Cretaceous Geology Of Texas
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The Cretaceous ( ) is a
geological period The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronol ...
that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire
Phanerozoic The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...
. The name is derived from the Latin ''creta'', " chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
marine reptile Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. The earliest marine reptile mesosaurus (not to be confused with mosasaurus), arose in the Permian period during the ...
s, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s and
birds Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth by the end of the Cretaceous, coincident with the decline and extinction of previously widespread gymnosperm groups. The Cretaceous (along with the Mesozoic) ended with the Cretaceous–Paleogene extinction event, a large mass extinction in which many groups, including non-avian dinosaurs, pterosaurs, and large marine reptiles, died out. The end of the Cretaceous is defined by the abrupt
Cretaceous–Paleogene boundary The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end of ...
(K–Pg boundary), a geologic signature associated with the mass extinction that lies between the Mesozoic and
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
Eras.


Etymology and history

The Cretaceous as a separate period was first defined by Belgian geologist
Jean d'Omalius d'Halloy Jean Baptiste Julien d'Omalius d'Halloy (17 February 1783 in Liège – 15 January 1875 in Brussels) was a Belgian geologist. He also wrote on races. Early life and education Born in Liège, he was the only son of an ancient and noble family ...
in 1822 as the ''Terrain Crétacé'', using
strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as ei ...
in the Paris Basin and named for the extensive beds of chalk ( calcium carbonate deposited by the shells of marine invertebrates, principally coccoliths), found in the upper Cretaceous of Western Europe. The name Cretaceous was derived from Latin ''creta'', meaning ''chalk''. The twofold division of the Cretaceous was implemented by Conybeare and Phillips in 1822. Alcide d'Orbigny in 1840 divided the French Cretaceous into five ''étages'' (stages): the Neocomian, Aptian, Albian, Turonian, and Senonian, later adding the ''Urgonian'' between Neocomian and Aptian and the Cenomanian between the Albian and Turonian.


Geology


Subdivisions

The Cretaceous is divided into
Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...
and Late Cretaceous epochs, or Lower and Upper Cretaceous series. In older literature, the Cretaceous is sometimes divided into three series: Neocomian (lower/early), Gallic (middle) and Senonian (upper/late). A subdivision into 12 stages, all originating from European stratigraphy, is now used worldwide. In many parts of the world, alternative local subdivisions are still in use. From youngest to oldest, the subdivisions of the Cretaceous period are:


Boundaries

The lower boundary of the Cretaceous is currently undefined, and the Jurassic–Cretaceous boundary is currently the only system boundary to lack a defined Global Boundary Stratotype Section and Point (GSSP). Placing a GSSP for this boundary has been difficult because of the strong regionality of most biostratigraphic markers, and the lack of any chemostratigraphic events, such as isotope excursions (large sudden changes in ratios of isotopes) that could be used to define or correlate a boundary. Calpionellids, an enigmatic group of planktonic protists with urn-shaped calcitic
tests Test(s), testing, or TEST may refer to: * Test (assessment), an educational assessment intended to measure the respondents' knowledge or other abilities Arts and entertainment * ''Test'' (2013 film), an American film * ''Test'' (2014 film), ...
briefly abundant during the latest Jurassic to earliest Cretaceous, have been suggested as the most promising candidates for fixing the Jurassic–Cretaceous boundary. In particular, the first appearance '' Calpionella alpina'', coinciding with the base of the eponymous Alpina subzone, has been proposed as the definition of the base of the Cretaceous. The working definition for the boundary has often been placed as the first appearance of the ammonite '' Strambergella jacobi'', formerly placed in the genus ''
Berriasella ''Berriasella'' is a discoidal evolute perisphinctacean ammonite, and type genus for the neocomitid subfamily Berriasellinae. Its ribbing is distinct, consisting of both simple and bifurcated ribs that extend from the umbilical seam across the ...
'', but its use as a stratigraphic indicator has been questioned, as its first appearance does not correlate with that of ''C. alpina''. The boundary is officially considered by the International Commission on Stratigraphy to be approximately 145 million years ago, but other estimates have been proposed based on U-Pb geochronology, ranging as young as 140 million years ago. The upper boundary of the Cretaceous is sharply defined, being placed at an iridium-rich layer found worldwide that is believed to be associated with the Chicxulub impact crater, with its boundaries circumscribing parts of the Yucatán Peninsula and extending into the Gulf of Mexico. This layer has been dated at 66.043 Mya. At the end of the Cretaceous, the impact of a large body with the Earth may have been the punctuation mark at the end of a progressive decline in biodiversity during the Maastrichtian age. The result was the extinction of three-quarters of Earth's plant and animal species. The impact created the sharp break known as the K–Pg boundary (formerly known as the K–T boundary). Earth's biodiversity required substantial time to recover from this event, despite the probable existence of an abundance of vacant ecological niches. Despite the severity of the K-Pg extinction event, there were significant variations in the rate of extinction between and within different
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
s. Species that depended on photosynthesis declined or became extinct as atmospheric particles blocked
solar energy Solar energy is radiant light and heat from the Sun that is harnessed using a range of technologies such as solar power to generate electricity, solar thermal energy (including solar water heating), and solar architecture. It is an essenti ...
. As is the case today, photosynthesizing organisms, such as
phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...
and land plants, formed the primary part of the food chain in the late Cretaceous, and all else that depended on them suffered, as well. Herbivorous animals, which depended on plants and plankton as their food, died out as their food sources became scarce; consequently, the top predators, such as '' Tyrannosaurus rex'', also perished. Yet only three major groups of tetrapods disappeared completely; the nonavian dinosaurs, the plesiosaurs and the pterosaurs. The other Cretaceous groups that did not survive into the Cenozoic the
ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...
s, last remaining temnospondyls (
Koolasuchus ''Koolasuchus'' is an extinct genus of brachyopoid temnospondyl in the family Chigutisauridae. Fossils have been found from Victoria, Australia and date back 120 Ma to the Aptian stage of the Early Cretaceous. ''Koolasuchus'' is the youngest kno ...
), and nonmammalian were already extinct millions of years before the event occurred.
Coccolithophorids Coccolithophores, or coccolithophorids, are single celled organisms which are part of the phytoplankton, the autotrophic (self-feeding) component of the plankton community. They form a group of about 200 species, and belong either to the kingd ...
and
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s, including
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
s, rudists,
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s, and
mussel Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and Freshwater bivalve, freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other ...
s, as well as organisms whose food chain included these shell builders, became extinct or suffered heavy losses. For example, ammonites are thought to have been the principal food of mosasaurs, a group of giant marine lizards related to snakes that became extinct at the boundary. Omnivores, insectivores, and
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
-eaters survived the extinction event, perhaps because of the increased availability of their food sources. At the end of the Cretaceous, there seem to have been no purely herbivorous or
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s. Mammals and birds that survived the extinction fed on insects, larvae, worms, and snails, which in turn fed on dead plant and animal matter. Scientists theorise that these organisms survived the collapse of plant-based food chains because they fed on
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commun ...
. In
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
communities A community is a Level of analysis, social unit (a group of living things) with commonality such as place (geography), place, Norm (social), norms, religion, values, Convention (norm), customs, or Identity (social science), identity. Communiti ...
, few groups of animals became extinct. Stream communities rely less on food from living plants and more on detritus that washes in from land. This particular ecological niche buffered them from extinction. Similar, but more complex patterns have been found in the oceans. Extinction was more severe among animals living in the water column than among animals living on or in the seafloor. Animals in the water column are almost entirely dependent on primary production from living phytoplankton, while animals living on or in the ocean floor feed on detritus or can switch to detritus feeding. The largest air-breathing survivors of the event,
crocodilian Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
s and champsosaurs, were semiaquatic and had access to detritus. Modern crocodilians can live as scavengers and can survive for months without food and go into hibernation when conditions are unfavorable, and their young are small, grow slowly, and feed largely on invertebrates and dead organisms or fragments of organisms for their first few years. These characteristics have been linked to crocodilian survival at the end of the Cretaceous.


