deimatic patterns on their wings, giving a sudden flash of bright colours that may startle predators long enough to give time to escape in a combination of jump and flight.Cott, p. 378 Some species are genuinely
aposematic Aposematism is the Advertising in biology, advertising by an animal, whether terrestrial or marine, to potential predation, predators that it is not worth attacking or eating. This unprofitability may consist of any defenses which make the pr ...
, having both bright warning coloration and sufficient toxicity to dissuade predators. '' Dictyophorus productus'' (Pyrgomorphidae) is a "heavy, bloated, sluggish insect" that makes no attempt to hide; it has a bright red abdomen. A '' Cercopithecus'' monkey that ate other grasshoppers refused to eat the species. Another species, the rainbow or painted grasshopper of Arizona, '' Dactylotum bicolor'' (Acridoidea), has been shown by experiment with a natural predator, the little striped whiptail lizard, to be aposematic. Atractomorpha lata, Burdwan, West Bengal, India 27 10 2012.jpg, Gaudy grasshopper, '' Atractomorpha lata'', evades predators with
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
. Titanacris Albipes Vol.jpg, Lubber grasshopper, '' Titanacris albipes'', has deimatically coloured wings, used to startle predators. LeafGrasshopper.jpg, Leaf grasshopper, '' Phyllochoreia ramakrishnai'', mimics a green leaf. Dactylotum bicolor.jpg, Painted grasshopper, '' Dactylotum bicolor'', deters predators with
warning coloration Aposematism is the Advertising in biology, advertising by an animal, whether terrestrial or marine, to potential predation, predators that it is not worth attacking or eating. This unprofitability may consist of any defenses which make the pr ...
. Aularches miliaris at Mangunan Orchard, Dlingo, Bantul, Yogyakarta 07.jpg, Spotted grasshopper, ''
Aularches miliaris ''Aularches miliaris'' is a grasshopper species of the monotypic genus ''Aularches'', belonging to the family Pyrgomorphidae.(2013''Aularches miliaris'' (Carl Linnaeus, Linnaeus, 10th edition of Systema Naturae, 1758) from Orthoptera Species File ...
'', defends itself with toxic foam and warning colours.


