Ficus indica
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''Ficus'' ( or ) is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of about 850
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of woody
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s,
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from tree ...
s,
vine A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themsel ...
s,
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
s and hemiepiphytes in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Moraceae The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 genera and over 1100 species. Most are widespread in tropical and subtropical regions, less so in temperate climates; however ...
. Collectively known as fig trees or figs, they are native throughout the
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also refer ...
with a few species extending into the semi-warm
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
zone. The
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the worl ...
(''F. carica'') is a temperate species native to southwest Asia and the
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
region (from Afghanistan to Portugal), which has been widely cultivated from ancient times for its
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
, also referred to as figs. The fruit of most other species are also edible though they are usually of only local economic importance or eaten as bushfood. However, they are extremely important food resources for wildlife. Figs are also of considerable cultural importance throughout the tropics, both as objects of worship and for their many practical uses.


Description

''Ficus'' is a
pantropical A pantropical ("all tropics") distribution is one which covers tropical regions of both hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and '' Bacopa''. ''Neotropical'' is a zoogeographic te ...
genus of trees, shrubs, and vines occupying a wide variety of
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...
s; most are
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
, but some deciduous species are found in areas outside of the tropics and to higher elevations. Fig species are characterized by their unique
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
and distinctive pollination syndrome, which uses wasp species belonging to the family Agaonidae for pollination. Specific identification of many of the species can be difficult, but members of the genus ''Ficus'' are relatively easy to recognize. Many have aerial roots and a distinctive shape or habit, and their fruits distinguish them from other plants. The fruit of ''Ficus'' is an
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
enclosed in an urn-like structure called a
syconium Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flow ...
, which is lined on the inside with the fig's tiny flowers that develop into multiple ovaries on the inside surface. In essence, the fig fruit is a fleshy stem with multiple tiny flowers that fruit and coalesce. The unique fig
pollination Pollination is the transfer of pollen from an Stamen, anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by Anemophily, wind. Pollinating agents can ...
system, involving tiny, highly specific wasps, known as fig wasps that enter via
ostiole An ''ostiole'' is a small hole or opening through which algae or fungi release their mature spores. The word is a diminutive of "ostium", "opening". The term is also used in higher plants, for example to denote the opening of the involuted ...
these subclosed inflorescences to both pollinate and lay their own eggs, has been a constant source of inspiration and wonder to biologists. Notably, three vegetative traits together are unique to figs. All figs present a white to yellowish
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms ...
, some in copious quantities; the twig shows paired
stipules In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...
—or circular scars if the stipules have fallen off; the lateral veins at the base of the leaf are steep, forming a tighter angle with the midrib than the other lateral veins, a feature referred to as "triveined". No unambiguous older
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s of ''Ficus'' are known. However, current
molecular clock The molecular clock is a figurative term for a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged. The biomolecular data used for such calculations are usually nucleo ...
estimates indicate that ''Ficus'' is a relatively ancient genus being at least 60 million years old, and possibly as old as 80 million years. The main radiation of
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
species, however, may have taken place more recently, between 20 and 40 million years ago. Some better-known species that represent the diversity of the genus include the
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the worl ...
, a small, temperate deciduous tree whose fingered fig leaf is well known in art and
iconography Iconography, as a branch of art history, studies the identification, description and interpretation of the content of images: the subjects depicted, the particular compositions and details used to do so, and other elements that are distinct fro ...
; the weeping fig (''F. benjamina''), a hemiepiphyte with thin, tough leaves on pendulous stalks adapted to its rain forest habitat; the rough-leaved
sandpaper fig The sandpaper figs are so named for their leaves, which are rough and sandpaper-like in texture. The common name may refer to a number of species in the genus ''Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vine ...
s from Australia; and the creeping fig (''F. pumila''), a vine whose small, hard leaves form a dense carpet of foliage over rocks or garden walls. Moreover, figs with different plant habits have undergone
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic in ...
in different biogeographic regions, leading to very high levels of
alpha diversity In ecology, alpha diversity (α-diversity) is the mean species diversity in a site at a local scale. The term was introduced by R. H. WhittakerWhittaker, R. H. (1960) Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monograp ...
. In the tropics, ''Ficus'' commonly is the most species-rich plant genus in a particular forest. In Asia, as many as 70 or more species can co-exist. ''Ficus'' species richness declines with an increase in latitude in both hemispheres. A description of fig tree cultivation is set out in Ibn al-'Awwam's 12th-century agricultural work entitled, ''Book on Agriculture''.


Ecology

Figs are
keystone species A keystone species is a species which has a disproportionately large effect on its natural environment relative to its abundance, a concept introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in maintaini ...
in many
tropical forest Tropical forests (a.k.a. jungle) are forested landscapes in tropical regions: ''i.e.'' land areas approximately bounded by the tropic of Cancer and Capricorn, but possibly affected by other factors such as prevailing winds. Some tropical fore ...
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s. Their fruit are a key resource for some frugivores including fruit bats, and primates including: capuchin monkeys, langurs,
gibbons Gibbons may refer to: * The plural of gibbon, an ape in the family Hylobatidae * Gibbons (surname) * Gibbons, Alberta * Gibbons (automobile), a British light car of the 1920s * Gibbons P.C., a leading American law firm headquartered in New Jerse ...
and mangabeys. They are even more important for
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s such as Asian barbets, pigeons,
hornbill Hornbills (Bucerotidae) are a family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a casque on the upper mandibl ...
s, fig-parrots and
bulbul The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical A ...
s, which may almost entirely subsist on figs when these are in plenty. Many
Lepidoptera Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described speci ...
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sy ...
s feed on fig leaves, for example several '' Euploea'' species (crow butterflies), the plain tiger (''Danaus chrysippus''), the giant swallowtail (''Papilio cresphontes''), the brown awl (''Badamia exclamationis''), and '' Chrysodeixis eriosoma'',
Choreutidae Choreutidae, or metalmark moths, are a family of insects in the lepidopteran order whose relationships have been long disputed. It was placed previously in the superfamily Yponomeutoidea in family Glyphipterigidae and in superfamily Sesioidea ...
and
Copromorphidae Copromorphidae, the "tropical fruitworm moths", is a family of insects in the lepidopteran order. These moths have broad, rounded forewings, and well-camouflaged scale patterns. Unlike Carposinidae the mouthparts include "labial palps" with th ...
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s. The citrus long-horned beetle (''Anoplophora chinensis''), for example, has larvae that feed on
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
, including that of fig trees; it can become a pest in fig plantations. Similarly, the sweet potato whitefly (''Bemisia tabaci'') is frequently found as a pest on figs grown as potted plants and is spread through the export of these plants to other localities. For a list of other diseases common to fig trees, see List of foliage plant diseases (Moraceae).


