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Azhdarchid
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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Phosphatodraco
''Phosphatodraco'' is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species ''Phosphatodraco mauritanicus'' in 2003; the genus name means " dragon from the phosphates", and the specific name refers to the region of Mauretania. ''Phosphatodraco'' was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur '' Tethydraco'' represent wing elements of ''Phosphatodraco''. Due to the fragmentary nature of the holotype cervical vertebrae, there has been co ...
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Hatzegopteryx
''Hatzegopteryx'' ("Hațeg basin wing") is a genus of azhdarchid pterosaur found in the late Maastrichtian deposits of the Densuş Ciula Formation, an outcropping in Transylvania, Romania. It is known only from the type species, ''Hatzegopteryx thambema'', named by Buffetaut ''et al.'' in 2002 based on parts of the skull and humerus. Additional specimens, including a neck vertebra, were later placed in the genus, representing a range of sizes. The largest of these remains indicate it was among the biggest pterosaurs, with an estimated wingspan of . Unusually among giant azhdarchids, ''Hatzegopteryx'' had a very wide skull bearing large muscular attachments; bones with a spongy internal texture instead of hollow; and a short, robust, and heavily muscled neck measuring long, which was about half the length of other azhdarchids with comparable wingspans, and was capable of withstanding strong bending forces. ''Hatzegopteryx'' inhabited Hațeg Island, an island situated in the Cret ...
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Quetzalcoatlus Northropi
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas at ...
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Quetzalcoatlus
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Cryodrakon
''Cryodrakon'' ("cold dragon") is a genus of azhdarchid pterosaur that lived during the Late Cretaceous period in what is now Canada. It contains a single species, ''Cryodrakon boreas'', recovered from the Dinosaur Park Formation. Discovery and history Since 1972, bones of large azhdarchids have been reported from Alberta, representing the first pterosaur finds from Canada. These were sometimes referred to a ''Quetzalcoatlus'' sp. Later research by Michael Habib indicated that they represented a taxon new to science. In 2019, the type species ''Cryodrakon boreas'' was named and described by David William Elliott Hone, Michael Habib and François Therrien. The generic name is derived from the Greek κρύος, ''kryos'', "icy cold", and δράκων, ''drakon'', "dragon". The specific name means "belonging to '' Boreas'', the north wind", or simply "northern". Habib had considered the name ''Cryodrakon viserion'', as a reference to the ice dragon in ''Game of Thrones''. The ...
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Wellnhopterus
''Wellnhopterus'' () is an Azhdarchidae, azhdarchid pterosaur recovered from the Late Cretaceous Javelina Formation in Texas that was previously identified as a thalassodrominae, thalassodromine. It consists of a set of upper and lower jaws (TMM 42489-2), as well as some cervical vertebrae (TMM 42489-2.3 to 42489-2.8) and a fragmentary long bone (TMM 42489-2.9). In July 2021, the jaws were given the genus name "''Javelinadactylus''" (meaning "Javelina Finger"), with the type species, type and only species as "''J. sagebieli''"; however, this article has now been Retractions in academic publishing, retracted. In a paper published in December 2021, the complete holotype was independently named ''Wellnhopterus'', with the only species being ''W. brevirostris''. As of 2022, this is the formal name of this pterosaur. Discovery and naming In March 1986, jaws and neck vertebrae of a large pterosaur were discovered in Brewster County by preparator Robert Rainey. In 1991, Peter Wellnhofer ...
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Alanqa
''Alanqa'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period ( Cenomanian stage) of what is now the Kem Kem Beds of southeastern Morocco. The name ''Alanqa'' comes from the Arabic word العنقاء ''al- ‘Anqā’'', for a mythical bird of Arabian culture. Discovery Aided by local villagers, a team of paleontologists had been excavating at several locations in the Kem Kem Beds during April, and November to December 2008, uncovering remains of several different pterosaurs. The material was fragmentary, and the type locality for ''Alanqa'' is Aferdou N'Chaft, near the village of Begaa and to the north-east of Taouz. Description ''Alanqa'' is known only from five fragments of the front upper and lower jaws, and possibly a neck vertebra, representing the single type species ''Alanqa saharica''. Two of these fragments were first described, but not named, by Wellnhofer and Buffetaut in 1999, and referred to a pteranodontid. Three additional jaw specimens, ...