Relationship with humans


In art and media

Grasshoppers are occasionally depicted in artworks, such as the
Dutch Golden Age The Dutch Golden Age ( ) was a period in the history of the Netherlands which roughly lasted from 1588, when the Dutch Republic was established, to 1672, when the '' Rampjaar'' occurred. During this period, Dutch trade, scientific development ...
painter Balthasar van der Ast's
still life A still life (: still lifes) is a work of art depicting mostly wikt:inanimate, inanimate subject matter, typically commonplace objects which are either natural (food, flowers, dead animals, plants, rocks, shells, etc.) or artificiality, human-m ...
oil painting, ''Flowers in a Vase with Shells and Insects'', c. 1630, now in the
National Gallery, London The National Gallery is an art museum in Trafalgar Square in the City of Westminster, in Central London, England. Founded in 1824, it houses a collection of more than 2,300 paintings dating from the mid-13th century to 1900. The current dire ...
, though the insect may be a bush-cricket. Another orthopteran is found in
Rachel Ruysch Rachel Ruysch (3 June 1664 – 12 October 1750) was a Dutch still-life painter from the Northern Netherlands. She specialized in flowers, inventing her own style and achieving international fame in her lifetime. Due to a long and successful car ...
's still life ''Flowers in a Vase'', c. 1685. The seemingly static scene is animated by a "grasshopper on the table that looks about ready to spring", according to the gallery curator Betsy Wieseman, with other invertebrates including a spider, an ant, and two caterpillars. Grasshoppers are also featured in cinema. The 1957 film '' Beginning of the End'' portrayed giant grasshoppers attacking
Chicago Chicago is the List of municipalities in Illinois, most populous city in the U.S. state of Illinois and in the Midwestern United States. With a population of 2,746,388, as of the 2020 United States census, 2020 census, it is the List of Unite ...
. In the 1998
Disney The Walt Disney Company, commonly referred to as simply Disney, is an American multinational mass media and entertainment industry, entertainment conglomerate (company), conglomerate headquartered at the Walt Disney Studios (Burbank), Walt Di ...
/
Pixar Pixar (), doing business as Pixar Animation Studios, is an American animation studio based in Emeryville, California, known for its critically and commercially successful computer-animated feature films. Pixar is a subsidiary of Walt Disney ...
animated film ''
A Bug's Life ''A Bug's Life'' (stylized in all lowercase) is a 1998 American animated comedy film directed by John Lasseter from a screenplay written by Andrew Stanton, Donald McEnery, and Bob Shaw, and a story conceived by Lasseter, Stanton, and Joe Ran ...
'', the antagonists are a gang of grasshoppers, with their leader Hopper serving as the main villain. The protagonists of the 1971 ''
tokusatsu is a Japanese term for live-action films or television programs that make heavy use of practical special effects. Credited to special effects director Eiji Tsuburaya, ''tokusatsu'' mainly refers to science fiction film, science fiction, War fi ...
'' series ''
Kamen Rider The , also known as ''Masked Rider Series'' (until ''Kamen Rider Decade, Decade'' and except Thailand), is a Japanese superhero fiction, superhero media franchise consisting of tokusatsu television programs, films, manga, and anime, created by ...
'' primarily carry a grasshopper motif (for example Kamen Rider Black's Batta Man form), which continues to serve as the baseline visual template for most entries in the
media franchise A media franchise, also known as a multimedia franchise, is a collection of related media in which several derivative works have been produced from an original creative work of fiction, such as a film, a work of literature, a television program, o ...
it has given birth to since.


Symbolism

Grasshoppers are sometimes used as symbols. During the Greek Archaic Era, the grasshopper was the symbol of the ''
polis Polis (: poleis) means 'city' in Ancient Greek. The ancient word ''polis'' had socio-political connotations not possessed by modern usage. For example, Modern Greek πόλη (polē) is located within a (''khôra''), "country", which is a πατ ...
'' of
Athens Athens ( ) is the Capital city, capital and List of cities and towns in Greece, largest city of Greece. A significant coastal urban area in the Mediterranean, Athens is also the capital of the Attica (region), Attica region and is the southe ...
, possibly because they were among the most common insects on the dry plains of
Attica Attica (, ''Attikḗ'' (Ancient Greek) or , or ), or the Attic Peninsula, is a historical region that encompasses the entire Athens metropolitan area, which consists of the city of Athens, the capital city, capital of Greece and the core cit ...
. Native Athenians for a while wore golden grasshopper brooches to symbolise that they were of pure Athenian lineage with no foreign ancestors. In addition,
Peisistratus Pisistratus (also spelled Peisistratus or Peisistratos; ;  – 527 BC) was a politician in ancient Athens, ruling as tyrant in the late 560s, the early 550s and from 546 BC until his death. His unification of Attica, the triangular ...
hung the figure of a kind of grasshopper before the
Acropolis of Athens The Acropolis of Athens (; ) is an ancient citadel located on a rocky outcrop above the city of Athens, Greece, and contains the remains of several Ancient Greek architecture, ancient buildings of great architectural and historical significance, ...
as
apotropaic magic Apotropaic magic (From ) or protective magic is a type of magic intended to turn away harm or evil influences, as in deflecting misfortune or averting the evil eye. Apotropaic observances may also be practiced out of superstition or out of tr ...
.A Dictionary of Greek and Roman Antiquities (1890), Fascinum
/ref> Another symbolic use of the grasshopper is Sir Thomas Gresham's gilded grasshopper in
Lombard Street, London Lombard Street () is a street notable for its connections with the City of London's merchant, banking and insurance industries, stretching back to medieval times. From Bank junction, where nine streets converge by the Bank of England, Lombard ...
, dating from 1563; the building was for a while the headquarters of the Guardian Royal Exchange, but the company declined to use the symbol for fear of confusion with the locust. Grasshoppers appearing in dreams have been interpreted as symbols of "Freedom, independence, spiritual enlightenment, inability to settle down or commit to decision". Locusts are taken literally to mean devastation of crops in the case of farmers; figuratively as "wicked men and women" for non-farmers; and "Extravagance, misfortune, & ephemeral happiness" by "gypsies".