Fig fruit and reproduction system

Many fig species are grown for their fruits, though only ''
Ficus carica The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
'' is cultivated to any extent for this purpose. A fig "fruit" is a type of
multiple fruit Multi-fruits, also called collective fruits, are fruiting bodies formed from a cluster of flowers, the '' inflorescence''. Each flower in the inflorescence produces a fruit, but these mature into a single mass. After flowering the mass is calle ...
known as a
syconium Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flow ...
, derived from an arrangement of many small flowers on an inverted, nearly closed receptacle. The many small flowers are unseen unless the fig is cut open. The fruit typically has a bulbous shape with a small opening (the
ostiole An ''ostiole'' is a small hole or opening through which algae or fungi release their mature spores. The word is a diminutive of "ostium", "opening". The term is also used in higher plants, for example to denote the opening of the involuted ...
) at the outward end that allows access to
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the m ...
s. The flowers are pollinated by very small wasps that crawl through the opening in search of a suitable place to lay eggs. Without this pollinator service fig trees could not reproduce by seed. In turn, the flowers provide a safe haven and nourishment for the next generation of wasps. This accounts for the frequent presence of wasp larvae in the fruit, and has led to a
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
ary relationship. Technically, a fig fruit proper would be only one of the many tiny matured, seed-bearing gynoecia found inside one fig – if you cut open a fresh fig, individual fruit will appear as fleshy "threads", each bearing a single seed inside. The genus '' Dorstenia'', also in the fig family (Moraceae), exhibits similar tiny flowers arranged on a receptacle but in this case the receptacle is a more or less flat, open surface. Fig plants can be
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is ...
(
hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have ...
) or gynodioecious (hermaphrodite and female). Nearly half of fig species are gynodioecious, and therefore have some plants with inflorescences (syconium) with long styled pistillate flowers, and other plants with staminate flowers mixed with short styled pistillate flowers. The long-styled flowers tend to prevent wasps from laying their eggs within the ovules, while the short-styled flowers are accessible for egg laying. All the native fig trees of the American continent are hermaphrodites, as well as species like Indian banyan (''F. benghalensis''), weeping fig (''F. benjamina''), Indian rubber plant (''F. elastica''), fiddle-leaved fig (''F. lyrata''),
Moreton Bay fig ''Ficus macrophylla'', commonly known as the Moreton Bay fig or Australian banyan, is a large evergreen banyan tree of the family Moraceae native to eastern Australia, from the Wide Bay–Burnett region in the north to the Illawarra in New S ...
(''F. macrophylla''), Chinese banyan (''F. microcarpa''),
sacred fig ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
(''F. religiosa'') and
sycamore fig ''Ficus sycomorus'', called the sycamore fig or the fig-mulberry (because the leaves resemble those of the mulberry), sycamore, or sycomore, is a fig species that has been cultivated since ancient times. The term '' sycamore'' spelled with an A ...
(''F. sycomorus''). The common fig (''Ficus carica'') is a gynodioecious plant, as well as lofty fig or clown fig ('' F. aspera''), Roxburgh fig ('' F. auriculata''), mistletoe fig ('' F. deltoidea''), '' F. pseudopalma'', creeping fig (''F. pumila'') and related species. The hermaphrodite common figs are called "inedible figs" or "caprifigs"; in traditional culture in the Mediterranean region they were considered food for
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s (''Capra aegagrus''). In the female fig trees, the male flower parts fail to develop; they produce the "'edible figs". Fig wasps grow in common fig caprifigs but not in the female syconiums because the female flower is too long for the wasp to successfully lay her eggs in them. Nonetheless, the wasp pollinates the flower with pollen from the caprifig it grew up in. When the wasp dies, it is broken down by enzymes ( Ficain) inside the fig. Fig wasps are not known to transmit any diseases harmful to humans. When a caprifig ripens, another caprifig must be ready to be pollinated. In temperate climes, wasps hibernate in figs, and there are distinct crops. Caprifigs have three crops per year; common figs have two. The first crop (
breba A breba (or more commonly ''breva'' in Spanish, and sometimes as ''taqsh'') is a fig that develops on a common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Medi ...
) is larger and juicier, and usually eaten fresh. In cold climates the breba crop is often destroyed by spring frosts. Some parthenocarpic
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s of common figs do not require pollination at all, and will produce a crop of figs (albeit
sterile Sterile or sterility may refer to: *Asepsis Asepsis is the state of being free from disease-causing micro-organisms (such as pathogenic bacteria, viruses, pathogenic fungi, and parasites). There are two categories of asepsis: medical and surgi ...
) in the absence of caprifigs and fig wasps. Depending on the species, each fruit can contain hundreds or even thousand of seeds. Figs can be propagated by seeds, cuttings, air-layering or grafting. However, as with any plant, figs grown from seed are not necessarily genetically identical to the parent and are only propagated this way for breeding purposes.