Geologic formations

The high sea level and warm climate of the Cretaceous meant large areas of the continents were covered by warm, shallow seas, providing habitat for many marine organisms. The Cretaceous was named for the extensive chalk deposits of this age in Europe, but in many parts of the world, the deposits from the Cretaceous are of
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
limestone, a rock type that is formed under warm, shallow marine conditions. Due to the high sea level, there was extensive space for such sedimentation. Because of the relatively young age and great thickness of the system, Cretaceous rocks are evident in many areas worldwide. Chalk is a rock type characteristic for (but not restricted to) the Cretaceous. It consists of coccoliths, microscopically small
calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...
skeletons of coccolithophores, a type of
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
that prospered in the Cretaceous seas. Stagnation of deep sea currents in middle Cretaceous times caused anoxic conditions in the sea water leaving the deposited organic matter undecomposed. Half of the world's petroleum reserves were laid down at this time in the anoxic conditions of what would become the Persian Gulf and the Gulf of Mexico. In many places around the world, dark anoxic
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especial ...
s were formed during this interval, such as the Mancos Shale of western North America. These shales are an important source rock for oil and gas, for example in the subsurface of the North Sea.


Europe

In northwestern Europe, chalk deposits from the Upper Cretaceous are characteristic for the
Chalk Group The Chalk Group (often just called the Chalk) is the lithostratigraphic unit (a certain number of rock strata) which contains the Upper Cretaceous limestone succession in southern and eastern England. The same or similar rock sequences occur acr ...
, which forms the white cliffs of Dover on the south coast of England and similar cliffs on the
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
Normandian coast. The group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily
consolidated Consolidated may refer to: *Consolidated (band) **'' ¡Consolidated!'', a 1989 extended play *Consolidated Aircraft (later Convair), an aircraft manufacturer *Consolidated city-county *Consolidated Communications * Consolidated school district *Co ...
and the Chalk Group still consists of loose sediments in many places. The group also has other limestones and arenites. Among the fossils it contains are
sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...
s, belemnites,
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
s and sea reptiles such as '' Mosasaurus''. In southern Europe, the Cretaceous is usually a marine system consisting of
competent Competence may refer to: *Competence (geology), the resistance of a rock against deformation or plastic flow. *Competence (human resources), a standardized requirement for an individual to properly perform a specific job *Competence (law), the me ...
limestone beds or incompetent
marl Marl is an earthy material rich in carbonate minerals, clays, and silt. When hardened into rock, this becomes marlstone. It is formed in marine or freshwater environments, often through the activities of algae. Marl makes up the lower part o ...
s. Because the Alpine mountain chains did not yet exist in the Cretaceous, these deposits formed on the southern edge of the European
continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
, at the margin of the Tethys Ocean.


North America

During the Cretaceous, the present North American continent was isolated from the other continents. In the Jurassic, the North Atlantic already opened, leaving a proto-ocean between Europe and North America. From north to south across the continent, the Western Interior Seaway started forming. This inland sea separated the elevated areas of Laramidia in the west and
Appalachia Appalachia () is a cultural region in the Eastern United States that stretches from the Southern Tier of New York State to northern Alabama and Georgia. While the Appalachian Mountains stretch from Belle Isle in Newfoundland and Labrador, Ca ...
in the east. Three dinosaur clades found in Laramidia (troodontids, therizinosaurids and oviraptorosaurs) are absent from Appalachia from the Coniacian through the Maastrichtian.


