Archosauromorph
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Archosauromorph
Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period. Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including ''Protorosaurus'', tanystropheids, and ''Prolacerta''. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors. A ...
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Protorosauria
Protorosauria is an extinct polyphyletic group of archosauromorph reptiles from the latest Middle Permian (Capitanian stage) to the end of the Late Triassic (Rhaetian stage) of Asia, Europe and North America. It was named by the English anatomist and paleontologist Thomas Henry Huxley in 1871 as an order, originally to solely contain ''Protorosaurus''. Other names which were once considered equivalent to Protorosauria include Prolacertiformes and Prolacertilia. Protorosaurs are distinguished by their long necks formed by elongated cervical vertebrae, which have ribs that extend backward to the vertebrae behind them. Protorosaurs also have a gap between the quadrate bones and the jugal bones in the back of the skull near the jaw joint, making their skulls resemble those of lizards. While previously thought to be monophyletic, the group is now though to consist of various groups of basal archosauromorph reptiles that lie outside Crocopoda. Classification Protorosauria was consi ...
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Elessaurus
''Elessaurus'' is an extinct genus of archosauromorph from the Early Triassic of Brazil. It contains a single species, ''Elessaurus gondwanoccidens''. It possessed a variety of features common to basal archosauromorphs, particularly basal tanystropheids such as '' Macrocnemus''. However, it is uncertain whether ''Elessaurus'' was a particularly close relative of tanystropheids, and it might instead be closer to other major archosauromorph clades. The genus name refers to "Elessar", an alternate name of the character Aragorn from J.R.R. Tolkien's Lord of the Rings trilogy. Discovery ''Elessaurus'' is known from a single holotype specimen, UFSM 11471, which consists of most of a leg connected to parts of the hip and the base of the tail. Despite being fairly complete by the standards of its locale, many of the specimen's bones are flattened or fragmentary. UFSM 11471 was collected from the Bica São Tomé site, an outcrop of the Sanga do Cabral Formation in Rio Grande do Su ...
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Aenigmastropheus
''Aenigmastropheus'' is an extinct genus of early archosauromorph reptiles known from the middle Late Permian Usili Formation of Songea District, southern Tanzania. It contains a single species, ''Aenigmastropheus parringtoni'', known solely from UMZC T836, a partial postcranial skeleton of a mature individual. It was collected in 1933, and first described in 1956, as a "problematic reptile" due to its unique morphology. Therefore, a binomial name was erected for this specimen in 2014. ''Aenigmastropheus'' was probably fully terrestrial. Discovery Fossils of ''Aenigmastropheus'' were first described by the British paleontologist Dr. Francis Rex Parrington in 1956, in an article titled as "A problematic reptile from the Upper Permian". Parrington reported collecting these remains in the Ruhuhu Valley in the Songea District of southern Tanzania in 1933, and considered them to come from a single individual. This specimen, UMZC T836, in currently housed at the University Museum of ...
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Boreopricea
''Boreopricea'' is an extinct genus of archosauromorph reptile from the Early Triassic of arctic Russia. It is known from a fairly complete skeleton discovered in a borehole on Kolguyev Island, though damage to the specimen and loss of certain bones has complicated study of the genus. ''Boreopricea'' shared many similarities with various other archosauromorphs, making its classification controversial. Various studies have considered it a close relative of ''Prolacerta'', tanystropheids, both, or neither. ''Boreopricea'' is unique among early archosauromorphs due to possessing contact between the jugal and squamosal bones at the rear half of the skull. History ''Boreopricea funerea'' was named and described by Soviet paleontologist L. P. Tatarinov in 1978. It was primarily based on a fairly complete skull and skeleton collected in 1972 from a borehole at Kolguyev Island in the Arctic Ocean. This holotype specimen, PIN 3708/1, included bones from most of the body, apart from the ...
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Tasmaniosaurus
''Tasmaniosaurus'' ('lizard from Tasmania', although this genus is not a true lizard) is an extinct genus of archosauromorph reptile known from the Knocklofty Formation (Early Triassic) of West Hobart, Tasmania, Australia. The type species is ''T. triassicus''. This genus is notable not only due to being one of the most complete Australian Triassic reptiles known, but also due to being a very close relative of Archosauriformes. Once believed to be a proterosuchid, this taxon is now believed to have been intermediate between advanced non-archosauriform archosauromorphs such as ''Prolacerta'', and basal archosauriforms such as ''Proterosuchus''. Features traditionally used to define Archosauria and later Archosauriformes, such as the presence of an antorbital fenestra and serrated teeth, are now known to have evolved prior to those groups due to their presence in ''Tasmaniosaurus''. History and Classification First named as a '' nomen nudum'' in 1974, the genus received a fo ...