Mutualism with the pollinating fig wasps

Each species of fig is pollinated by one or a few specialised wasp species, and therefore plantings of fig species outside of their native range results in effectively sterile individuals. For example, in
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
, some 60 species of figs have been introduced, but only four of the wasps that fertilize them, so only those species of figs produce viable seeds there and can become
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...
. This is an example of mutualism, in which each organism (fig plant and fig wasp) benefit each other, in this case reproductively. The intimate association between fig species and their wasp pollinators, along with the high incidence of a one-to-one plant-pollinator ratio have long led scientists to believe that figs and wasps are a clear example of
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
. Morphological and reproductive behavior evidence, such as the correspondence between fig and wasp larvae maturation rates, have been cited as support for this hypothesis for many years. Additionally, recent genetic and molecular dating analyses have shown a very close correspondence in the character evolution and
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
phylogenies of these two clades. According to meta-analysis of molecular data for 119 fig species 35% (41) have multiple pollinator wasp species. The real proportion is higher because not all wasp species were detected. On the other hand, species of wasps pollinate multiple host fig species. Molecular techniques, like
microsatellite A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. ...
markers and mitochondrial sequence analysis, allowed a discovery of multiple genetically distinct,
cryptic Cryptic may refer to: In science: * Cryptic species complex, a group of species that are very difficult to distinguish from one another * Crypsis, the ability of animals to blend in to avoid observation * Cryptic era, earliest period of the Earth ...
wasp species. Not all these cryptic species are sister taxa and thus must have experienced a host fig shift at some point. These cryptic species lacked evidence of genetic
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intr ...
or backcrosses indicating limited fitness for hybrids and effective
reproductive isolation The mechanisms of reproductive isolation are a collection of evolutionary mechanisms, behaviors and physiological processes critical for speciation. They prevent members of different species from producing offspring, or ensure that any offsprin ...
and
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
. The existence of cryptic species suggests that neither the number of symbionts nor their evolutionary relationships are necessarily fixed ecologically. While the morphological characteristics that facilitate the fig-wasp mutualisms are likely to be shared more fully in closer relatives, the absence of unique pairings would make it impossible to do a one-to-one tree comparison and difficult to determine cospeciation.


Systematics

With 800 species, ''Ficus'' is by far the largest genus in the Moraceae, and is one of the largest genera of flowering plants currently described. The species currently classified within ''Ficus'' were originally split into several genera in the mid-1800s, providing the basis for a subgeneric classification when reunited into one genus in 1867. This classification put functionally
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
species into four subgenera based on floral characters. In 1965, E. J. H. Corner reorganized the genus on the basis of breeding system, uniting these four dioecious subgenera into a single dioecious subgenus ''Ficus''.
Monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is ...
figs were classified within the subgenera ''
Urostigma A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as ...
'', '' Pharmacosycea'' and '' Sycomorus''. This traditional classification has been called into question by recent
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...
studies employing genetic methods to investigate the relationships between representative members of the various sections of each subgenus. Of Corner's original subgeneric divisions of the genus, only ''Sycomorus'' is supported as monophyletic in the majority of phylogenetic studies. Notably, there is no clear split between dioecious and monoecious lineages. One of the two sections of ''Pharmacosycea'', a monoecious group, form a monophyletic clade basal to the rest of the genus, which includes the other section of ''Pharmacosycea'', the rest of the monoecious species, and all of the dioecious species. These remaining species are divided into two main monophyletic lineages (though the statistical support for these lineages isn't as strong as for the monophyly of the more derived clades within them). One consists of all sections of ''Urostigma'' except for section ''Urostigma s. s.''. The other includes section ''Urostigma s. s.'', subgenus ''Sycomorus'', and the species of subgenus ''Ficus'', though the relationships of the sections of these groups to one another are not well resolved.


Selected species

There are 875 accepted ''Ficus'' species, as of March 2021, according to
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
.


Subgenus ''Ficus''

*'' Ficus amplissima'' Sm. – bat fig *''
Ficus carica The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
'' L. – common fig *'' Ficus daimingshanensis'' Chang *''
Ficus deltoidea ''Ficus deltoidea'', commonly known as mistletoe fig (''mas cotek'' in Malay language, Malaysian, ''tabat barito'' in Indonesia, or ''สาลิกาลิ้นทอง'' in Thai language, Thai) is a species of flowering plant in the family ...
'' Jack – mistletoe fig *''
Ficus erecta ''Ficus erecta'' (syn. ''Ficus beecheyana''), the Japanese fig, is a species of flowering plant in the family Moraceae. It is found in the eastern Himalayas, Assam, Bangladesh, Vietnam, southern China, Taiwan, Jeju Island of South Korea, the Ry ...
'' Thunb. – Japanese fig *'' Ficus fulva'' Reinw. ex Blume *'' Ficus grossularioides'' Burman f. – white-leaved fig *''
Ficus neriifolia ''Ficus neriifolia'' is a species of fig (''Ficus''). It is native to Asia, including Bhutan, Burma, China, India, and Nepal.
'' Sm. *'' Ficus palmata'' Forssk. *'' Ficus pandurata'' Hance *''
Ficus simplicissima ''Ficus simplicissima'' is an Asian species of fig tree in the family Moraceae. This species is similar to '' Ficus triloba'' and synonyms include ''Ficus hirta''; its native range is Nepal to southern China and Indo-China, Sumatra and Java. ...
'' Lour. (synonym '' Ficus hirta'' Vahl) *''
Ficus triloba ''Ficus triloba''Voigt JO (1845) In: ''Hort. Suburb. Calcutt.'': 284. is an Asian species of fig tree in the family Moraceae. It is dioecious, with male and female flowers produced on separate individuals. Nomenclature This species is similar ...
'' Buch.-Ham. ex Voigt


