Fig Wasp
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Fig Wasp
Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while the pollinators are in the family Agaonidae. While pollinating fig wasps are gall-makers, the remaining types either make their own galls or usurp the galls of other fig wasps; reports of their being parasitoids are considered dubious. History Aristotle recorded in his ''History of Animals'' that the fruits of the wild fig (the caprifig) contain ''psenes'' (fig wasps); these begin life as grubs (larvae), and the adult ''psen'' splits its "skin" (pupa) and flies out of the fig to find and enter a cultivated fig, saving it from dropping. He believed that the ''psen'' was generated spontaneously; he did not recognise that the fig was reproducing sexually and that the ''psen'' was assisting in that process. Taxonomy The fig wasps are a pol ...
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Agaonidae
The family Agaonidae is a group of pollinating and nonpollinating fig wasps. They spend their larval stage inside the fruits of figs. The pollinating wasps (Agaoninae, Kradibiinae, and Tetrapusiinae) are the mutualistic partners of the fig trees. The nonpollinating fig wasps are parasitic. Extinct forms from the Eocene and Miocene are nearly identical to modern forms, suggesting that the niche has been stable over geologic time. Taxonomy The family has changed several times since its taxonomic appearance after the work of Francis Walker in 1846 described from the wasp genus '' Agaon''. Previously the subfamilies Epichrysomallinae, Otitesellinae, Sycoecinae, Sycoryctinae, Sycophaginae, and Agaoninae were the subdivisions of the family. Recent works building strong molecular phylogenies with an extended sampling size have changed the composition of Agaonidae. First, the paraphyletic groups have been excluded (Epichrysomallinae, Otitesellinae, Sycoecinae, and Sycoryctinae) and ne ...
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Sycophaginae
The family Agaonidae is a group of pollinating and nonpollinating fig wasps. They spend their larval stage inside the fruits of figs. The pollinating wasps (Agaoninae, Kradibiinae, and Tetrapusiinae) are the mutualistic partners of the fig trees. The nonpollinating fig wasps are parasitic. Extinct forms from the Eocene and Miocene are nearly identical to modern forms, suggesting that the niche has been stable over geologic time. Taxonomy The family has changed several times since its taxonomic appearance after the work of Francis Walker in 1846 described from the wasp genus '' Agaon''. Previously the subfamilies Epichrysomallinae, Otitesellinae, Sycoecinae, Sycoryctinae, Sycophaginae, and Agaoninae were the subdivisions of the family. Recent works building strong molecular phylogenies with an extended sampling size have changed the composition of Agaonidae. First, the paraphyletic groups have been excluded (Epichrysomallinae, Otitesellinae, Sycoecinae, and Sycoryctinae) and ne ...
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Blastophaga Psenes
''Blastophaga psenes'' is a wasp species in the genus '' Blastophaga''. It pollinates the common fig ''Ficus carica'' and the closely related ''Ficus palmata''. Without a colony or nest, these wasps breed in figs and the adults live for only a few days or weeks. They locate the fig they wish to pollinate through olfactory senses. Taxonomy and phylogenetics Mutualism occurs between fig and fig wasps, which creates a need for specific species of figs to be pollinated by specific species of wasps. The origin of mutualism is also the beginning of the fig wasp phylogeny. In the phylogenetic tree, the genus of '' Blastophaga'' and ''Wiebesia'' are very similar. Both of these genera pollinate ''Ficus'' species of figs. Description and identification ''B. psenes'' are small wasps, approximately only in length. The females are black wasps and seem shiny while the males are smaller than females. While males are wingless, females have wings that are transparent and very thin. Yet fe ...
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Blastophaga Psenes2
''Blastophaga'' is a wasp genus in the family Agaonidae (fig wasps) which pollinate figs or are otherwise associated with figs, a coevolutional relationship that has been developing for at least 80 million years. Pollinating fig wasps are specific to specific figs. The common fig ''Ficus carica'' is pollinated by ''Blastophaga psenes ''Blastophaga psenes'' is a wasp species in the genus '' Blastophaga''. It pollinates the common fig ''Ficus carica'' and the closely related ''Ficus palmata''. Without a colony or nest, these wasps breed in figs and the adults live for only a f ...''. References * Proctor, M., Yeo, P. & Lack, A. (1996). ''The Natural History of Pollination''. Timber Press, Portland, OR. External links The Fig Web. ''Blastophaga'' Agaonidae Hymenoptera genera Taxa named by Johann Ludwig Christian Gravenhorst {{Chalcidoidea-stub ...