As food

In some countries, grasshoppers are used as food. In southern
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, grasshoppers, known as '' chapulines'', are eaten in a variety of dishes, such as in
tortilla A tortilla (, ) is a thin, circular unleavened flatbread from Mesoamerica originally made from maize hominy meal, and now also from wheat flour. The Aztecs and other Nahuatl speakers called tortillas ''tlaxcalli'' (). First made by the indi ...
s with chilli sauce. Grasshoppers are served on skewers in some Chinese food markets, like the Donghuamen Night Market. Fried grasshoppers (''walang goreng'') are eaten in the
Gunung Kidul Regency Gunůngkidůl () is a Regency (Indonesia), regency area located in the Special Province of the Daerah Istimewa Yogyakarta, Indonesia. The administrative centre is the town of Wonosari. The name "Gunungkidul" comes from the Javanese language ''moun ...
,
Yogyakarta Yogyakarta is the capital city of the Special Region of Yogyakarta in Indonesia, in the south-central part of the island of Java. As the only Indonesian royal city still ruled by Hamengkubuwono, a monarchy, Yogyakarta is regarded as an importan ...
,
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
in Indonesia. Grasshoppers are a beloved delicacy in Uganda; they are usually eaten fried (most commonly in November and May after the rains). In America, the
Ohlone The Ohlone ( ), formerly known as Costanoans (from Spanish meaning 'coast dweller'), are a Native Americans in the United States, Native American people of the Northern California coast. When Spanish explorers and missionaries arrived in the l ...
burned grassland to herd grasshoppers into pits where they could be collected as food. It is recorded in the Bible that
John the Baptist John the Baptist ( – ) was a Jewish preacher active in the area of the Jordan River in the early first century AD. He is also known as Saint John the Forerunner in Eastern Orthodoxy and Oriental Orthodoxy, John the Immerser in some Baptist ...
ate locusts and wild honey (Greek: ἀκρίδες καὶ μέλι ἄγριον, ''akrídes kaì méli ágrion'') while living in the wilderness. However, because of a tradition of depicting him as an
ascetic Asceticism is a lifestyle characterized by abstinence from worldly pleasures through self-discipline, self-imposed poverty, and simple living, often for the purpose of pursuing spiritual goals. Ascetics may withdraw from the world for their pra ...
, attempts have been made to explain that the ''locusts'' were in fact a suitably ascetic
vegetarian Vegetarianism is the practice of abstaining from the Eating, consumption of meat (red meat, poultry, seafood, insects as food, insects, and the flesh of any other animal). It may also include abstaining from eating all by-products of animal slau ...
food such as
carob The carob ( ; ''Ceratonia siliqua'') is a flowering evergreen tree or shrub in the Caesalpinioideae sub-family of the legume family, Fabaceae. It is widely cultivated for its edible fruit, which takes the form of seed pods, and as an ornam ...
beans, notwithstanding the fact that the word ''ἀκρίδες'' means plainly ''grasshoppers''. In recent years, with the search for alternative healthy and sustainable protein sources, grasshoppers are being cultivated by commercial companies operating grasshopper farms and are being used as food and protein supplements.