Subgenus ''Pharmacosycea''

*'' Ficus crassiuscula'' Standl. *'' Ficus gigantosyce'' Dugand *''
Ficus insipida ''Ficus insipida'' is a common tropical tree in the fig genus of the family Moraceae growing in forest habitats along rivers. It ranges from Mexico to northern South America. Taxonomy The tree was described in 1806 under the scientific name ' ...
'' Willd. *''
Ficus lacunata ''Ficus lacunata'' is a species of plant in the family Moraceae which is endemic to Ecuador. ''F. lacunata'' is a free-standing tree which grows up to 25 m (82 ft) tall in wet forests in the Andes. ''Ficus lacunata'' is member of ...
'' Kvitvik *'' Ficus maxima'' Mill. *'' Ficus mutabilis'' Bureau *'' Ficus nervosa'' Heyne ex Roth *'' Ficus pulchella'' Schott *''
Ficus yoponensis ''Ficus yoponensis'' is a species of fig tree found in Central and South America. It can grow to heights of tall, having a trunk diameter of . The trunk is buttressed, light grey in colour and reasonably smooth. Its petioles are long, the sti ...
'' Desv.


Subgenus ''Sycidium''

*'' Ficus andamanica'' Corner *'' Ficus aspera'' G.Forst. *'' Ficus assamica'' Miq. *'' Ficus bojeri'' Baker *''
Ficus capreifolia The river sandpaper fig (''Ficus capreifolia'') is a fig shrub or small tree of the western and eastern Afrotropics The Afrotropical realm is one of Earth's eight biogeographic realms. It includes Africa south of the Sahara Desert, the majo ...
'' Delile *''
Ficus coronata ''Ficus coronata'', commonly known as the sandpaper fig or creek sandpaper fig, is a species of fig tree, native to Australia. It is found along the east coast from Mackay in Central Queensland, through New South Wales and just into Victoria ...
'' Spin – creek sandpaper fig *''
Ficus fraseri ''Ficus fraseri'', the white sandpaper fig or shiny sandpaper fig, is one of several fig species commonly known as sandpaper figs. It is native to New South Wales, Queensland and the Northern Territory in Australia and to New Caledonia and ...
'' Miq. – shiny sandpaper fig *'' Ficus heterophylla'' L.f. *'' Ficus lateriflora'' Vahl *'' Ficus montana'' Burm.f. – oakleaf fig *''
Ficus opposita ''Ficus opposita'' is one of several fig species commonly known as sandpaper fig The sandpaper figs are so named for their leaves, which are rough and sandpaper-like in texture. The common name may refer to a number of species in the genus ''Fic ...
'' Miq. – sweet sandpaper fig *'' Ficus phaeosyce'' K.Schum. & Lauterb. *''
Ficus tinctoria ''Ficus tinctoria'', also known as dye fig, or humped fig is a hemiepiphytic tree of genus ''Ficus''. It is also one of the species known as '' strangler fig''. It is found in Asia, Malesia, northern Australia, and the South Pacific islands. ...
'' G.Forst. – dye fig *'' Ficus ulmifolia'' Lam. *'' Ficus wassa'' Roxb. *'' Ficus parietalis'' *'' Ficus sinuata'' *'' Ficus hampelas''


Subgenus ''Sycomorus''

*''
Ficus auriculata ''Ficus auriculata'', the Roxburgh fig, is a type of fig tree, native to Asia, noted for its big and round leaves. Description This plant is a small tree of high with numerous bristle-covered branches. The leaves are big and round, and are up t ...
'' Lour. – Roxburgh fig *'' Ficus bernaysii'' King *'' Ficus dammaropsis'' Diels – highland breadfruit, ''kapiak'' *'' Ficus fistulosa'' Blume *'' Ficus hispida'' L. *'' Ficus nota'' Merr. – ''tibig'' *'' Ficus pseudopalma'' Blanco *'' Ficus racemosa'' L. – cluster fig *'' Ficus septica'' Burm.f. – hauli tree *'' Ficus sycomorus'' L., 1753 – sycamore fig (Africa) *'' Ficus variegata'' Blume


Subgenus ''Synoecia''

The following species are typically spreading or climbing
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a t ...
s: *'' Ficus hederacea'' Roxb. *''
Ficus pantoniana ''Ficus pantoniana'', commonly known as the climbing fig, is a species of fig tree, native to Australia. It is found in lowland rainforests on the Cape York Peninsula in northern Queensland. It is a woody climber which may reach 3–6 m in hei ...
'' King – climbing fig *'' Ficus pumila'' L. – creeping fig ** ''Ficus pumila'' var. ''awkeotsang'' (Makino) Corner – jelly fig *'' Ficus punctata'' Thunb. *'' Ficus sagittata'' J. König ex
Vahl Vahl is a surname. Notable people with the surname include: * Christian-Friedrich Vahl (born 1955), German cardiac surgeon * Emanuel Vahl (born 1938), Ukrainian-Israeli composer *Herbert-Ernst Vahl, German SS general *Jens Vahl (1796–1854), Danis ...
*'' Ficus sarmentosa'' Buch.-Ham. ex Sm. *'' Ficus trichocarpa'' Blume *'' Ficus villosa'' Blume