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Azhdarcho Lancicollis
''Azhdarcho'' is a genus of azhdarchid pterosaur from the late Cretaceous Period of the Bissekty Formation (middle Turonian stage, about 92 million years ago) of Uzbekistan, as well as the Zhirkindek Formation of Kazakhstan. It is known from fragmentary remains including the distinctive, elongated neck vertebrae that characterizes members of the family Azhdarchidae, a family that includes many giant pterosaurs such as ''Quetzalcoatlus''. The name ''Azhdarcho'' comes from the Persian word '' azhdar'' (), a dragon-like creature in Persian mythology. The type species is ''Azhdarcho lancicollis''. The specific epithet ''lancicollis'' is derived from the Latin words ''lancea'' (meaning "lance" or "spear") and ''collum'' ("neck"). History The fossil remains of ''Azhdarcho'' were recovered in the Kyzyl Kum desert (from the Taykarshinskaya unit of the Bissekty Formation) by Lev A. Nesov during expeditions to Central Asia in 1974–1981. The type specimen, given the catalog number Ц ...
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Azhdarcho
''Azhdarcho'' is a genus of azhdarchid pterosaur from the late Cretaceous Period of the Bissekty Formation (middle Turonian stage, about 92 million years ago) of Uzbekistan, as well as the Zhirkindek Formation of Kazakhstan. It is known from fragmentary remains including the distinctive, elongated neck vertebrae that characterizes members of the family Azhdarchidae, a family that includes many giant pterosaurs such as ''Quetzalcoatlus''. The name ''Azhdarcho'' comes from the Persian word '' azhdar'' (), a dragon-like creature in Persian mythology. The type species is ''Azhdarcho lancicollis''. The specific epithet ''lancicollis'' is derived from the Latin words ''lancea'' (meaning "lance" or "spear") and ''collum'' ("neck"). History The fossil remains of ''Azhdarcho'' were recovered in the Kyzyl Kum desert (from the Taykarshinskaya unit of the Bissekty Formation) by Lev A. Nesov during expeditions to Central Asia in 1974–1981. The type specimen, given the catalog number ...
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Eurazhdarcho
''Eurazhdarcho'' is a genus of azhdarchid pterosaur from the Late Cretaceous period (Maastrichtian stage) of what is now the Transylvanian Basin of Romania. Its fossil remains dated back 69 million years ago. Discovery and naming In 2009, Mátyás Vremir at Lancrăm near Sebeş-Glod in Transylvania at the SbG-B site uncovered the remains of a pterosaur. He donated these to the ''Erdélyi Múzeum'', of the '' Societății Muzeului Ardelean'' (Transylvanian Museum Society). Subsequent excavations by Vremir discovered additional bones of the same individual animal and were added by him to the collection of the Babeș-Bolyai University. In 2013, Vremir, Alexander Wilhelm Armin Kellner, Darren Naish, and Gareth Dyke named and described the type species ''Eurazhdarcho langendorfensis''. The generic name combines the name of Europe with that of the related form ''Azhdarcho''. The specific name refers to Langendorf, the name of Lancrǎm in the language of the German ethnic minor ...
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Arambourgiania
''Arambourgiania'' is an extinct genus of azhdarchid pterosaur from the Late Cretaceous period (Maastrichtian stage) of Jordan, and possibly the United States.Harrell, T. Lynn Jr.; Gibson, Michael A.; Langston, Wann Jr. (2016). "A cervical vertebra of ''Arambourgiania philadelphiae'' (Pterosauria, Azhdarchidae) from the Late Campanian micaceous facies of the Coon Creek Formation in McNairy County, Tennessee, USA" ''Bull. Alabama Mus. Nat. Hist.'' 33:94–103 ''Arambourgiania'' was among the largest members of its family, the Azhdarchidae, and it is also one of the largest flying animals ever known. The incomplete left ulna of the "Sidi Chennane azhdarchid" from Morocco may have also belonged to ''Arambourgiania''. History of discovery In the early 1940s, a railway worker during repairs on the Amman-Damascus railroad near Russeifa found a two foot long fossil bone. In 1943 this was acquired by the director of a nearby phosphate mine, Amin Kawar, who brought it to the attention ...
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