Tikisuchus
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''Tikisuchus'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of rauisuchid
archosauromorph Archosauromorpha ( Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, l ...
. It is known from the
Late Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. ...
Tiki Formation in the
Shahdol District Shahdol District is a district of Madhya Pradesh state in east central India. It old name was Matsya raj or Virat kingdom in the name of great king Maharaja Virata. With a total area of and a population of 10,66,063. Shahdol is an important dist ...
of central
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
and was the first rauisuchid to have been found in Asia. The horizon from which remains have been found is
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227 m ...
in age. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
is ''T. romeri'', named in honor of American paleontologist
Alfred Romer Alfred Sherwood Romer (December 28, 1894 – November 5, 1973) was an American paleontologist and biologist and a specialist in vertebrate evolution. Biography Alfred Romer was born in White Plains, New York, the son of Harry Houston Romer an ...
. Romer was present at the Tiki locality during the excavation of the fossil, but died before the description of the genus in 1987. ''Tikisuchus'' is known only from one specimen, called ISI R 305, which consists of the skull and some postcranial elements of a young individual.


Description

Compared to other rausuchids, the skull of ''Tikisuchus'' was very large. The skull's length is around 40% of the length of the presacral area between the head and the sacrum. The skull is deep, being wide at the back with a narrow rostrum. The teeth are large, recurved and serrated. Like other rauisuchids, it has rows of
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinc ...
s, or bony
scute A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior po ...
s, along its back. There are two rows of osteoderms. Each osteoderm is rectangular in shape and imbricates, or articulates tightly, with the ones around it. In other rauisuchids, the osteoderms are leaf-shaped rather than rectangular.


Paleobiology

Many other tetrapods were found in association with ''Tikisuchus'' representing a diverse Carnian paleofauna. Tetrapods from the Tiki site include '' Paleorhinus'', a
phytosaur Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent g ...
, '' Metoposaurus'', a
temnospondyl Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carbo ...
, and '' Paradapedon'', a
rhynchosaur Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs ...
. The Tiki fauna is similar to that of the
German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...
Keuper The Keuper is a lithostratigraphic unit (a sequence of rock strata) in the subsurface of large parts of west and central Europe. The Keuper consists of dolomite, shales or claystones and evaporites that were deposited during the Middle and Late T ...
.
Theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s were also present in the Tiki Formation. Both ''Tikisuchus'' and the theropods were large terrestrial predators, and, having been found at the same locality, likely came in close contact with one another. The similar lifestyles of the two carnivores may have resulted in competition for the same food sources. Possible prey would have included
rhynchosaur Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs ...
s, trilophosaurs,
dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typic ...
s, and
aetosaur Aetosaurs () are heavily armored reptiles belonging to the extinct order (biology), order Aetosauria (; from Ancient Greek, Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized Omnivore, omnivorous or Herbivore, herbivoro ...
s. The authors of the original description of ''Tikisuchus'',
Sankar Chatterjee Sankar Chatterjee (born May 28, 1943) is a paleontologist, and is the Paul W. Horn Professor of Geosciences at Texas Tech University and Curator of Paleontology at the Museum of Texas Tech University. He earned his Ph. D. from the University of ...
and Pranab Majumdar, suggested that competition between ''Tikisuchus'' and theropods was low because of abundant food resources and stabilized ecological interactions. Chatterjee and Majumdar thought that there was an "ecological balance" during the Carnian stage on the basis of little change in the paleofauna of the time. They considered the paleoclimate to have been warm with seasonal wet and dry seasons conducive to the growth of tropical forests. At the end of the Carnian, however, the authors claimed that many prey resources became extinct and the forest environment was replaced by a more open environment. The limited resources would have heightened competition between theropods and rauisuchids like ''Tikisuchus''. Chatterjee and Majumdar considered theropods to be agile pursuit predators while rauisuchids were considered slow ambush predators. Therefore, they suggested that theropods, which were more suited to living in an open environment, outcompeted rauisuchids at the end of the Triassic to become the dominant large land carnivores by the beginning of the Jurassic. However, more recent studies suggest that dinosaurs gained dominance only after the Triassic-Jurassic extinction event in a case of opportunism with no other large archosaurs such as rauisuchids to compete with.


References

{{Taxonbar, from=Q3281023 Rauisuchids Prehistoric reptiles of Asia Fossil taxa described in 1987 Prehistoric pseudosuchian genera