Subgenus ''

Urostigma A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as ...
''

*''
Ficus abutilifolia ''Ficus abutilifolia'', the large-leaved rock fig, is a species of African rock-splitting fig that occurs in two disjunct regions, one population north, and another south of the equator. The two populations are pollinated by different fig wasps, ...
'' Miq. *''
Ficus albert-smithii ''Ficus albert-smithii'' is a species of plant in the family Moraceae. It is found in Brazil, Colombia, Guyana, Peru, and Venezuela. References

Ficus, albert-smithii Least concern plants Trees of Peru Taxonomy articles created by Polbot ...
'' Standl. *'' Ficus altissima'' Blume *''
Ficus amazonica ''Ficus amazonica'' is a species of flowering plant in the family Moraceae. It is found in Brazil, Guyana, Trinidad and Tobago, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Boliva ...
'' Miq. *'' Ficus americana'' Aubl. *''
Ficus aripuanensis ''Ficus aripuanensis'' is a species of plant in the family Moraceae. It is endemic to Mato Grosso and Pará Pará is a state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of ...
'' Berg & Kooy *''
Ficus arpazusa ''Ficus pertusa'' is a species of tree in the family Moraceae. It is found in Mexico and Central and South America. Description Trees up to 25 m tall. Leaves lanceolate, elliptic or ovate, with acuminate or acute apex. Figs edible, globose, 0. ...
'' Carauta and Diaz – Brazil *'' Ficus aurea'' Nutt. – Florida strangler fig *''
Ficus beddomei ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species exten ...
'' King – ''thavital'' *'' Ficus benghalensis'' L. – Indian banyan *'' Ficus benjamina'' L. – weeping fig *'' Ficus binnendijkii'' Miq. *'' Ficus bizanae'' Hutch. & Burtt-Davy *'' Ficus blepharophylla'' Vázquez Avila *''
Ficus broadwayi ''Ficus broadwayi'' is a species of plant in the family Moraceae. It is found in Brazil, Colombia, Suriname, and Venezuela. References broadwayi Least concern plants Taxonomy articles created by Polbot {{Moraceae-stub ...
'' Urb. *''
Ficus burtt-davyi ''Ficus burtt-davyi'' is a fig species endemic to Southern Africa, belonging to the Mulberry family of Moraceae. It grows in coastal and inland forests up to 1500m, from the vicinity of Mossel Bay in the Southern Cape to southern Mozambique - ...
'' Hutch. *''
Ficus calyptroceras ''Ficus calyptroceras'' is a species of Ficus, fig in the family Moraceae, in Brazil. Description The tree is endemic to Brazil, in the states of Bahia, Goiás, Minas Gerais, Mato Grosso do Sul, and Piauí. It is an IUCN Red List Vulnerable spe ...
'' Miq. *'' Ficus castellviana'' Dugand *'' Ficus catappifolia'' Kunth & Bouché *'' Ficus citrifolia'' Mill. – short-leaved fig *'' Ficus consociata'' Bl. *''
Ficus cordata ''Ficus cordata'', the Namaqua rock fig, or Namaqua fig is a species of fig that occurs in two disjunct populations in Africa, one in the arid southwest of the continent, and a second in the northern subtropics. In the south it is often the large ...
'' Thunb. *'' Ficus costata'' Ait. *'' Ficus crassipes'' F.M.Bailey – round-leaved banana fig *'' Ficus craterostoma'' Mildbr. & Burret *'' Ficus cyathistipula'' Warb. *'' Ficus cyclophylla'' (Miq.) Miq. *''
Ficus dendrocida ''Ficus dendrocida'' is a species of plant in the family Moraceae. It is found in Brazil, Colombia, Panama, Venezuela, and eastern Bolivia. In Bolivia, it is one of a few closely related trees in the genus ''Ficus ''Ficus'' ( or ) is a genus ...
'' Kunth *'' Ficus depressa'' Bl. *''
Ficus destruens ''Ficus destruens'' is a hemiepiphytic fig that is endemic to the wet tropical rainforests of northeastern Queensland, Australia. Description ''Ficus destruens'' is a monoecious tree which grows up to tall. Its leaves are long and wide. ...
'' F.White *'' Ficus drupacea'' Thunb. *'' Ficus elastica'' Hornem. – rubber plant *'' Ficus exasperata'' Vahl. *'' Ficus faulkneriana'' Berg *'' Ficus fergusonii'' (King) T.B.Worth. ex Corner *''
Ficus glaberrima ''Ficus glaberrima'' is an Asian species of fig tree in the family Moraceae. The native range of this species is India, S. China and tropical Asia: Indo-China to the Lesser Sunda Islands (but not Borneo, Sulawesi or the Philippines). The spec ...
'' Blume *''