As pests

Grasshoppers eat large quantities of foliage both as adults and during their development, and can be serious pests of arid land and prairies. Pasture, grain, forage, vegetable and other crops can be affected. Grasshoppers often bask in the sun, and thrive in warm sunny conditions, so drought stimulates an increase in grasshopper populations. A single season of drought is not normally sufficient to stimulate a major population increase, but several successive dry seasons can do so, especially if the intervening winters are mild so that large numbers of nymphs survive. Although sunny weather stimulates growth, there needs to be an adequate food supply for the increasing grasshopper population. This means that although precipitation is needed to stimulate plant growth, prolonged periods of cloudy weather will slow nymphal development.Capinera, 2008. pp. 1710–1712 Grasshoppers can best be prevented from becoming pests by manipulating their environment. Shade provided by trees will discourage them and they may be prevented from moving onto developing crops by removing coarse vegetation from fallow land and field margins and discouraging thick growth beside ditches and on roadside verges. With increasing numbers of grasshoppers, predator numbers may increase, but this seldom happens rapidly enough to have much effect on populations. Biological control is being investigated, and spores of the protozoan parasite '' Nosema locustae'' can be used mixed with bait to control grasshoppers, being more effective with immature insects. On a small scale, neem products can be effective as a feeding deterrent and as a disruptor of nymphal development.
Insecticide Insecticides are pesticides used to kill insects. They include ovicides and larvicides used against insect eggs and larvae, respectively. The major use of insecticides is in agriculture, but they are also used in home and garden settings, i ...
s can be used, but adult grasshoppers are difficult to kill, and as they move into fields from surrounding rank growth, crops may soon become reinfested. Some grasshopper species, like the Chinese rice grasshopper, are a pest in
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
paddies. Ploughing exposes the eggs on the surface of the field, to be destroyed by sunshine or eaten by natural enemies. Some eggs may be buried too deeply in the soil for hatching to take place. Locust plagues can have devastating effects on human populations, causing
famine A famine is a widespread scarcity of food caused by several possible factors, including, but not limited to war, natural disasters, crop failure, widespread poverty, an Financial crisis, economic catastrophe or government policies. This phenom ...
s and population upheavals. They are mentioned in both the
Qur’an The Quran, also Romanization, romanized Qur'an or Koran, is the central religious text of Islam, believed by Muslims to be a Waḥy, revelation directly from God in Islam, God (''Allah, Allāh''). It is organized in 114 chapters (, ) which ...
and the
Bible The Bible is a collection of religious texts that are central to Christianity and Judaism, and esteemed in other Abrahamic religions such as Islam. The Bible is an anthology (a compilation of texts of a variety of forms) originally writt ...
and have also been held responsible for
cholera Cholera () is an infection of the small intestine by some Strain (biology), strains of the Bacteria, bacterium ''Vibrio cholerae''. Symptoms may range from none, to mild, to severe. The classic symptom is large amounts of watery diarrhea last ...
epidemics, resulting from the corpses of locusts drowned in the Mediterranean Sea and decomposing on beaches.Capinera, 2008. pp 1181–1183 The
FAO The Food and Agriculture Organization of the United Nations; . (FAO) is a List of specialized agencies of the United Nations, specialized agency of the United Nations that leads international efforts to defeat hunger and improve nutrition ...
and other organisations monitor locust activity around the world. Timely application of pesticides can prevent nomadic bands of hoppers from forming before dense swarms of adults can build up. Besides conventional control using contact insecticides,
biological pest control Biological control or biocontrol is a method of controlling pests, whether pest animals such as insects and mites, weeds, or pathogens affecting animals or plants by using other organisms. It relies on predation, parasitism, herbivory, or ot ...
using the entomopathogenic fungus '' Metarhizium acridum'', which specifically infects grasshoppers, has been used with some success.