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Prolacertoides
''Prolacertoides'' is an extinct genus of archosauromorph reptile from the Early Triassic of China, the type species being ''Prolacertoides jimusarensis''. ''Prolacertoides'' means 'like ''Prolacerta''', in reference to ''Prolacerta'', another genus of archosauromorph which ''Prolacertoides'' was once believed to have been closely related to. ''Prolacertoides'' is known from a single partial skull, IVPP V3233, which was discovered in Xinjiang in northwestern China. The locality of its discovery belongs to the Cangfanggou Group of the Jiucaiyuan Formation, which is dated to the Induan age of the very early Triassic. Description IVPP V3233 is a partially flattened and incomplete skull, with some bones being well preserved and others being badly damaged. The snout is tapering and lacks antorbital fenestrae. The anterodorsal edge of each maxilla is convex, in contrast to the concave edge of other basal archosauromorphs. The ascending process of each maxilla is tall and anteropos ...
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Teyujagua
''Teyujagua'' (named for Teyú Yaguá, a legendary beast from local Guaraní mythology) is an extinct genus of small, probably semi-aquatic archosauromorph reptile that lived in Brazil during the Early Triassic period. The genus contains the type and only known species, ''T. paradoxa''. It is known from a well-preserved skull, and probably resembled a crocodile in appearance. It was an intermediary between the primitive archosauromorphs and the more advanced Archosauriformes, revealing the mosaic evolution of how the key features of the archosauriform skull were acquired. ''Teyujagua'' also provides additional support for a two-phase model of archosauriform radiation, with an initial diversification in the Permian followed by a second adaptive radiation in the Early Triassic. Description ''Teyujagua'' is known only from a well preserved skull with four associated cervical vertebrae, the only known postcranial material, but it is inferred to have been a small, carnivorous quadrupe ...
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Elachistosuchus
''Elachistosuchus'' is an extinct genus of neodiapsid reptile, most likely basal archosauromorph, known from the Late Triassic Arnstadt Formation of Saxony-Anhalt, central Germany. It contains a single species, ''Elachistosuchus huenei'', known from a single individual. ''E. huenei'', originally considered a pseudosuchian archosaur and then a rhynchocephalian lepidosaur, was largely ignored in the scientific literature, as its small size and fragility did not permit further mechanical preparation and examination. More recently however a non-invasive μCT scanning was performed to resolve its placement within Reptilia, and found it to represent a more basal reptile, potentially closely related to several early archosauromorph clades. Discovery Fossils of ''Elachistosuchus'' were first described and named by the German paleontologist Werner Janensch in 1949, with the type species being ''Elachistosuchus huenei''. The generic name is derived from Greek ''ἐλάχιστος'' / ' ...
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Allokotosauria
Allokotosauria is a clade of early archosauromorph reptiles from the Middle to Late Triassic known from Asia, Africa, North America and Europe. Allokotosauria was first described and named when a new monophyletic grouping of specialized herbivorous archosauromorphs was recovered by Sterling J. Nesbitt, John J. Flynn, Adam C. Pritchard, J. Michael Parrish, Lovasoa Ranivoharimanana and André R. Wyss in 2015. The name Allokotosauria is derived from Greek meaning "strange reptiles" in reference to unexpected grouping of early archosauromorph with a high disparity of features typically associated with herbivory. History Nesbitt ''et al.'' (2015) defined the group as a stem-based taxon containing ''Azendohsaurus madagaskarensis'' and '' Trilophosaurus buettneri'' and all taxa more closely related to them than to ''Tanystropheus longobardicus'', ''Proterosuchus fergusi'', ''Protorosaurus speneri'' or ''Rhynchosaurus articeps''. Therefore, Allokotosauria includes the families Azendoh ...
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Archosaur
Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, possessed these features yet originated pri ...
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Reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians (tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, historically combined with that of modern amphibians, is called herpetology. The earliest known proto-reptiles originated around ...
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Archosauriformes
Archosauriformes (Greek for 'ruling lizards', and Latin for 'form') is a clade of diapsid reptiles that developed from archosauromorph ancestors some time in the Latest Permian (roughly 252 million years ago). It was defined by Jacques Gauthier (1994) as the clade stemming from the last common ancestor of Proterosuchidae and Archosauria (the group that contains crocodiles, pterosaurs and dinosaurs bird.html"_;"title="ncluding_bird">ncluding_birds;_Phil_Senter.html" ;"title="bird">ncluding_birds.html" ;"title="bird.html" ;"title="ncluding bird">ncluding birds">bird.html" ;"title="ncluding bird">ncluding birds; Phil Senter">bird">ncluding_birds.html" ;"title="bird.html" ;"title="ncluding bird">ncluding birds">bird.html" ;"title="ncluding bird">ncluding birds; Phil Senter (2005) defined it as the most exclusive clade containing ''Proterosuchus'' and Archosauria. These reptiles, which include members of the Family (biology), family Proterosuchidae and more advanced forms, were ...
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