Ficus glumosa ''Ficus glumosa'' also known as the Mountain or Hairy Rock Fig is an Afrotropical fig shrub or tree, growing up to 20 m tall. It is found over a range of altitudes and broken terrain types, including kopjes, outcrops, escarpments and lava flows, ...
'' Delile *'' Ficus greiffiana'' Dugand *'' Ficus hirsuta'' Schott *'' Ficus ilicina'' Miq. *'' Ficus kerkhovenii'' Valeton – Johore fig *'' Ficus kurzii'' King *'' Ficus luschnathiana'' Miq. *'' Ficus ingens'' Miq. *'' Ficus krukovii'' Standl. *''
Ficus lacor ''Ficus lacor'' is a large evergreen tree of the family Moraceae. It is the city tree of Chongqing Chongqing ( or ; ; Sichuanese pronunciation: , Standard Mandarin pronunciation: ), alternately romanized as Chungking (), is a munici ...
'' Buch.-Ham. *''
Ficus lapathifolia ''Ficus lapathifolia'' is a species of plant in the family Moraceae. It is endemic to Mexico. Location Ficus lapathifolia is common to the Gulf region and is also found in the Jalisco Dry Forest habitat that extends along the Pacific coast o ...
'' Miq. *'' Ficus lauretana'' Vázquez Avila *'' Ficus lutea'' Vahl *'' Ficus lyrata'' Warb. – fiddle-leaved fig *'' Ficus maclellandii'' King – Alii fig *''
Ficus macrophylla ''Ficus macrophylla'', commonly known as the Moreton Bay fig or Australian banyan, is a large evergreen banyan tree of the family Moraceae native to eastern Australia, from the Wide Bay–Burnett region in the north to the Illawarra in New ...
'' Desf. ex Pers. – Moreton Bay fig *'' Ficus malacocarpa'' Standl. *''
Ficus mariae ''Ficus mariae'' is a species of tree in the family Moraceae. It is native to South America. References mariae Trees of Peru Trees of Brazil Trees of Bolivia {{Moraceae-stub ...
'' Berg, Emygdio & Carauta *'' Ficus mathewsii'' Miq. *'' Ficus matiziana'' Dugand *'' Ficus mexiae'' Standl. *'' Ficus microcarpa'' L. – Chinese banyan *'' Ficus muelleriana'' Berg *'' Ficus natalensis'' Hochst. – Natal fig *'' Ficus obliqua'' G.Forst. – small-leaved fig *'' Ficus obtusifolia'' Kunth *'' Ficus pakkensis'' Standl. *'' Ficus pallida'' Vahl *'' Ficus panurensis'' Standl. *'' Ficus pertusa'' L.f. *'' Ficus petiolaris'' Kunth *'' Ficus pisocarpa'' Bl. *'' Ficus platypoda'' Cunn. – desert fig *'' Ficus pleurocarpa'' DC. – banana fig *'' Ficus polita'' Vahl *''
Ficus religiosa ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
'' L. – sacred fig *'' Ficus roraimensis'' Berg *'' Ficus rubiginosa'' Desf. – Port Jackson fig *'' Ficus rumphii'' Blume *'' Ficus salicifolia'' Vahl – willow-leaved fig *'' Ficus sansibarica'' Warb. *'' Ficus schippii'' Standl. *'' Ficus schultesii'' Dugand *'' Ficus schumacheri'' Griseb. *''
Ficus sphenophylla ''Ficus sphenophylla'' is a species of fig tree in the family Moraceae. It is found in Bolivia, Brazil, Colombia, and Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_sym ...
'' Standl. *'' Ficus stuhlmannii'' Warb. *'' Ficus subcordata'' Bl. *'' Ficus subpisocarpa'' Gagnep. *'' Ficus subpuberula'' Corner *'' Ficus sumatrana'' Miq. *''
Ficus superba ''Ficus superba'', also known as sea fig or deciduous fig, is a hemiepiphytic tree of genus ''Ficus''. It is one of the species known as banyans or "strangler figs" because of its potential to grow as a hemi-epiphyte and eventually progress to ...
'' Miq. ** ''Ficus superba'' var. ''henneana'' (Miq.) Corner *'' Ficus thonningii'' Blume *'' Ficus trichopoda'' Baker *''
Ficus trigona Ficus trigona is a species of tree in the family Moraceae. It is native to South America. Characteristics ''Ficus trigona'' typically starts life as an epiphyte on another tree. Eventually the plant will send roots to the ground in order to s ...
'' L.f. *'' Ficus trigonata'' L. *'' Ficus triradiata'' Corner – red-stipule fig *'' Ficus ursina'' Standl. *''
Ficus velutina ''Ficus velutina'' is a species of plant in the family Moraceae. It is found in Brazil and Costa Rica. References velutina Least concern plants Taxonomy articles created by Polbot {{Moraceae-stub ...
'' Willd. *''
Ficus verruculosa ''Ficus verruculosa'', the water fig, is a species of fig from sub-saharan Africa. It is found from north eastern South Africa, northern Botswana and Namibia to Uganda and west to Nigeria in riverine and swamp fringes or grassland, always near w ...
'' Warb. *'' Ficus virens'' Aiton – white fig ** ''Ficus virens'' var. ''sublanceolata'' (Miq.) Corner – sour fig *'' Ficus watkinsiana'' F.M.Bailey – Watkins's fig