Paleogeography

During the Cretaceous, the late- Paleozoic-to-early-Mesozoic supercontinent of Pangaea completed its tectonic breakup into the present-day continents, although their positions were substantially different at the time. As the Atlantic Ocean widened, the convergent-margin mountain building (
orogenies Orogeny is a mountain building process. An orogeny is an event that takes place at a convergent plate margin when plate motion compresses the margin. An ''orogenic belt'' or ''orogen'' develops as the compressed plate crumples and is uplifted t ...
) that had begun during the Jurassic continued in the
North American Cordillera The North American Cordillera, sometimes also called the Western Cordillera of North America, the Western Cordillera or the Pacific Cordillera, is the North American portion of the American Cordillera, the mountain chain system (cordillera) alon ...
, as the
Nevadan orogeny The Nevadan orogeny occurred along the western margin of North America during the Middle Jurassic to Early Cretaceous time which is approximately from 155 Ma to 145 Ma. Throughout the duration of this orogeny there were at least two different ki ...
was followed by the
Sevier Sevier ( ) is an unincorporated community in southwestern Sevier County, Utah, United States. It lies in the valley of the Sevier River along U.S. Route 89 southwest of the city of Richfield, the county seat of Sevier County. Its elevation is ...
and Laramide orogenies.
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
had begun to break up during the Jurassic Period, but its fragmentation accelerated during the Cretaceous and was largely complete by the end of the period. South America, Antarctica, and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
rifted away from Africa (though India and Madagascar remained attached to each other until around 80 million years ago); thus, the South Atlantic and Indian Oceans were newly formed. Such active rifting lifted great undersea mountain chains along the welts, raising eustatic sea levels worldwide. To the north of Africa the Tethys Sea continued to narrow. During the most of the Late Cretaceous, North America would be divided in two by the Western Interior Seaway, a large interior sea, separating Laramidia to the west and
Appalachia Appalachia () is a cultural region in the Eastern United States that stretches from the Southern Tier of New York State to northern Alabama and Georgia. While the Appalachian Mountains stretch from Belle Isle in Newfoundland and Labrador, Ca ...
to the east, then receded late in the period, leaving thick marine deposits sandwiched between coal beds. At the peak of the Cretaceous
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, one-third of Earth's present land area was submerged. The Cretaceous is justly famous for its chalk; indeed, more chalk formed in the Cretaceous than in any other period in the
Phanerozoic The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...
. Mid-ocean ridge activity—or rather, the circulation of seawater through the enlarged ridges—enriched the oceans in calcium; this made the oceans more saturated, as well as increased the bioavailability of the element for calcareous nanoplankton. These widespread carbonates and other
sedimentary deposits Superficial deposits (or surficial deposits) refer to geological deposits typically of Quaternary age (less than 2.6 million years old). These geologically recent unconsolidated sediments may include stream channel and floodplain deposits, beach ...
make the Cretaceous rock record especially fine. Famous formations from North America include the rich marine fossils of Kansas's Smoky Hill Chalk Member and the terrestrial fauna of the late Cretaceous Hell Creek Formation. Other important Cretaceous exposures occur in Europe (e.g., the Weald) and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
(the Yixian Formation). In the area that is now India, massive lava beds called the
Deccan Traps The Deccan Traps is a large igneous province of west-central India (17–24°N, 73–74°E). It is one of the largest volcanic features on Earth, taking the form of a large shield volcano. It consists of numerous layers of solidified flood ...
were erupted in the very late Cretaceous and early Paleocene.


Climate

Palynological evidence indicates the Cretaceous climate had three broad phases: a Berriasian–Barremian warm-dry phase, a Aptian–Santonian warm-wet phase, and a Campanian–Maastrichtian cool-dry phase. The cooling trend of the last epoch of the Jurassic continued into the Berriasian, the first age of the Cretaceous. There is evidence that snowfalls were common in the higher latitudes during this age, and the tropics became wetter than during the Triassic and Jurassic. Glaciation was however restricted to high- latitude mountains, though seasonal snow may have existed farther from the poles. Rafting by ice of stones into marine environments occurred during much of the Cretaceous, but evidence of deposition directly from glaciers is limited to the Early Cretaceous of the Eromanga Basin in southern
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. After the end of the first age, however, temperatures increased again, and these conditions were almost constant until the end of the period. The warming may have been due to intense volcanic activity which produced large quantities of carbon dioxide. Between 70 and 69 Ma and 66–65 Ma, isotopic ratios indicate elevated atmospheric CO2 pressures with levels of 1000–1400 ppmV and mean annual temperatures in west Texas between . Atmospheric CO2 and temperature relations indicate a doubling of pCO2 was accompanied by a ~0.6 °C increase in temperature. The production of large quantities of magma, variously attributed to mantle plumes or to extensional tectonics, further pushed sea levels up, so that large areas of the continental crust were covered with shallow seas. The Tethys Sea connecting the tropical oceans east to west also helped to warm the global climate. Warm-adapted plant fossils are known from localities as far north as Alaska and Greenland, while dinosaur fossils have been found within 15 degrees of the Cretaceous south pole. It was suggested that there was
Antarctic The Antarctic ( or , American English also or ; commonly ) is a polar region around Earth's South Pole, opposite the Arctic region around the North Pole. The Antarctic comprises the continent of Antarctica, the Kerguelen Plateau and other ...
marine glaciation in the Turonian Age, based on isotopic evidence. However, this has subsequently been suggested to be the result of inconsistent isotopic proxies, with evidence of polar rainforests during this time interval at 82° S. A very gentle
temperature gradient A temperature gradient is a physical quantity that describes in which direction and at what rate the temperature changes the most rapidly around a particular location. The temperature gradient is a dimensional quantity expressed in units of degree ...
from the
equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can als ...
to the poles meant weaker global winds, which drive the ocean currents, resulted in less upwelling and more stagnant oceans than today. This is evidenced by widespread black
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especial ...
deposition and frequent anoxic events. Sediment cores show that tropical sea surface temperatures may have briefly been as warm as , warmer than at present, and that they averaged around . Meanwhile, deep ocean temperatures were as much as warmer than today's.


