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The karuka (''Pandanus julianettii'', also called karuka nut and ''Pandanus'' nut) is a species of tree in the family
Pandanaceae Pandanaceae is a family of flowering plants native to the tropics and subtropics of the Old World, from West Africa through the Pacific. It contains 982 known species in five genera, of which the type genus, ''Pandanus'', is the most important, wi ...
and an important regional food crop in New Guinea. The nuts are more nutritious than
coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family ( Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the ...
s, and are so popular that villagers in the highlands will move their entire households closer to trees for the harvest season.


Names

The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
"''julianettii''" honors naturalist
Amedeo Giulianetti Amedeo is an Italian given name meaning "lover of God", "loves God", or more correctly "for the love of God" and cognate to the Latin name Amadeus and the Spanish and Portuguese Amadeo. People with this name include: * A number of rulers and nob ...
, who found the original type specimens. is a loanword from Tok Pisin. Sometimes the tree is called '' or 'karuka nut pandanus'. The term 'karuka' can apply to both ''Pandanus julianettii'' and '' P. brosimos'', though the latter is usually called 'wild karuka'. Both species, as well as '' P. dubius'', can be called 'pandanus nut'. In addition to ''P. brosimos'', 'wild karuka' can also refer to '' P. antaresensis'', '' P. iwen'', and '' P. limbatus'', but nuts from these trees are a much smaller part of the local diet. In contexts where multiple karuka species are discussed, ''P. julianettii'' is sometimes termed 'planted karuka'. ''P. julianettii'', ''P. iwen'', and ''P. brosimos'' are also in the subsection named '' Karuka'', which is in the monotypic section also named '' Karuka''. In New Guinea it goes by different names among each of the
Papuan people The indigenous peoples of West Papua in Indonesia and Papua New Guinea, commonly called Papuans, are Melanesians. There is genetic evidence for two major historical lineages in New Guinea and neighboring islands: a first wave from the Malay Arc ...
s. In the
Ankave language Ankave or Angave is a Papuan language spoken by the approximately 1,600 () Angave people in Kerema District, Gulf Province, Papua New Guinea. Phonology Vowels Diphthongs: Consonants Writing system An orthography using the Latin script has ...
it is . It is in the
Baruya language Yipma (pronounced as ''Hipma'') is an Angan language of Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilo ...
. The Huli language word is , and it is also in the Duna language. In Kewa language it is ''aga'', but it is unclear which dialect(s). In the Kewa pandanus language it is ''rumala agaa''. The Kalam language term, in both standard and pandanus languages, is , but it can also be called or . The plant is called in the
Wiru language Wiru or Witu is the language spoken by the Wiru people of Ialibu-Pangia District of the Southern Highlands Province of Papua New Guinea. The language has been described by Harland Kerr, a missionary who lived in the Wiru community for many years. ...
. In the
Pole language Kewa is an Engan language complex of the Southern Highlands province of Papua New Guinea. A dictionary of the western dialect of Kewa has been compiled by . Kewa pandanus register Kewa's elaborate pandanus avoidance register, which is used on ...
it's called ''maisene''. It goes by ''ank'' in Angal language, and in the Wola dialect. The
Imbongu language Kaugel (Gawigl) is one of the languages spoken in the Southern Highlands (Papua New Guinea), Southern Highlands province of Papua New Guinea. Native speakers call the area on the Southern Highlands (Papua New Guinea), Southern Highlands side of t ...
word is . The plant also has many names on the other half of the island. In Indonesian it is called ( lit.) and (), but the latter can also refer to ''P. brosimos'' and ''P. iwen''. The Dani people call it ''tuke''. The
Lani people The Lani are an indigenous people in Puncak, Central Papua and Lanny Jaya, Highland Papua, usually labelled 'Western Dani' by foreign missionaries, or grouped—inaccurately—with the Dani people who inhabit the Baliem Valley to the east. Po ...
call it , but this might be a separate species in the complex.


Description

The species was originally described by
Ugolino Martelli Ugolino Martelli (1860–1934) was an Italian botanist, biologist, and mycology, mycologist. Martelli is known for his studies of and contributions to the systematics of the tropical genus ''Pandanus'' and his taxonomic definition of the flora of ...
from only a few
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
s in the collections of the Royal Botanic Gardens, Kew He was hesitant to describe it as a new species from only that, but the characteristics were so salient he published his description. The tree is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
(individual plants either have male flowers or female ones), with male trees uncommon compared to females. It reaches in height, with a grey
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...
of in diameter and supported by buttress roots. The trunk has white mottling and is generally smooth with occasional warts or small knobs as well as rings of leaf scars. Inside the trunk is pithy and lacking cambium. The top of the tree sometimes branches, producing three or four crowns of leaves. Each crown will produce a single cluster of nuts, typically once every other season. Production is affected by the seasonality of local rainfall.
Leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
spiral up the trunk in opposite pairs. The large leathery leaves are long and wide. The apex of the leaf is attenuate and doubly-pleated, with prickles pointing up at the tip and along the
margins Margin may refer to: Physical or graphical edges *Margin (typography), the white space that surrounds the content of a page *Continental margin, the zone of the ocean floor that separates the thin oceanic crust from thick continental crust *Leaf ...
and midrib. The leaves are dark green on top and dull cyan underneath. The inflorescence on male trees is a densely-branched spadix with a dozen long spikes, each containing many staminate phalanges. In each phalange is a
column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member. ...
3 mm long topped by up to 9
subsessile In botany, sessility (meaning "sitting", used in the sense of "resting on the surface") is a characteristic of plant parts (such as flowers and leaves) that have no stalk. Plant parts can also be described as subsessile, that is, not completely ...
anther The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
s. The male flowers are white, and the whole male flowering organ may be up to long. The
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
has a
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