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Polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of convergent evolution. The arrangement of the members of a polyphyletic group is called a polyphyly .. ource for pronunciation./ref> It is contrasted with monophyly and paraphyly. For example, the biological characteristic of warm-bloodedness evolved separately in the ancestors of mammals and the ancestors of birds; "warm-blooded animals" is therefore a polyphyletic grouping. Other examples of polyphyletic groups are algae, C4 photosynthetic plants, and edentates. Many taxonomists aim to avoid homoplasies in grouping taxa together, with a goal to identify and eliminate groups that are found to be polyphyletic. This is often the stimulus for major revisions of the classification schemes. Researchers concerned more with ecology than with systema ...
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Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism for the union of sperm with eggs. Flowers may facilitate outcrossing (fusion of sperm and eggs from different individuals in a population) resulting from cross-pollination or allow selfing (fusion of sperm and egg from the same flower) when self-pollination occurs. There are two types of pollination: self-pollination and cross-pollination. Self-pollination occurs when the pollen from the anther is deposited on the stigma of the same flower, or another flower on the same plant. Cross-pollination is when pollen is transferred from the anther of one flower to the stigma of another flower on a different individual of the same species. Self-pollination happens in flowers where the stamen and carpel mature at the same time, and are positi ...
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Ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typically its form is adapted to functions such as preparing a place for the egg, transmitting the egg, and then placing it properly. For most insects, the organ is used merely to attach the egg to some surface, but for many parasitic species (primarily in wasps and other Hymenoptera), it is a piercing organ as well. Some ovipositors only retract partly when not in use, and the basal part that sticks out is known as the scape, or more specifically oviscape, the word ''scape'' deriving from the Latin word '' scāpus'', meaning "stalk" or "shaft". In insects Grasshoppers use their ovipositors to force a burrow into the earth to receive the eggs. Cicadas pierce the wood of twigs with their ovipositors to insert the eggs. Sawflies slit the ...
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Bagworm
The Psychidae (bagworm moths, also simply bagworms or bagmoths) are a family of the Lepidoptera (butterflies and moths). The bagworm family is fairly small, with about 1,350 species described. Bagworm species are found globally, with some, such as the snailcase bagworm (''Apterona helicoidella''), in modern times settling continents where they are not native. Another common name for the Psychidae is "case moths", but this is just as well used for the case-bearers (Coleophoridae). The names refer to the habits of caterpillars of these two families, which build small protective cases in which they can hide. The bagworms belong to the superfamily Tineoidea, which is a basal lineage of the Ditrysia (as is Gelechioidea, which includes case-bearers). This means that the bagworms and case-bearers are only as closely related to each other as either is to butterflies (Rhopalocera). Most bagworms are inoffensive to humans and inconspicuous; some are occasional nuisance pests. However ...
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Strepsiptera
The Strepsiptera are an order of insects with eleven extant families that include about 600 described species. They are endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches. Females of most species never emerge from the host after entering its body, finally dying inside it. The early-stage larvae do emerge because they must find an unoccupied living host, and the short-lived males must emerge to seek a receptive female in her host. They are believed to be most closely related to beetles, from which they diverged 300–350 million years ago, but do not appear in the fossil record until the mid-Cretaceous around 100 million years ago. The order is not well known to non-specialists, and the nearest they have to a common name is stylops. The name of the order translates to "twisted wing"', giving rise to other common names used for the order, twisted-wing insects and twisted-winged parasites. Adult males are rarely observed, although specimen ...
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Syconium
Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flowers, so it is considered both a multiple and accessory fruit. Etymology The term ''syconium'' comes from the Ancient Greek word (''sykon''), meaning "fig". Morphology The syconium is an urn-shaped receptacle which contains between 50 and 7000 (depending on the species) highly simplified uniovulate flowers or florets on its inner surface. It is closed off from most organisms by the ostiole, fringed by scale-like bracts. Syconia can be monoecious or functionally dioecious: the former contain female flowers with variable style length and few male flowers, and produce seeds and pollen. The latter have male and female forms in different plants: seed figs contain female flowers with long styles and produce seeds; gall figs contain female ...
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