Flora

Flowering plants (angiosperms) make up around 90% of living plant species today. Prior to the rise of angiosperms, during the Jurassic and the Early Cretaceous, the higher flora was dominated by gymnosperm groups, including cycads,
conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
, ginkgophytes, gnetophytes and close relatives, as well as the extinct Bennettitales. Other groups of plants included
pteridosperms The term Pteridospermatophyta (or "seed ferns" or "Pteridospermatopsida") is a polyphyletic group of extinct seed-bearing plants (spermatophytes). The earliest fossil evidence for plants of this type is the genus ''Elkinsia'' of the late Devonia ...
or "seed ferns", a collective term that refers to disparate groups of extinct seed plants with fern-like foliage, including groups such as Corystospermaceae and Caytoniales. The exact origins of angiosperms are uncertain, although molecular evidence suggests that they are not closely related to any living group of gymnosperms. The earliest widely accepted evidence of flowering plants are monosulcate (single-grooved)
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
grains from the late Valanginian (~ 134 million years ago) found in Israel and Italy, initially at low abundance. Molecular clock estimates conflict with fossil estimates, suggesting the diversification of crown-group angiosperms during the Upper Triassic or Jurassic, but such estimates are difficult to reconcile with the heavily sampled pollen record and the distinctive tricolpate to tricolporoidate (triple grooved) pollen of
eudicot The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicot ...
angiosperms. Among the oldest records of Angiosperm macrofossils are ''
Montsechia ''Montsechia'' is an extinct genus of aquatic plants containing the species ''Montsechia vidalii'', discovered in Spain. ''Montsechia vidalii'' lived about 130 million years ago, during the Barremian age, and appears to be the earliest known flo ...
'' from the Barremian aged Las Hoyas beds of Spain and '' Archaefructus'' from the Barremian-Aptian boundary Yixian Formation in China. Tricolpate pollen distinctive of eudicots first appears in the Late Barremian, while the earliest remains of monocots are known from the Aptian. Flowering plants underwent a rapid radiation beginning during the middle Cretaceous, becoming the dominant group of land plants by the end of the period, coincident with the decline of previously dominant groups such as conifers. The oldest known fossils of grasses are from the Albian, with the family having diversified into modern groups by the end of the Cretaceous. The oldest large angiosperm trees are known from the Turonian (c. 90 Mya) of New Jersey, with the trunk having a preserved diameter of and an estimated height of . During the Cretaceous, Polypodiales ferns, which make up 80% of living fern species, would also begin to diversify.