Unknown subgenus

*'' Ficus bibracteata'' *'' Ficus callosa'' Willd. *''
Ficus cristobalensis ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species exten ...
'' *'' Ficus hebetifolia'' *'' Ficus tsjahela'' Burm.f. * '' Ficus nymphaeifolia'' Mill.


Uses

The wood of fig trees is often soft and the
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosperms ...
precludes its use for many purposes. It was used to make
mummy A mummy is a dead human or an animal whose soft tissues and organs have been preserved by either intentional or accidental exposure to chemicals, extreme cold, very low humidity, or lack of air, so that the recovered body does not decay fu ...
caskets in Ancient Egypt. Certain fig species (mainly ''F. cotinifolia'', '' F. insipida'' and '' F. padifolia'') are traditionally used in
Mesoamerica Mesoamerica is a historical region and cultural area in southern North America and most of Central America. It extends from approximately central Mexico through Belize, Guatemala, El Salvador, Honduras, Nicaragua, and northern Costa Rica. Wit ...
to produce '' papel amate'' (
Nahuatl Nahuatl (; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahua peoples, most of whom live mainly in Central Mexico and have small ...
: ''āmatl''). ''Mutuba'' ('' F. natalensis'') is used to produce barkcloth in Uganda. ''Pou'' ('' F. religiosa'') leaves' shape inspired one of the standard '' kbach rachana'', decorative elements in Cambodian architecture. Indian banyan (''F. benghalensis'') and the Indian rubber plant, as well as other species, have use in
herbalism Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern reme ...
. The inner bark of an unknown type of wild fig, locally known as ''urú'', was once used by the of Bolivia to produce a fibrous cloth used for clothing. Figs have figured prominently in some human cultures. There is evidence that figs, specifically the
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the worl ...
(''F. carica'') and sycamore fig ('' Ficus sycomorus''), were among the first – if not the very first – plant species that were deliberately bred for agriculture in the Middle East, starting more than 11,000 years ago. Nine
subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
''F. carica'' figs dated to about 9400–9200 BCE were found in the early
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several pa ...
village Gilgal I (in the Jordan Valley, 13 km, or 8.1 mi, north of
Jericho Jericho ( ; ar, أريحا ; he, יְרִיחוֹ ) is a Palestinian city in the West Bank. It is located in the Jordan Valley, with the Jordan River to the east and Jerusalem to the west. It is the administrative seat of the Jericho ...
). These were a
parthenogenetic Parthenogenesis (; from the Greek grc, παρθένος, translit=parthénos, lit=virgin, label=none + grc, γένεσις, translit=génesis, lit=creation, label=none) is a natural form of asexual reproduction in which growth and development ...
type and thus apparently an early cultivar. This find predates the first known cultivation of
grain A grain is a small, hard, dry fruit ( caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
in the Middle East by many hundreds of years. Fig is a popular species in the practice of bonsai. It is particularly popular with beginners, as it is widely available and survives well as an indoor plant.


Cultivation

Numerous species of fig are found in cultivation in domestic and office environments, including: *'' F. carica'', common fig – hardy to . Shrub or small tree which can be grown outdoors in mild temperate regions, producing substantial harvests of fruit. Many cultivars are available. *'' F. benjamina'', weeping fig, ficus – hardy to . Widely used as an indoor plant for the home or the office. It benefits from the dry, warm atmosphere of centrally-heated interiors, and can grow to substantial heights in a favoured position. Several variegated cultivars are available. *'' F. elastica'', rubber plant – hardy to : widely cultivated as a houseplant; several cultivars with variegated leaves *'' F. lyrata'', fiddle-leaf fig – hardy to *'' F. maclellandii'' – hardy to *'' F. microcarpa'', Indian laurel – hardy to *'' F. pumila'', creeping fig – hardy to *'' F. rubiginosa'', Port Jackson fig – hardy to


Cultural and spiritual significance

Fig trees have profoundly influenced culture through several religious traditions. Among the more famous species are the
sacred fig ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
tree (Pipal, bodhi, bo, or po, ''Ficus religiosa'') and other banyan figs such as ''Ficus benghalensis''. The oldest living plant of known planting date is a ''Ficus religiosa'' tree known as the Sri Maha Bodhi planted in the temple at
Anuradhapura Anuradhapura ( si, අනුරාධපුරය, translit=Anurādhapuraya; ta, அனுராதபுரம், translit=Aṉurātapuram) is a major city located in north central plain of Sri Lanka. It is the capital city of North Central ...
, Sri Lanka by King Tissa in 288 BCE. The common fig is one of two significant trees in
Islam Islam (; ar, ۘالِإسلَام, , ) is an Abrahamic monotheistic religion centred primarily around the Quran, a religious text considered by Muslims to be the direct word of God (or '' Allah'') as it was revealed to Muhammad, the ...
, and there is a
sura A ''surah'' (; ar, سورة, sūrah, , ), is the equivalent of "chapter" in the Qur'an. There are 114 ''surahs'' in the Quran, each divided into '' ayats'' (verses). The chapters or ''surahs'' are of unequal length; the shortest surah ('' Al-K ...
in Quran named "The Fig" or At-Tin (سوره تین). In Asia, figs are important in
Buddhism Buddhism ( , ), also known as Buddha Dharma and Dharmavinaya (), is an Indian religion or philosophical tradition based on teachings attributed to the Buddha. It originated in northern India as a -movement in the 5th century BCE, and ...
and
Hinduism Hinduism () is an Indian religion or '' dharma'', a religious and universal order or way of life by which followers abide. As a religion, it is the world's third-largest, with over 1.2–1.35 billion followers, or 15–16% of the global p ...
. In
Jainism Jainism ( ), also known as Jain Dharma, is an Indian religion. Jainism traces its spiritual ideas and history through the succession of twenty-four tirthankaras (supreme preachers of ''Dharma''), with the first in the current time cycle bein ...
, the consumption of any fruit belonging to this genus is prohibited. The
Buddha Siddhartha Gautama, most commonly referred to as the Buddha, was a wandering ascetic and religious teacher who lived in South Asia during the 6th or 5th century BCE and founded Buddhism. According to Buddhist tradition, he was born in L ...
is traditionally held to have found ''
bodhi The English term enlightenment is the Western translation of various Buddhist terms, most notably bodhi and vimutti. The abstract noun ''bodhi'' (; Sanskrit: बोधि; Pali: ''bodhi''), means the knowledge or wisdom, or awakened intellec ...
'' (enlightenment) while meditating for 49 days under a
sacred fig ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
. The same species was ''
Ashvattha According to Hindu scriptures, Aśvattha, ( sa, अश्वत्थ) or ''Sacred fig'' (''Ficus religiosa''), is a sacred tree for the Hindus and has been extensively mentioned in texts pertaining to Hinduism, for example as ''peepul'' in Rig V ...
'', the "
world tree The world tree is a motif present in several religions and mythologies, particularly Indo-European religions, Siberian religions, and Native American religions. The world tree is represented as a colossal tree which supports the heavens, thereb ...
" of Hinduism. The '' Plaksa Pra-sravana'' was said to be a fig tree between the roots of which the
Sarasvati River The Sarasvati River () is a deified river first mentioned in the Rigveda and later in Vedic and post-Vedic texts. It played an important role in the Vedic religion, appearing in all but the fourth book of the Rigveda. As a physical river, ...
sprang forth; it is usually held to be a sacred fig but more probably is '' Ficus virens''. According to the
Kikuyu people The Kikuyu (also ''Agĩkũyũ/Gĩkũyũ'') are a Bantu ethnic group native to Central Kenya. At a population of 8,148,668 as of 2019, they account for 17.13% of the total population of Kenya, making them Kenya's largest ethnic group. The t ...
, sacrifices to Ngai were performed under a sycomore tree (Mũkũyũ) and if one was not available, a fig tree (Mũgumo) would be used. The common fig tree is cited in the
Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity Christianity is an Abrahamic monotheistic religion based on the life and teachings of Jesus ...
, where in Genesis 3:7,
Adam and Eve Adam and Eve, according to the creation myth of the Abrahamic religions, were the first man and woman. They are central to the belief that humanity is in essence a single family, with everyone descended from a single pair of original ancestors. ...
cover their nakedness with fig leaves. The fig fruit is also one of the traditional crops of Israel, and is included in the list of food found in the Promised Land, according to the Torah ( Deut. 8). Jesus cursed a fig tree for bearing no fruit (). The fig tree was sacred in ancient
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
and
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ...
, where it was a symbol of
fertility Fertility is the capability to produce offspring through reproduction following the onset of sexual maturity. The fertility rate is the average number of children born by a female during her lifetime and is quantified demographically. Ferti ...
. File:Ficus religiosa Bo.jpg, Leaves of the sacred fig (''
Ficus religiosa ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
'') File:เศียรพระพุทธรูปในรากโพธิ์.jpg, Fig tree roots overgrowing a sandstone Buddha statue, near Wat Maha That in Ayutthaya province, Thailand Image:Sarkaradevi Temple Ficus Tree.jpg, Ficus tree in front of Sarkaradevi Temple,
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South Ca ...
, India