Detection of explosives

In February 2020, researchers from Washington University in St. Louis announced they had engineered "cyborg grasshoppers" capable of accurately detecting explosives. In the project, funded by the
US Office of Naval Research The Office of Naval Research (ONR) is an organization within the United States Department of the Navy responsible for the science and technology programs of the U.S. Navy and United States Marine Corps, Marine Corps. Established by United States ...
, researchers fitted grasshoppers with lightweight sensor backpacks that recorded and transmitted the electrical activity of their antennal lobes to a computer. According to the researchers, the grasshoppers were able to detect the location of the highest concentration of explosives. The researchers also tested the effect of combining sensorial information from several grasshoppers on detection accuracy. The neural activity from seven grasshoppers yielded an average detection accuracy rate of 80%, whereas a single grasshopper yielded a 60% rate.


In literature

The Egyptian word for locust or grasshopper was written ''snḥm'' in the consonantal hieroglyphic writing system. The pharaoh
Ramesses II Ramesses II (sometimes written Ramses or Rameses) (; , , ; ), commonly known as Ramesses the Great, was an Pharaoh, Egyptian pharaoh. He was the third ruler of the Nineteenth Dynasty of Egypt, Nineteenth Dynasty. Along with Thutmose III of th ...
compared the armies of the
Hittites The Hittites () were an Anatolian peoples, Anatolian Proto-Indo-Europeans, Indo-European people who formed one of the first major civilizations of the Bronze Age in West Asia. Possibly originating from beyond the Black Sea, they settled in mo ...
to locusts: "They covered the mountains and valleys and were like locusts in their multitude." One of
Aesop's Fables Aesop's Fables, or the Aesopica, is a collection of fables credited to Aesop, a Slavery in ancient Greece, slave and storyteller who lived in ancient Greece between 620 and 564 Before the Common Era, BCE. Of varied and unclear origins, the stor ...
, later retold by La Fontaine, is the tale of ''
The Ant and the Grasshopper The Ant and the Grasshopper, alternatively titled The Grasshopper and the Ant (or Ants), is one of Aesop's Fables, numbered 373 in the Perry Index. The fable describes how a hungry grasshopper begs for food from an ant when winter comes and is ...
''. The ant works hard all summer, while the grasshopper plays. In winter, the ant is ready but the grasshopper starves. Somerset Maugham's short story "The Ant and the Grasshopper" explores the fable's symbolism via complex framing. Other human weaknesses besides improvidence have become identified with the grasshopper's behaviour. So an unfaithful woman (hopping from man to man) is "a grasshopper" in "Poprygunya", an 1892 short story by
Anton Chekhov Anton Pavlovich Chekhov (; ; 29 January 1860 – 15 July 1904) was a Russian playwright and short-story writer, widely considered to be one of the greatest writers of all time. His career as a playwright produced four classics, and his b ...
, and in
Jerry Paris William Gerald Paris (July 25, 1925 – March 31, 1986) was an American actor and director best known for playing Jerry Helper, the dentist and next-door neighbor of Rob and Laura Petrie, on ''The Dick Van Dyke Show'', and for directing the majo ...
's 1969 film '' The Grasshopper''.


In mechanical engineering

The name "Grasshopper" was given to the Aeronca L-3 and Piper L-4 light aircraft, both used for
reconnaissance In military operations, military reconnaissance () or scouting is the exploration of an area by military forces to obtain information about enemy forces, the terrain, and civil activities in the area of operations. In military jargon, reconnai ...
and other support duties in
World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
. The name is said to have originated when Major General Innis P. Swift saw a Piper making a rough landing and remarked that it looked like a grasshopper for its bouncing progress. Grasshopper beam engines were
beam engine A beam engine is a type of steam engine where a pivoted overhead Beam (structure), beam is used to apply the force from a vertical piston to a vertical connecting rod. This configuration, with the engine directly driving a pump, was first used b ...
s pivoted at one end, the long horizontal arm resembling the hind leg of a grasshopper. The type was patented by William Freemantle in 1803.


Notes


References


Sources

* * * *


External links

* * * * {{Authority control Acrididea Extant Triassic first appearances Herbivorous insects Insects in culture Insect common names