Terrestrial fauna

On land,
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s were generally small sized, but a very relevant component of the fauna, with
cimolodont Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order (biology), order Multituberculata. They probably lived something of a rodent-like existence u ...
multituberculates outnumbering dinosaurs in some sites. Neither true marsupials nor placentals existed until the very end, but a variety of non-marsupial metatherians and non-placental eutherians had already begun to diversify greatly, ranging as carnivores ( Deltatheroida), aquatic foragers ( Stagodontidae) and herbivores ('' Schowalteria'', Zhelestidae). Various "archaic" groups like eutriconodonts were common in the Early Cretaceous, but by the Late Cretaceous northern mammalian faunas were dominated by multituberculates and therians, with dryolestoids dominating South America. The apex predators were
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
ian
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s, especially dinosaurs, which were at their most diverse stage. Avians such as the ancestors of modern-day
birds Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
also diversified. They inhabited every continent, and were even found in cold polar latitudes. Pterosaurs were common in the early and middle Cretaceous, but as the Cretaceous proceeded they declined for poorly understood reasons (once thought to be due to competition with early birds, but now it is understood avian adaptive radiation is not consistent with pterosaur decline). By the end of the period only three highly specialized families remained; Pteranodontidae, Nyctosauridae, and
Azhdarchidae Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cret ...
. The
Liaoning Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmost ...
lagerstätte ( Yixian Formation) in China is an important site, full of preserved remains of numerous types of small dinosaurs, birds and mammals, that provides a glimpse of life in the Early Cretaceous. The coelurosaur dinosaurs found there represent types of the group Maniraptora, which includes modern birds and their closest non-avian relatives, such as dromaeosaurs, oviraptorosaurs, therizinosaurs, troodontids along with other avialans. Fossils of these dinosaurs from the
Liaoning Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmost ...
lagerstätte are notable for the presence of hair-like
feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier ...
s. Insects diversified during the Cretaceous, and the oldest known ants, termites and some
lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
ns, akin to
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
and moths, appeared. Aphids,
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s and gall wasps appeared. File:Tyrannosaurus-rex-Profile-steveoc86.png, '' Tyrannosaurus rex'', one of the largest land predators of all time, lived during the Late Cretaceous File: Velociraptor Restoration.png, Up to 2 m long and 0.5 m high at the hip, '' Velociraptor'' was feathered and roamed the Late Cretaceous File: Triceratops by Tom Patker.png, '' Triceratops'', one of the most recognizable genera of the Cretaceous File:Quetzalcoatlus07.jpg, The azhdarchid ''
Quetzalcoatlus ''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, ...
'', one of the largest animals to ever fly, lived during the Late Cretaceous File:Confuciusornis sanctus mmartyniuk.png, '' Confuciusornis'', a genus of crow-sized birds from the Early Cretaceous File:Ichthyornis restoration.jpeg, '' Ichthyornis'' was a toothed, seabird-like ornithuran from the Late Cretaceous


Rhynchocephalians

Rhynchocephalia Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse g ...
ns (which today only includes the Tuatara) disappeared from North America and Europe after the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
, and were absent from North Africa and northern South America by the early Late Cretaceous. The cause of the decline of Rhynchocephalia remains unclear, but has often been suggested to be due to competition with advanced lizards and mammals. They appear to have remained diverse in high-latitude southern South America during the Late Cretaceous, where lizards remained rare, with their remains outnumbering terrestrial lizards 200:1.


Choristodera

Choristoderes Choristodera (from the Greek χωριστός ''chōristos'' + δέρη ''dérē'', 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the late Miocene (168 to ...
, a group of freshwater aquatic reptiles that first appeared during the preceding Jurassic, underwent a major evolutionary radiation in Asia during the Early Cretaceous, which represents the high point of choristoderan diversity, including long necked forms such as '' Hyphalosaurus'' and the first records of the gharial-like Neochoristodera, which appear to have evolved in the regional absence of aquatic neosuchian crocodyliformes. During the Late Cretaceous the neochoristodere '' Champsosaurus'' was widely distributed across western North America.