List of famous fig trees

* ''
Ashvattha According to Hindu scriptures, Aśvattha, ( sa, अश्वत्थ) or ''Sacred fig'' (''Ficus religiosa''), is a sacred tree for the Hindus and has been extensively mentioned in texts pertaining to Hinduism, for example as ''peepul'' in Rig V ...
'' – the
world tree The world tree is a motif present in several religions and mythologies, particularly Indo-European religions, Siberian religions, and Native American religions. The world tree is represented as a colossal tree which supports the heavens, thereb ...
of
Hinduism Hinduism () is an Indian religion or '' dharma'', a religious and universal order or way of life by which followers abide. As a religion, it is the world's third-largest, with over 1.2–1.35 billion followers, or 15–16% of the global p ...
, held to be a supernatural ''F. religiosa'' * Bodhi tree – a ''F. religiosa'' * Charybdis Fig Tree of
Homer Homer (; grc, Ὅμηρος , ''Hómēros'') (born ) was a Greek poet who is credited as the author of the ''Iliad'' and the ''Odyssey'', two epic poems that are foundational works of ancient Greek literature. Homer is considered one of the ...
's ''
Odyssey The ''Odyssey'' (; grc, Ὀδύσσεια, Odýsseia, ) is one of two major ancient Greek epic poems attributed to Homer. It is one of the oldest extant works of literature still widely read by modern audiences. As with the '' Iliad'', ...
'', presumably a ''F. carica'' *
Curtain Fig Tree Curtain Fig Tree is a heritage-listed tree at Curtain Fig Tree Road, Yungaburra, Tablelands Region, Queensland, Australia. It is one of the largest trees in Tropical North Queensland, Australia, and one of the best known attractions on the ...
– a ''F. virens'' * Ficus Ruminalis – a ''F. carica'' * '' Plaksa'' – another supernatural fig in Hinduism; usually identified as ''F. religiosa'' but is probably ''F. virens'' *
Santa Barbara's Moreton Bay Fig Tree Santa Barbara's Moreton Bay Fig Tree located in Santa Barbara, California is believed to be the largest ''Ficus macrophylla'' in the United States.Days, M. L. (1977). ''Histories of individual parks Santa Barbara California''. Santa Barbara, CA: ...
– a ''F. macrophylla'' * Sri Maha Bodhi – another ''F. religiosa'', planted in 288 BCE, the oldest human-planted tree on record * '' The Barren Fig Tree'' – Matthew 21:19 of The Holy Bible, Jesus put a curse on the tree and used this as an example for believers of the promise of the power faith in the only true God. '' * The Great Banyan – a ''F. benghalensis'', a
clonal colony A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in ...
and once the largest organism known * Vidurashwatha – " Vidura's Sacred Fig Tree", a village in India named after a famous ''F. religiosa'' that until recently stood there * Wonderboom – the largest fig tree in Pretoria, South Africa


Citations


General references

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Supporting Online Material
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External links



Major reference site for the genus ''Ficus''
World checklist of Ficus species from the Catalogue of Life
845 species supplied by M. Hassler's World Plants.

Multi-award-winning documentary


BBC: Fig fossil clue to early farming
Video
How the fig tree strangles other plants for survival in the rainforest
{{Authority control Moraceae genera Taxa named by Carl Linnaeus