Marine fauna

In the seas, rays, modern sharks and teleosts became common. Marine reptiles included
ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...
s in the early and mid-Cretaceous (becoming extinct during the late Cretaceous Cenomanian-Turonian anoxic event),
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
s throughout the entire period, and mosasaurs appearing in the Late Cretaceous. Sea turtles in the form of Cheloniidae and Panchelonioidea lived during the period and survived the extinction event. Panchelonioidea is today represented by a single species; the leatherback sea turtle. '' Baculites'', an
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
genus with a straight shell, flourished in the seas along with reef-building rudist clams. The
Hesperornithiformes Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as ''Hesperorni ...
were flightless, marine diving birds that swam like grebes. Globotruncanid Foraminifera and
echinoderms An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea li ...
such as sea urchins and starfish (sea stars) thrived. Thylacocephala, a class of crustaceans, went extinct in the Late Cretaceous. The first radiation of the
diatom A diatom (Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising sev ...
s (generally siliceous shelled, rather than calcareous) in the oceans occurred during the Cretaceous; freshwater diatoms did not appear until the Miocene. The Cretaceous was also an important interval in the evolution of
bioerosion Bioerosion describes the breakdown of hard ocean substrates – and less often terrestrial substrates – by living organisms. Marine bioerosion can be caused by mollusks, polychaete worms, phoronids, sponges, crustaceans, echinoids, and ...
, the production of borings and scrapings in rocks,
hardgrounds Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor (Wilson and Palmer, 1992). A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limesto ...
and shells. File:Kronosaurus hunt1DB.jpg, A scene from the early Cretaceous: a ''
Woolungasaurus ''Woolungasaurus'' ('Woolunga lizard', named after an Aboriginal mythical reptile, Persson 1960) is a plesiosaur, an extinct marine reptile, belonging to the Elasmosauridae. The type species, ''Woolungasaurus glendowerensis'', was named after ...
'' is attacked by a '' Kronosaurus''. File:Tylosaurus pembinensis 1DB.jpg, '' Tylosaurus'' was a large mosasaur, carnivorous marine reptiles that emerged in the late Cretaceous. File:Hesperornis BW (white background).jpg, Strong-swimming and toothed predatory waterbird '' Hesperornis'' roamed late Cretacean oceans. File:DiscoscaphitesirisCretaceous.jpg, The
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
'' Discoscaphites iris'', Owl Creek Formation (Upper Cretaceous), Ripley, Mississippi File:The fossils from Cretaceous age found in Lebanon.jpg, A plate with ''
Nematonotus ''Nematonotus'' is an extinct genus of prehistoric bony fish that lived during the Lower Cenomanian. See also * Prehistoric fish * List of prehistoric bony fish A ''list'' is any set of items in a row. List or lists may also refer to: Pe ...
sp.'', ''Pseudostacus sp.'' and a partial ''Dercetis triqueter'', found in Hakel, Lebanon File:Cretoxyrhina attacking Pteranodon.png, ''
Cretoxyrhina ''Cretoxyrhina'' (; meaning 'Cretaceous sharp-nose') is an extinct genus of large mackerel shark that lived about 107 to 73 million years ago during the late Albian to late Campanian of the Late Cretaceous. The type species, ''C. mantelli'', is m ...
'', one of the largest Cretaceous sharks, attacking a ''
Pteranodon ''Pteranodon'' (); from Ancient Greek (''pteron'', "wing") and (''anodon'', "toothless") is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of . They lived during the late Cr ...
'' in the Western Interior Seaway


See also

*
Mesozoic Era The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising ...
* Cretaceous-Paleogene extinction * Chalk Formation *
Cretaceous Thermal Maximum The Cretaceous Thermal Maximum (CTM), also known as Cretaceous Thermal Optimum, was a period of climatic warming that reached its peak approximately 90 million years ago (90 Ma) during the Turonian age of the Late Cretaceous epoch. The CTM is notab ...
* List of fossil sites (with link directory) *
South Polar region of the Cretaceous The South Polar region of the Cretaceous comprised the continent of East Gondwana–modern day Australia and Antarctica–a product of the break-up of Gondwana in the Cretaceous Period. The southern region, during this time, was much warmer than i ...


References


Citations


Bibliography

* * * * * —detailed coverage of various aspects of the evolutionary history of the insects. * * *


External links


UCMP Berkeley Cretaceous pageCretaceous Microfossils: 180+ images of Foraminifera
* {{Authority control Geological periods